View source for Biology:2013 in paleobotany

This list of '''2013 in [[Biology:Paleobotany|paleobotany]]''' records new  [[Biology:Fossil|fossil]] [[Biology:Plant|plant]] taxa that were [[Biology:Binomial nomenclature|described]] during 2013, as well as other significant discoveries and events related to [[Biology:Paleobotany|paleobotany]] that occurred in the year.

==[[Biology:Chlorophyta|Chlorophyta]]==
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Unit
! Location
! Notes
! Images
|-
|
''Cymatiosphaera scitula''<ref name=WicanderPlayford2013>{{cite journal |author1=R. Wicander |author2=G. Playford |year=2013 |title=Marine and terrestrial palynofloras from transitional Devonian–Mississippian strata, Illinois Basin, U.S.A. |journal=Boletín Geológico y Minero |volume=124 |issue=4 |pages=589–637 |url=http://aiplanetatierra.igme.es/Boletin/2013/124_4/6_ARTICULO%205.pdf }}</ref>
|
Sp. nov.
|
Valid
|
Wicander & Playford
|
[[Earth:Devonian|Devonian]] ([[Earth:Famennian|Famennian]])
|
Saverton Shale
|
{{Flag|United States}}
|
A member of [[Biology:Chlorophyta|Chlorophyta]] belonging to the group [[Biology:Prasinophyceae|Prasinophyceae]].
|
|-
|}

==[[Biology:Moss|Bryophyta]]==
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Unit
! Location
! Notes
! Images
|-
|
''Brachytheciites sachalinensis''<ref name=Pseudoleskeellites>{{Cite journal|author1=Michael S. Ignatov |author2=Evgeny E. Perkovsky |year=2013 |title=Mosses from Sakhalinian amber (Russian Far East) |journal=Arctoa: A Journal of Bryology |volume=22 |pages=79–82 |doi=10.15298/arctoa.22.11 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Ignatov & Perkovsky
|
Middle [[Earth:Eocene|Eocene]]
|
Sakhalinian amber
|
{{Flag|Russia}}
|
A [[Biology:Moss|moss]] belonging to the family [[Biology:Brachytheciaceae|Brachytheciaceae]].
|
|-
|
''Bryiidites''<ref>{{cite journal |author1=Richard S. Barclay |author2=Jennifer C. McElwain |author3=Jeffrey G. Duckett |author4=Maarten H. van Es |author5=Anika S. Mostaert |author6=Silvia Pressel |author7=Bradley B. Sageman |year=2013 |title=New methods reveal oldest known fossil epiphyllous moss: ''Bryiidites utahensis'' gen. et sp. nov. (Bryidae) |journal=American Journal of Botany |volume=100 |issue=12 |pages=2450–2457 |doi=10.3732/ajb.1300209 |pmid=24302691|s2cid=7217440 }}</ref>
|
Gen. et sp. nov.
|
Valid
|
Barclay ''et al.''
|
Late Cretaceous (late Cenomanian)
|
Middle Dakota Formation
|
{{Flag|United States}}
|
A member of [[Biology:Bryidae|Bryidae]] of uncertain phylogenetic placement. The type specimen is ''Bryiidites utahensis''.
| 
|-
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''Bryokhutuliinia crassimarginata''<ref>{{Cite journal|author1=Michael S. Ignatov |author2=Eugeny E. Karasev |author3=Sofia M. Sinitsa |author4=Elena V. Maslova |year=2013 |title=New ''Bryokhutuliinia'' species (Bryophyta) with sporophytes from the Upper Jurassic of Transbaikalia |journal=Arctoa: A Journal of Bryology |volume=22 |pages=69–78 |doi=10.15298/arctoa.22.10 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Ignatov ''et al.''
|
Late Jurassic
|
|
{{Flag|Russia}}
|
A [[Biology:Moss|moss]].
|
|-
|
''Calymperites''<ref>{{Cite journal|author1=Michael S. Ignatov |author2=Evgeny E. Perkovsky |year=2013 |title=Mosses from Rovno amber (Ukraine), 2 |journal=Arctoa: A Journal of Bryology |volume=22 |pages=83–92 |doi=10.15298/arctoa.22.12 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Ignatov & Perkovsky
|
Late Eocene
|
Rovno amber
|
{{Flag|Ukraine}}
|
A [[Biology:Moss|moss]] belonging to the group [[Biology:Dicranales|Dicranales]] (''sensu lato''). Genus includes new species ''C. ucrainicus''.
|
|-
|
''Pseudoleskeellites''<ref name=Pseudoleskeellites />
|
Gen. et sp. nov
|
Valid
|
Ignatov & Perkovsky
|
Middle [[Earth:Eocene|Eocene]]
|
Sakhalinian amber
|
{{Flag|Russia}}
|
A [[Biology:Moss|moss]] belonging to the order [[Biology:Hypnales|Hypnales]]. Genus includes new species ''P. obscurus''.
|
|-
|}

==Ferns and fern allies==

{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Unit
! Location
! Notes
! Images
|-
|
''Ashicaulis beipiaoensis''<ref>{{Cite journal|author1=Ning Tian |author2=Yongdong Wang |author3=Wu Zhang |author4=Zikun Jiang |author5=David L. Dilcher |year=2013 |title=''Ashicaulis beipiaoensis'' sp. nov., a New Osmundaceous Fern Species from the Middle Jurassic of Liaoning Province, Northeastern China |journal=International Journal of Plant Sciences |volume=174 |issue=3 |pages=328–339 |doi=10.1086/668868 |s2cid=83565443 }}</ref>
|
Sp. nov
|
Valid
|
Tian ''et al.''
|
Middle Jurassic
|
[[Earth:Tiaojishan Formation|Tiaojishan Formation]]
|
{{Flag|China}}
|
A member of [[Biology:Osmundaceae|Osmundaceae]]. Originally described as a species of ''Ashicaulis''; transferred to the genus ''Millerocaulis'' by Bomfleur, Grimm & McLoughlin (2017).<ref>{{Cite journal|author1=Benjamin Bomfleur |author2=Guido W. Grimm |author3=Stephen McLoughlin |year=2017 |title=The fossil Osmundales (Royal Ferns)—a phylogenetic network analysis, revised taxonomy, and evolutionary classification of anatomically preserved trunks and rhizomes |journal=PeerJ |volume=5 |pages=e3433 |doi=10.7717/peerj.3433 |pmid=28713650 |pmc=5508817 |doi-access=free }}</ref>
|
|-
|
''Mygdykitia''<ref name=Mygdykitia>{{cite journal |author1=L.B. Golovneva |author2=S.V. Shczepetov |year=2013 |title=The Yana floristic assemblage from the Upper Cretaceous volcanic deposits of the northern coast of Sea of Okhotsk |journal=Paleobotanika |volume=4 |pages=5–35 |doi=10.31111/palaeobotany/2013.4.5 |url=https://www.researchgate.net/publication/262918366 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Golovneva & Shczepetov
|
[[Earth:Late Cretaceous|Late Cretaceous]] ([[Earth:Campanian|Campanian]])
|
Ola Formation
|
{{Flag|Russia}}
|
A fern belonging to the family [[Biology:Dipteridaceae|Dipteridaceae]]. Genus includes new species ''M. ochotica''.
|
|-
|
''Palaeosorum''<ref>{{Cite journal|author1=Frédéric M. B. Jacques |author2=Tao Su |author3=Zhe-Kun Zhou |year=2013 |title=The first fossil Microsoroid fern (''Palaeosorum ellipticum'' gen. et sp. nov.) from the middle Miocene of Yunnan, SW China |journal=Journal of Systematics and Evolution |volume=51 |issue=6 |pages=758–764 |doi=10.1111/jse.12030 |s2cid=82491404 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Jacques & Zhou ''in'' Jacques, Su & Zhou
|
Miocene
|
Dajie Formation
|
{{Flag|China}}
|
A member of [[Biology:Polypodiaceae|Polypodiaceae]]. The type specimen is ''Palaeosorum ellipticum''.
|
|-
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''Todea amissa''<ref>{{cite journal |author1=Mónica R. Carvalho |author2=Peter Wilf |author3=Elizabeth J. Hermsen |author4=Maria A. Gandolfo |author5=N. Rubén Cúneo |author6=Kirk R. Johnson |year=2013 |title=First record of ''Todea'' (Osmundaceae) in South America, from the early Eocene paleorainforests of Laguna del Hunco (Patagonia, Argentina) |journal=American Journal of Botany |volume=100 |issue=9 |pages=1831–1848 |doi=10.3732/ajb.1200637 |pmid=24018858|hdl=11336/5393 |hdl-access=free }}</ref>
|
Sp. nov.
|
Valid
|
Carvalho ''in'' Carvalho ''et al.''
|
Early [[Earth:Eocene|Eocene]]
|
|
{{Flag|Argentina}}
|
A species of ''[[Biology:Todea|Todea]]''.
| 
|-
|
''[[Biology:Yavanna chimaerica|Yavanna]]''<ref>{{Cite journal|author=Ezequiel Ignacio Vera |year=2013 |title=New cyathealean tree fern, ''Yavanna chimaerica'' gen. et sp. nov., from the Early Cretaceous of Livingston Island, Antarctica |journal=Cretaceous Research |volume=44 |pages=214–222 |doi=10.1016/j.cretres.2013.04.011 |bibcode=2013CrRes..44..214V |hdl=11336/8854 |hdl-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Vera
|
Early Cretaceous ([[Earth:Aptian|Aptian]])
|
Cerro Negro Formation
|
{{Flag|Antarctica}}<br>(Livingston Island)
|
A [[Biology:Cyatheales|cyathealean]] of uncertain placement. Type species is ''Y. chimaerica''.
|
|-
|}

==Ginkgophytes==
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Unit
! Location
! Notes
! Images
|-
|
''Allicospermum angrenicum''<ref name=Nagrenia>{{cite journal |author=Natalya Nosova |year=2013 |title=Revision of the genus ''Grenana'' Samylina from the Middle Jurassic of Angren, Uzbekistan |journal=Review of Palaeobotany and Palynology |volume=197 |pages=226–252 |doi=10.1016/j.revpalbo.2013.06.005 |bibcode=2013RPaPa.197..226N }}</ref>
|
Sp. nov.
|
Valid
|
Nosova
|
Middle Jurassic
|
Angren Formation
|
{{Flag|Uzbekistan}}
|
A ''[[Biology:Ginkgo|Ginkgo]]''-like plant described on the basis of seeds.
|
|-
|
''Nagrenia''<ref name=Nagrenia />
|
Gen. et sp. nov.
|
Valid
|
Nosova
|
Middle Jurassic
|
Angren Formation
|
{{Flag|Uzbekistan}}
|
A member of [[Biology:Ginkgoales|Ginkgoales]] described on the basis of [[Biology:Peduncle (botany)|peduncles]] furcating into several [[Biology:Pedicel (botany)|pedicels]]. Genus includes new species ''N. samylinae''.
|
|-
|}

==[[Biology:Gnetophyta|Gnetophyta]]==
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Unit
! Location
! Notes
! Images
|-
|
''Chengia''<ref>{{cite journal |author=Yong Yang |author2=Longbiao Lin |author3=Qi Wang |name-list-style=amp |year=2013 |title=''Chengia laxispicata'' gen. et sp. nov., a new ephedroid plant from the Early Cretaceous Yixian Formation of western Liaoning, Northeast China: evolutionary, taxonomic, and biogeographic implications |journal=BMC Evolutionary Biology |volume=13 |issue=1 |pages=72 |doi=10.1186/1471-2148-13-72 |pmid=23530702 |pmc=3626868 |bibcode=2013BMCEE..13...72Y |doi-access=free }}</ref>
|
Gen. et sp. nov.
|
Valid
|
Yang, Lin & Wang
|
Early Cretaceous ([[Earth:Aptian|Aptian]])
|
[[Earth:Yixian Formation|Yixian Formation]]
|
{{Flag|China}}
|
A member of [[Biology:Ephedraceae|Ephedraceae]]. The type specimen is ''Chengia laxispicata''.
|
|-
|
''Friedsellowia gracilifolia''<ref name="Löwe2013">{{cite journal |last1=Löwe |first1=S. |last2=Mohr |first2=B.A.R. |last3=Coiffard|first3=C.|last4=Bernardes-de-Oliveira |first4=M.E.C. |year=2013 |title=''Friedsellowia gracilifolia'' gen. nov. et sp. nov., a new gnetophyte from the Lower Cretaceous Crato Formation (Brazil)  |journal= Palaeontographica Abteilung B  |volume=289 |issue=4–6 |pages=139–177 |url= http://www.schweizerbart.de/papers/palb/detail/289/80732/Friedsellowia_gracilifolia_gen_nov_et_sp_nov_a_new}}</ref>
|
Gen. nov. et sp. nov.
|
Valid
|
Löwe et al.
|
Early Cretaceous
|
[[Earth:Crato Formation|Crato Formation]]
|
{{Flag|Brazil}}
|
A [[Biology:Gnetophyta|gnetophyte]]. The type of a new genus.
| 
|-
|
''Itajuba''{{Citation needed|date=December 2019|reason=removed citation to predatory publisher content}}
|
Gen. et sp. nov
|
Valid
|
Ricardi-Branco ''et al.''
|
Early Cretaceous (Aptian or Albian)
|
Crato Member of the Santana Formation
|
{{Flag|Brazil}}
|
A member of [[Biology:Gnetales|Gnetales]]. The type specimen is ''Itajuba yansanae''.
|
|-
|}

==Pachytestopsida== 
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Unit
! Location
! Notes
! Images
|-
|
''Stephanospermum braidwoodensis''<ref>{{cite journal |author1=Alan R.T. Spencer |author2=Jason Hilton |author3=Mark D. Sutton |year=2013 |title=Combined methodologies for three-dimensional reconstruction of fossil plants preserved in siderite nodules: ''Stephanospermum braidwoodensis'' nov. sp. (Medullosales) from the Mazon Creek lagerstätte |journal=Review of Palaeobotany and Palynology |volume=188 |pages=1–17 |doi=10.1016/j.revpalbo.2012.09.001 |bibcode=2013RPaPa.188....1S }}</ref>
|
Sp. nov.
|
Valid
|
Spencer, Hilton & Sutton
|
Carboniferous
|
[[Earth:Mazon Creek fossil beds|Mazon Creek fossil beds]]
|
{{Flag|United States}}
|
A member of [[Biology:Medullosales|Medullosales]], a species of ''Stephanospermum''.
|
|-
|
''Stephanospermum shuichengensis''<ref>{{cite journal |author1=Alan R.T. Spencer |author2=Shi-Jun Wang |author3=Michael T. Dunn |author4=Jason Hilton |year=2013 |title=Species of the medullosan ovule ''Stephanospermum'' from the Lopingian (late Permian) floras of China |journal=Journal of Asian Earth Sciences |volume=76 |pages=59–69 |doi=10.1016/j.jseaes.2013.07.030 |bibcode=2013JAESc..76...59S }}</ref>
|
Sp. nov.
|
Valid
|
Spencer ''et al.''
|
Permian ([[Earth:Lopingian|Lopingian]])
|
Wangjiazhai Formation
|
{{Flag|China}}
|
A member of [[Biology:Medullosales|Medullosales]], a species of ''Stephanospermum''.
|
|-
|}

==[[Biology:Pinophyta|Pinophyta]]==
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Unit
! Location
! Notes
! Images
|-
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''Bancroftiastrobus''<ref>{{cite journal |author=Gar W. Rothwell |author2=Gene Mapes |author3=Ruth A. Stockey |author4=Jason Hilton |name-list-style=amp |year=2013 |title=Diversity of ancient conifers: the Jurassic seed cone ''Bancroftiastrobus digitata'' gen. et sp. nov. (Coniferales) |journal=International Journal of Plant Sciences |volume=174 |issue=6 |pages=937–946 |doi=10.1086/670368 |s2cid=85605229 |url=http://pure-oai.bham.ac.uk/ws/files/9880605/IJPS_Bancroftiastrobus_2013.pdf }}</ref>
|
Gen. et sp. nov.
|
Valid
|
Rothwell ''et al.''
|
Late Jurassic (early Kimmeridgian)
|
[[Earth:Kimmeridge Clay|Kimmeridge Clay]] Formation
|
{{Flag|United Kingdom}}
|
A [[Biology:Pinophyta|pinophyte]] of uncertain phylogenetic placement.<br/>  Type species is ''B. digitata''.
|
|-
|
''Brachyoxylon currumilii''<ref>{{cite journal |author1=Josefina Bodnar |author2=Ignacio Escapa |author3=Néstor Rubén Cúneo |author4=Silvia Gnaedinger |year=2013 |title=First record of conifer wood from the Cañadón Asfalto Formation (Early-Middle Jurassic), Chubut Province, Argentina |journal=Ameghiniana |volume=50 |issue=2 |pages=227–239 |doi=10.5710/AMGH.26.04.2013.620 |hdl=11336/3391 |s2cid=130814427 |hdl-access=free }}</ref>
|
Sp. nov.
|
Valid
|
Bodnar ''et al.''
|
Early–Middle Jurassic
|
[[Earth:Cañadón Asfalto Formation|Cañadón Asfalto Formation]]
|
{{Flag|Argentina}}
|
A member of [[Biology:Pinophyta|Pinophyta]] described on the basis of [[Biology:Fossil wood|fossil wood]].
|
|-
|
''Cunninghamia taylorii''<ref>{{cite journal |author=Rudolph Serbet |author2=Benjamin Bomfleur |author3=Gar W. Rothwell |name-list-style=amp |year=2013 |title=''Cunninghamia taylorii'' sp. nov., a Structurally Preserved Cupressaceous Conifer from the Upper Cretaceous (Campanian) Horseshoe Canyon Formation of Western North America |journal=International Journal of Plant Sciences |volume=174 |issue=3 |pages=471–488 |doi=10.1086/668695 |s2cid=83890547 }}</ref>
|
Sp. nov.
|
Valid
|
Serbet, Bomfleur & Rothwell
|
Late Cretaceous ([[Earth:Campanian|Campanian]])
|
[[Earth:Horseshoe Canyon Formation|Horseshoe Canyon Formation]]
|
{{Flag|Canada}}
|
A species of ''[[Biology:Cunninghamia|Cunninghamia]]''.
| 
|-
|
''Elatocladus anorovii''<ref name=Mygdykitia />
|
Sp. nov
|
Valid
|
Golovneva & Shczepetov
|
[[Earth:Late Cretaceous|Late Cretaceous]] ([[Earth:Campanian|Campanian]])
|
Ola Formation
|
{{Flag|Russia}}
|
A [[Biology:Conifer|conifer]] of uncertain phylogenetic placement.
|
|-
|
''[[Biology:Pagiophyllum|Pagiophyllum heshanense]]''<ref>{{cite journal |author1=Lu-jun Liu |author2=Zhao-qi Yao |year=2013 |title=The conifer-remains from the Permian of South China |journal=Acta Palaeontologica Sinica |volume=52 |issue=2 |pages=182–201 |url=http://eng.oversea.cnki.net/kcms/detail/detail.aspx?dbCode=cjfd&QueryID=62&CurRec=4&filename=GSWX201302006&dbname=CJFD2013 }}</ref>
|
Sp. nov.
|
Valid
|
Liu & Yao
|
Permian
|
|
{{Flag|China}}
|
A species of ''Pagiophyllum''.
| 
|-
|
''[[Biology:Pararaucaria|Pararaucaria delfueyoi]]''<ref>{{cite journal |author1=Ignacio H. Escapa |author2=Néstor R. Cúneo |author3=Gar Rothwell |author4=Ruth A. Stockey |year=2013 |title=''Pararaucaria delfueyoi'' sp. nov. from the Late Jurassic Cañadón Calcáreo Formation, Chubut, Argentina: Insights into the Evolution of the Cheirolepidiaceae |journal=International Journal of Plant Sciences |volume=174 |issue=3 |pages=458–470 |doi=10.1086/668612 |hdl=11336/1531 |s2cid=54809961 |hdl-access=free }}</ref>
|
Sp. nov.
|
Valid
|
Escapa ''et al.''
|
Late Jurassic
|
Cañadón Calcáreo Formation
|
{{Flag|Argentina}}
|
A member of [[Biology:Pinophyta|Pinophyta]] belonging to the family [[Biology:Cheirolepidiaceae|Cheirolepidiaceae]], a species of ''Pararaucaria''.
| 
|-
|
''Picea yanensis''<ref name=Mygdykitia />
|
Sp. nov
|
Valid
|
Golovneva & Shczepetov
|
[[Earth:Late Cretaceous|Late Cretaceous]] ([[Earth:Campanian|Campanian]])
|
Ola Formation
|
{{Flag|Russia}}
|
A [[Biology:Spruce|spruce]].
|
|-
|
''Prumnopitys anglica''<ref>{{cite journal |author=David R. Greenwood |author2=Christopher R. Hill |author3=John G. Conran |name-list-style=amp |year=2013 |title=''Prumnopitys anglica'' sp. nov. (Podocarpaceae) from the Eocene of England |journal=Taxon |volume=62 |issue=3 |pages=565–580 |doi=10.12705/623.15 |doi-access=free }}</ref>
|
Sp. nov.
|
Valid
|
Greenwood, Hill & Conran
|
Eocene
|
Branksome Formation
|
{{Flag|United Kingdom}}
|
A species of ''[[Biology:Prumnopitys|Prumnopitys]]''.
|
|-
|
''Schizolepidopsis canicularis''<ref>{{cite journal |author1=Andrew B. Leslie |author2=Ian Glasspool |author3=Patrick S. Herendeen |author4=Niiden Ichinnorov |author5=Patrick Knopf |author6=Masamichi Takahashi |author7=Peter R. Crane |year=2013 |title=Pinaceae-like reproductive morphology in ''Schizolepidopsis canicularis'' sp. nov. from the Early Cretaceous (Aptian-Albian) of Mongolia |journal=American Journal of Botany |volume=100 |issue=12 |pages=2426–2436 |doi=10.3732/ajb.1300173 |pmid=24285570}}</ref>
|
Sp. nov.
|
Valid
|
Leslie ''et al.''
|
Early Cretaceous (probably Aptian-Albian)
|
Tevshiin Govi Formation
|
{{Flag|Mongolia}}
|
A member of [[Biology:Pinophyta|Pinophyta]], possibly a member of the [[Biology:Crown group#Stem groups|stem group]] of [[Biology:Pinaceae|Pinaceae]]; a species of ''Schizolepidopsis''.
| 
|-
|
''Xuanweioxylon''<ref>{{cite journal |author1=Jian He |author2=Shi-Jun Wang |author3=Jason Hilton |author4=Longyi Shao |year=2013 |title=''Xuanweioxylon scalariforme'' gen. et sp. nov.: Novel Permian coniferophyte stems with scalariform bordered pitting on secondary xylem tracheids |journal=Review of Palaeobotany and Palynology |volume=197 |pages=152–165 |doi=10.1016/j.revpalbo.2013.05.010 |bibcode=2013RPaPa.197..152H |doi-access=free }}</ref>
|
Gen. et sp. nov.
|
Valid
|
He ''et al.''
|
[[Earth:Permian|Permian]] (late [[Earth:Wuchiapingian|Wuchiapingian]]-[[Earth:Changhsingian|Changhsingian]])
|
Xuanwei Formation
|
{{Flag|China}}
|
A member of [[Biology:Pinophyta|Pinophyta]]. Genus includes new species ''X. scalariforme''.
|
|-
|}

==Angiosperms==

{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Unit
! Location
! Notes
! Images
|-
|
''Anonaspermum warmanensis''<ref name=Lauroflorum>{{cite journal |author1=Hongshan Wang |author2=Jane Blanchard |author3=David L. Dilcher |year=2013 |title=Fruits, seeds, and flowers from the Warman clay pit (middle Eocene Claiborne Group), western Tennessee, USA |journal=Palaeontologia Electronica |volume=16 |issue=3 |pages=Article number 16.3.31A |url=http://palaeo-electronica.org/content/2013/545-eocene-plants-from-tennessee }}</ref>
|
Sp. nov.
|
Valid
|
Wang, Blanchard & Dilcher
|
Eocene
|
Cockfield Formation
|
{{Flag|United States}}
|
A member of [[Biology:Annonaceae|Annonaceae]].
|
|-
|
''Atli''<ref>{{cite journal |author1=Selena Y. Smith |author2=Stefan A. Little |author3=Ranessa L. Cooper |author4=Robyn J. Burnham |author5=Ruth A. Stockey |year=2013 |title=A Ranunculalean Liana Stem from the Cretaceous of British Columbia, Canada: ''Atli morinii'' gen. et sp. nov. |journal=International Journal of Plant Sciences |volume=174 |issue=5 |pages=818–831 |doi=10.1086/669925 |s2cid=84787422 }}</ref>
|
Gen. et sp. nov.
|
Valid
|
Smith ''et al.''
|
Late Cretaceous (Campanian)
|
|
{{Flag|Canada}}
|
A [[Biology:Ranunculales|ranunculalean]] known from a stem with [[Biology:Liana|lianoid]] anatomy. The type specimen is ''Atli morinii''.
| 
|-
|
''Bambusiculmus''<ref name=RPP197bamboos>{{cite journal |author1=Li Wang |author2=Frédéric M.B. Jacques |author3=Tao Su |author4=Yaowu Xing |author5=Shitao Zhang |author6=Zhekun Zhou |year=2013 |title=The earliest fossil bamboos of China (middle Miocene, Yunnan) and their biogeographical importance |journal=Review of Palaeobotany and Palynology |volume=197 |pages=253–265 |doi=10.1016/j.revpalbo.2013.06.004 |bibcode=2013RPaPa.197..253W }}</ref>
|
Gen. et 2 sp. nov.
|
Valid
|
Wang & Zhou ''in'' Wang ''et al.''
|
Miocene
|
Dajie Formation
|
{{Flag|China}}
|
A [[Biology:Bamboo|bamboo]]. The type specimen is ''Bambusiculmus angustus''; genus also includes ''Bambusiculmus latus''.
|
|-
|
''Bambusium angustifolia''<ref name=RPP197bamboos />
|
Sp. nov.
|
Valid
|
Wang & Zhou ''in'' Wang ''et al.''
|
Miocene
|
Dajie Formation
|
{{Flag|China}}
|
A [[Biology:Bamboo|bamboo]], a species of ''Bambusium''.
|
|-
|
''Bambusium latifolia''<ref name=RPP197bamboos />
|
Sp. nov.
|
Valid
|
Wang & Zhou ''in'' Wang ''et al.''
|
Miocene
|
Dajie Formation
|
{{Flag|China}}
|
A [[Biology:Bamboo|bamboo]], a species of ''Bambusium''.
|
|-
|
''Bombacacioxylon''<ref>{{cite journal |author1=Rakesh Chandra Mehrotra |author2=R. P. Tiwari |author3=Gaurav Srivastava |author4=Anumeha Shukla |year=2013 |title=Further contribution to the Neogene petrified wood forest of Mizoram, India |journal=Chinese Science Bulletin |volume=58 |issue=Supplement I |pages=104–110 |doi=10.1360/tb-2013-suppl006 |url=https://www.academia.edu/7903479 }}</ref>
|
Gen. et sp. nov.
|
Valid
|
Mehrotra ''et al.''
|
Late Miocene to early Pliocene
|
Tipam Group
|
{{Flag|India}}
|
A possible relative of ''[[Biology:Bombax|Bombax]]''. The type specimen is ''Bombacacioxylon tertiarum''.
| 
|-
|
''Carpolithus warmanensis''<ref name=Lauroflorum />
|
Sp. nov.
|
Valid
|
Wang, Blanchard & Dilcher
|
Eocene
|
Cockfield Formation
|
{{Flag|United States}}
|
A fossil fruit.
|
|-
|
''Cornerocarpon''<ref name=Lauroflorum />
|
Gen. et sp. nov.
|
Valid
|
Grote ''in'' Wang, Blanchard & Dilcher
|
Eocene
|
Cockfield Formation
|
{{Flag|United States}}
|
A member of [[Biology:Rosales|Rosales]] belonging or related to the family [[Biology:Moraceae|Moraceae]]. Genus includes new species ''C. copiosum''.
|
|-
|
''Corylopsis grotei''<ref name=Lauroflorum />
|
Sp. nov.
|
Valid
|
Wang, Blanchard & Dilcher
|
Eocene
|
Cockfield Formation
|
{{Flag|United States}}
|
A species of ''[[Biology:Corylopsis|Corylopsis]]''.
|
|-
|
''Diospyros neyveliensis''<ref>{{cite journal |author1=Debi Mukherjee |author2=Mahesh Prasad |year=2013 |title=An ebenaceous fossil wood from the Neyveli lignite, South Arcot District, Tamil Nadu, India |journal=Geophytology |volume=42 |issue=2 |pages=127–133 }}</ref>
|
Sp. nov.
|
Valid
|
Mukherjee & Prasad
|
Miocene
|
Neyveli Formation
|
{{Flag|India}}
|
A member of the family [[Biology:Ebenaceae|Ebenaceae]] described on the basis of [[Biology:Fossil wood|fossil wood]]. Originally described as a species of ''[[Biology:Diospyros|Diospyros]]''; Srivastava ''et al.'' (2018) transferred this species to the genus ''Ebenoxylon''.<ref>{{Cite journal|author1=Gaurav Srivastava |author2=R. C. Mehrotra |author3=C. Srikarni |year=2018 |title=Fossil wood flora from the Siwalik Group of Arunachal Pradesh, India and its climatic and phytogeographic significance |journal=Journal of Earth System Science |volume=127 |issue=1 |pages=Article 2 |doi=10.1007/s12040-017-0903-2 |bibcode=2018JESS..127....2S |s2cid=134314695 |doi-access=free }}</ref>
|
|-
|
''Firmiana oligocenica''<ref name=JESS1225>{{cite journal |author1=Gaurav Srivastava |author2=Rakesh Chandra Mehrotra |year=2013 |title=Further contribution to the low latitude leaf assemblage from the late Oligocene sediments of Assam and its phytogeographical significance |journal=Journal of Earth System Science |volume=122 |issue=5 |pages=1341–1357 |doi=10.1007/s12040-013-0354-3 |bibcode=2013JESS..122.1341S |s2cid=128613123 |doi-access=free }}</ref>
|
Sp. nov.
|
Valid
|
Srivastava & Mehrotra
|
Oligocene ([[Earth:Chattian|Chattian]])
|
Tikak Parbat Formation
|
{{Flag|India}}
|
A species of ''[[Biology:Firmiana|Firmiana]]''.
| 
|-
|
''Grotea''<ref name=Lauroflorum />
|
Gen. et sp. nov.
|
Valid
|
Wang, Blanchard & Dilcher
|
Eocene
|
Cockfield Formation
|
{{Flag|United States}}
|
A member of the family [[Biology:Bignoniaceae|Bignoniaceae]]. Genus includes new species ''G. warmanensis''.
|
|-
|
''Guadua morronei''<ref>{{cite journal |author1=Mariana Brea |author2=Alejandro F. Zucol |author3=María Jimena Franco |year=2013 |title=A new Bambusoideae (Poaceae: Bambusoideae: Bambuseae: Guaduinae) from the Ituzaingó Formation (Pliocene–Pleistocene), Entre Ríos, Argentina |journal=Review of Palaeobotany and Palynology |volume=192 |pages=1–9 |doi=10.1016/j.revpalbo.2012.12.006 |bibcode=2013RPaPa.192....1B |hdl=11336/78560 |hdl-access=free }}</ref>
|
Sp. nov.
|
Valid
|
Brea, Zucol & Franco
|
Pliocene–Pleistocene
|
[[Earth:Ituzaingó Formation|Ituzaingó Formation]]
|
{{Flag|Argentina}}
|
A [[Biology:Bamboo|bamboo]], a species of ''[[Biology:Guadua|Guadua]]''.
|
|-
|
''Hamamelis warmanensis''<ref name=Lauroflorum />
|
Sp. nov.
|
Valid
|
Wang, Blanchard & Dilcher
|
Eocene
|
Cockfield Formation
|
{{Flag|United States}}
|
A [[Biology:Witch-hazel|witch-hazel]].
|
|-
|
''Indovitis''<ref>{{cite journal |author1=Steven R. Manchester |author2=Dashrath K. Kapgate |author3=Jun Wen |year=2013 |title=Oldest fruits of the grape family (Vitaceae) from the Late Cretaceous Deccan Cherts of India |journal=American Journal of Botany |volume=100 |issue=9 |pages=1849–1859 |doi=10.3732/ajb.1300008 |pmid=24036414}}</ref>
|
Gen. et sp. nov.
|
Valid
|
Manchester, Kapgate & Wen
|
Late Cretaceous ([[Earth:Maastrichtian|Maastrichtian]])
|
[[Earth:Deccan Traps|Deccan]] [[Earth:Intertrappean Beds|Intertrappean Beds]]
|
{{Flag|India}}
|
A member of the family [[Biology:Vitaceae|Vitaceae]]. Genus includes new species ''I. chitaleyae''.
| 
|-
|
''Iterophyllum''<ref>{{cite journal |author1=Abel Barral |author2=Bernard Gomez |author3=Taylor S. Feild |author4=Clément Coiffard |author5=Véronique Daviero-Gomez |year=2013 |title=Leaf architecture and ecophysiology of an early basal eudicot from the Early Cretaceous of Spain |journal=Botanical Journal of the Linnean Society |volume=173 |issue=4 |pages=594–605 |doi=10.1111/boj.12095 |doi-access=free }}</ref>
|
Gen. et sp. nov.
|
Valid
|
Barral ''et al.''
|
Early Cretaceous (late Barremian)
|
|
{{Flag|Spain}}
|
A basal [[Biology:Eudicots|eudicot]]. The type specimen is ''Iterophyllum lobatum''.
| 
|-
|
''Jaguariba wiersemana''<ref name="Coiffard2013">{{cite journal |last1=Coiffard |first1=C. |last2=Mohr |first2=B.A.R. |last3=Bernardes-de-Oliveira |first3=M.E.C. |year=2013 |title=''Jaguariba wiersemana'' gen. nov. et sp. nov., an Early Cretaceous member of crown group Nymphaeales (Nymphaeaceae) from northern Gondwana |journal= Taxon |volume=62 |issue=1 |pages=141–151 |doi=10.1002/tax.621012 }}</ref>
|
Gen. nov. et sp. nov.
|
Valid
|
Coiffard, Mohr & Bernardes-de-Oliveira
|
Early Cretaceous
|
[[Earth:Crato Formation|Crato Formation]]
|
{{Flag|Brazil}}
|
Fossil [[Biology:Nymphaeales|Nymphaeales]].
| 
|-
|
''Lagerstroemia corvinusii''<ref name=HimalayanGeology>{{cite journal |author=Mahesh Prasad |year=2013 |title=Record of leaf impressions from Middle Churia Formation of Arjun Khola area in the Sub-Himalayan zone of Nepal and their palaeoclimatic and palaeophytogeographical implications |journal=Himalayan Geology |volume=34 |issue=2 |pages=158–167 |url=https://www.researchgate.net/publication/286341115 }}</ref>
|
Sp. nov.
|
Valid
|
Prasad
|
Mio-Pliocene
|
Churia Formation
|
{{Flag|Nepal}}
|
A species of ''[[Biology:Lagerstroemia|Lagerstroemia]]''.
|
|-
|
''Laurelia otagoensis''<ref>{{Cite journal|author1=John G. Conran |author2=Jennifer M. Bannister |author3=Daphne E. Lee |year=2013 |title=Fruits and leaves with cuticle of ''Laurelia otagoensis'' sp. nov. (Atherospermataceae) from the early Miocene of Otago (New Zealand) |journal=Alcheringa: An Australasian Journal of Palaeontology |volume=37 |issue=4 |pages=496–509 |doi=10.1080/03115518.2013.798765 |bibcode=2013Alch...37..496C |s2cid=140637946 }}</ref>
|
Sp. nov
|
Valid
|
Conran, Bannister & Lee
|
Early Miocene
|
|
{{Flag|New Zealand}}
|
A species of ''[[Biology:Laurelia|Laurelia]]''.
|
|-
|
''Lauroflorum''<ref name=Lauroflorum />
|
Gen. et sp. nov.
|
Valid
|
Wang, Blanchard & Dilcher
|
Eocene
|
Cockfield Formation
|
{{Flag|United States}}
|
A member of [[Biology:Lauraceae|Lauraceae]]. Genus includes new species ''L. warmanensis''.
| 
|-
|
''Lindera acuminatissima''<ref>{{Cite journal|author1=Kequn Dao |author2=Junlin Chen |author3=Peihong Jin |author4=Chong Dong |author5=Yi Yang |author6=Xiaohui Xu |author7=Jingyu Wu |author8=Sanping Xie |author9=Zhicheng Lin |author10=Bainian Sun |year=2013 |title=A New Material of ''Lindera'' (Lauraceae) of the Late Pliocene from Tengchong, Yunnan and the Genus' Biogeography Significance |journal=Acta Geologica Sinica (English Edition) |volume=87 |issue=3 |pages=690–706 |url=http://www.geojournals.cn/dzxben/ch/reader/view_abstract.aspx?file_no=201303007&flag=1 |doi=10.1111/1755-6724.12081 |bibcode=2013AcGlS..87..690D |s2cid=129635617 }}</ref>
|
Sp. nov
|
Valid
|
Dao & Sun ''in'' Dao ''et al.''
|
Late Pliocene
|
Mangbang Formation
|
{{Flag|China}}
|
A species of ''[[Biology:Lindera|Lindera]]''.
|
|-
|
''"Magnolia" szakmanycsabae''<ref name=GH59 />
|
Sp. nov.
|
Valid
|
Hably
|
Late Miocene
|
|
{{Flag|Hungary}}
|
A plant of uncertain phylogenetic placement, possibly a relative of [[Biology:Magnoliaceae|Magnoliaceae]]–[[Biology:Lauraceae|Lauraceae]].
|
|-
|
''Micropetasos''<ref>{{Cite journal |author1=George O. Poinar, Jr. |author2=Kenton L. Chambers |author3=Joerg Wunderlich |year=2013 |title=''Micropetasos'', a new genus of angiosperms from mid-cretaceous Burmese amber |journal=Journal of the Botanical Research Institute of Texas |volume=7 |issue=2 |pages=745–750 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Poinar, Chambers & Wunderlich
|
[[Earth:Late Cretaceous|Late Cretaceous]] ([[Earth:Cenomanian|Cenomanian]])
|
[[Earth:Burmese amber|Burmese amber]]
|
{{Flag|Myanmar}}
|
An early flowering plant of uncertain phylogenetic placement. Genus includes new species ''M. burmensis''.
|
|-
|
''Myrtineoxylon''<ref>{{cite journal |author1=Alexei A. Oskolski |author2=Xin-Xin Feng |author3=Jian-Hua Jin |year=2013 |title=''Myrtineoxylon'' gen. nov.: The first fossil wood record of the tribe Myrteae (Myrtaceae) in eastern Asia |journal=Taxon |volume=62 |issue=4 |pages=771–778 |doi=10.12705/624.7 }}</ref>
|
Gen. et sp. nov.
|
Valid
|
Oskolski, Feng & Jin
|
Late [[Earth:Eocene|Eocene]]
|
Youganwo Formation
|
{{Flag|China}}
|
A member of [[Biology:Myrtaceae|Myrtaceae]] described on the basis of fossil wood. Genus includes new species ''M. maomingensis''.
|
|-
|
''Palmoxylon vastanensis''<ref>{{cite journal |author1=M. Prasad |author2=H. Singh |author3=S. K. Singh |author4=Debi Mukherjee |author5=Emilio Estrada Ruiz |year=2013 |title=Early Eocene arecoid palm wood, ''Palmoxylon vastanensis'' n. sp. from Vastan Lignite, Gujarat, India: its palaeoenvironmental implications |journal=Journal of the Palaeontological Society of India |volume=58 |issue=1 |pages=115–123 |url=http://palaeontologicalsociety.in/aug14/vol58_1/12.pdf }}</ref>
|
Sp. nov.
|
Valid
|
Prasad ''et al.''
|
Eocene
|
Cambay Shale Formation
|
{{Flag|India}}
|
A [[Biology:Arecaceae|palm tree]], a species of ''[[Biology:Palmoxylon|Palmoxylon]]''.
| 
|-
|
''Paranephelium makumensis''<ref name=JESS1225 />
|
Sp. nov.
|
Valid
|
Srivastava & Mehrotra
|
Oligocene ([[Earth:Chattian|Chattian]])
|
Tikak Parbat Formation
|
{{Flag|India}}
|
A species of ''[[Biology:Paranephelium|Paranephelium]]''.
| 
|-
|
''Poeciloneuron preindicum''<ref>{{cite journal |author1=Gaurav Srivastava |author2=R C Mehrotra |year=2013 |title=Endemism due to climate change: Evidence from ''Poeciloneuron'' Bedd. (Clusiaceae) leaf fossil from Assam, India |journal=Journal of Earth System Science |volume=122 |issue=2 |pages=283–288 |doi=10.1007/s12040-013-0277-z |bibcode=2013JESS..122..283S |s2cid=67769307 |doi-access=free }}</ref>
|
Sp. nov.
|
Valid
|
Srivastava & Mehrotra
|
Late Oligocene
|
|
{{Flag|India}}
|
A species of ''[[Biology:Poeciloneuron|Poeciloneuron]]''.
| 
|-
|
''Pongamia arjunkholaensis''<ref name=HimalayanGeology />
|
Sp. nov.
|
Valid
|
Prasad
|
Mio-Pliocene
|
Churia Formation
|
{{Flag|Nepal}}
|
A species of ''[[Biology:Pongamia|Pongamia]]''.
|
|-
|
''Potomacapnos''<ref>{{cite journal |author1=Nathan A. Jud |author2=Leo J. Hickey |year=2013 |title=''Potomacapnos apeleutheron'' gen. et sp. nov., a new Early Cretaceous angiosperm from the Potomac Group and its implications for the evolution of eudicot leaf architecture |journal=American Journal of Botany |volume=100 |issue=12 |pages=2437–2449 |doi=10.3732/ajb.1300250 |pmid=24287268}}</ref>
|
Gen. et sp. nov.
|
Valid
|
Jud & Hickey
|
Early Cretaceous (Aptian)
|
Potomac Group
|
{{Flag|United States}}
|
A close relative of ranunculalean [[Biology:Eudicots|eudicots]]. The type specimen is ''Potomacapnos apeleutheron''.
| 
|-
|
''Pterygota palaeoalata''<ref name=JESS1225 />
|
Sp. nov.
|
Valid
|
Srivastava & Mehrotra
|
Oligocene ([[Earth:Chattian|Chattian]])
|
Tikak Parbat Formation
|
{{Flag|India}}
|
A species of ''Pterygota''.
| 
|-
|
''Sapindus palaeoemarginatus''<ref name=JESS1225 />
|
Sp. nov.
|
Valid
|
Srivastava & Mehrotra
|
Oligocene ([[Earth:Chattian|Chattian]])
|
Tikak Parbat Formation
|
{{Flag|India}}
|
A species of ''[[Biology:Sapindus|Sapindus]]''.
| 
|-
|
''Spixiarum kipea''<ref name="Coiffard2013b">{{cite journal |last1=Coiffard |first1=C. |last2=Mohr |first2=B.A.R. |last3=Bernardes-de-Oliveira |first3=M.E.C. |year=2013 |title=The Early Cretaceous Aroid, ''Spixiarum kipea'' gen. et sp nov. and implications on early dispersal and ecology of basal monocots |journal= Taxon |volume=62 |issue=5 |pages=997–1008 |doi=10.12705/625.21|doi-access=free }}</ref>
|
Gen. nov. et sp. nov.
|
Valid
|
Coiffard, Mohr & Bernardes-de-Oliveira
|
Early Cretaceous
|
[[Earth:Crato Formation|Crato Formation]]
|
{{Flag|Brazil}}
|
Fossil Aroid.
| 
|-
|
''Trapa pannonica''<ref name=GH59>{{cite journal |author=Lilla Hably |year=2013 |title=The Late Miocene Flora of Hungary |journal=Geologica Hungarica Series Palaeontologica |volume=59 |pages=1–175 |url=https://epa.oszk.hu/02900/02989/00044/pdf/EPA02989_geologica_hungarica_ser_paleo_59_2013.pdf }}</ref>
|
Sp. nov.
|
Valid
|
Hably
|
Late Miocene
|
|
{{Flag|Hungary}}
|
A [[Biology:Water caltrop|water caltrop]].
|
|-
|
''Trapa rozsaszentmartoni''<ref name=GH59 />
|
Sp. nov.
|
Valid
|
Hably
|
Late Miocene
|
|
{{Flag|Hungary}}
|
A [[Biology:Water caltrop|water caltrop]].
|
|-
|
''Virola dominicana''<ref>{{cite journal |author1=George Poinar Jr. |author2=Royce Steeves |year=2013 |title=''Virola dominicana'' sp. nov. (Myristicaceae) from Dominican amber |journal=Botany |volume=91 |issue=8 |pages=530–534 |doi=10.1139/cjb-2013-0019 }}</ref>
|
Sp. nov.
|
Valid
|
Poinar & Steeves
|
Cenozoic (Eocene or Miocene)
|
|
{{Flag|Dominican Republic}}
|
A member of [[Biology:Myristicaceae|Myristicaceae]] found in [[Earth:Dominican amber|Dominican amber]], a species of ''[[Biology:Virola|Virola]]''.
| 
|-
|
''Vitis szakmanygyorgyi''<ref name=GH59 />
|
Sp. nov.
|
Valid
|
Hably
|
Late Miocene
|
|
{{Flag|Hungary}}
|
A species of ''[[Biology:Vitis|Vitis]]''.
|
|-
|}

==Other seed plants==
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Unit
! Location
! Notes
! Images
|-
|
''Donponoxylon''<ref>{{cite journal |author1=William D. Tidwell |author2=Brooks B. Britt |author3=W. Walton Wright |year=2013 |title=''Donponoxylon'' gen. nov., a new spermatophyte axis from the Middle to Late Jurassic of Australia and New Zealand |journal=Review of Palaeobotany and Palynology |volume=196 |pages=36–50 |doi=10.1016/j.revpalbo.2013.03.008 |bibcode=2013RPaPa.196...36T }}</ref>
|
Gen. et 2 sp. nov.
|
Valid
|
Tidwell, Britt & Wright
|
Middle to Late Jurassic
|
|
{{Flag|Australia}}<br>
{{Flag|New Zealand}}
|
A [[Biology:Spermatophyte|seed plant]] of uncertain phylogenetic placement. The type specimen is ''Donponoxylon bennettii''; genus also contains ''Donponoxylon jacksonii''.
| 
|-
|
''Dordrechtites arcanus''<ref>{{cite journal |author1=Julie A. Bergene |author2=Edith L. Taylor |author3=Thomas N. Taylor |year=2013 |title=''Dordrechtites arcanus'' sp. nov., an Anatomically Preserved Gymnospermous Reproductive Structure from the Middle Triassic of Antarctica |journal=International Journal of Plant Sciences |volume=174 |issue=2 |pages=250–265 |doi=10.1086/668792 |hdl=1808/13678 |s2cid=73561042 |hdl-access=free }}</ref>
|
Sp. nov.
|
Valid
|
Bergene, Taylor & Taylor 
|
Middle Triassic (Anisian)
|
[[Earth:Fremouw Formation|Fremouw Formation]]
|
{{Flag|Antarctica}}
|
A [[Biology:Gymnosperm|gymnosperm]] of uncertain phylogenetic placement.
| 
|-
|}

==Other plants==
{| class="wikitable sortable" align="center" width="100%"
|-
! Name
! Novelty
! Status
! Authors
! Age
! Unit
! Location
! Notes
! Images
|-
|
''Acinosporites tristratus''<ref name=SPP89>{{cite journal |author1=Pierre Breuer |author2=Philippe Steemans |title=Devonian spore assemblages from northwestern Gondwana: taxonomy and biostratigraphy |journal=Special Papers in Palaeontology |volume=89 |pages=5–163 |doi=10.1111/pala.12031 |doi-broken-date=31 January 2024 | url = https://www.palass.org/publications/special-papers-palaeontology/archive/89/article_pp1-163}}</ref>
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jubah Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Alatisporites? trisacculus''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Biornatispora elegantula''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation
|
{{Flag|Belgium}}<br>
{{Flag|Germany}}<br>
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Biornatispora microclavata''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Brochotriletes crameri''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation<br>
Jubah Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Brochotriletes tenellus''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Brochotriletes tripapillatus''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Awaynat Wanin II Formation
|
{{Flag|Libya}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Camarozonotriletes asperulus''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Awaynat Wanin I Formation<br>
Awaynat Wanin II Formation<br>
Awaynat Wanin III Formation
|
{{Flag|Brazil}}<br>
{{Flag|Tunisia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Chelinospora carnosa''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Chelinospora condensata''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Chelinospora densa''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Chelinospora laxa''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Chelinospora vulgata''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Coronaspora inornata''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Cristatisporites streelii''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Awaynat Wanin II Formation
|
{{Flag|Brazil}}<br>
{{Flag|Tunisia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Cymbosporites variabilis''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Cymbosporites variegatus''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Awaynat Wanin II Formation
|
{{Flag|Brazil}}<br>
{{Flag|Libya}}<br>
{{Flag|Saudi Arabia}}<br>
{{Flag|Tunisia}}<br>
{{Flag|United Kingdom}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Cymbosporites wellmanii''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Cyrtospora tumida''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Awaynat Wanin I Formation<br>
Awaynat Wanin II Formation<br>
Awaynat Wanin III Formation
|
{{Flag|Saudi Arabia}}<br>
{{Flag|Tunisia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Diaphanospora milleri''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation<br>
Ouan Kasa Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Dibolisporites tuberculatus''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Awaynat Wanin I Formation<br>
Awaynat Wanin II Formation<br>
Jauf Formation<br>
Jubah Formation
|
{{Flag|Brazil}}<br>
{{Flag|Libya}}<br>
{{Flag|Saudi Arabia}}<br>
{{Flag|Spain}}<ref name=PPAskewWellman /><br>
{{Flag|Tunisia}}<br>
{{Flag|United States}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Dibolisporites verecundus''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Dictyotriletes hemeri''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jubah Formation
|
{{Flag|Saudi Arabia}}<br>
{{Flag|Spain}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Dictyotriletes marshallii''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation<br>
Ouan Kasa Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Elenisporis gondwanensis''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Awaynat Wanin I Formation<br>
Awaynat Wanin II Formation<br>
Jubah Formation<br>
|
{{Flag|Saudi Arabia}}<br>
{{Flag|Tunisia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Emphanisporites laticostatus''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Awaynat Wanin III Formation<br>
Jubah Formation<br>
|
{{Flag|Brazil}}<br>
{{Flag|Saudi Arabia}}<br>
{{Flag|Tunisia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Emphanisporites plicatus''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation<br>
Jubah Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''[[Biology:Franhueberia|Franhueberia]]''<ref>{{cite journal |author1=Laurel A. Hoffman |author2=Alexandru M. F. Tomescu |year=2013 |title=An early origin of secondary growth: ''Franhueberia gerriennei'' gen. et sp. nov. from the Lower Devonian of Gaspé (Quebec, Canada) |journal=American Journal of Botany |volume=100 |issue=4 |pages=754–763 |doi=10.3732/ajb.1300024 |pmid=23535772}}</ref>
|
Gen. et sp. nov.
|
Valid
|
Hoffman & Tomescu
|
Devonian ([[Earth:Emsian|Emsian]])
|
Battery Point Formation
|
{{Flag|Canada}}
|
A basal member of Euphyllophytina. The type specimen is ''F. gerriennei''.
| 
|-
|
''Geminospora convoluta''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Awaynat Wanin II Formation<br>
Jubah Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Grandispora maura''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Awaynat Wanin I Formation<br>
Awaynat Wanin II Formation<br>
Awaynat Wanin III Formation
|
{{Flag|Saudi Arabia}}<br>
{{Flag|Tunisia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Granulatisporites concavus''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Awaynat Wanin I Formation<br>
Awaynat Wanin II Formation<br>
Jauf Formation<br>
Jubah Formation
|
{{Flag|Libya}}<br>
{{Flag|Saudi Arabia}}<br>
{{Flag|Spain}}<ref name=PPAskewWellman />
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Hystricosporites brevispinus''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Awaynat Wanin II Formation
|
{{Flag|Tunisia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Raistrickia commutata''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Awaynat Wanin II Formation<br>
Awaynat Wanin III Formation
|
{{Flag|Saudi Arabia}}<br>
{{Flag|Tunisia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Raistrickia jaufensis''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Retusotriletes atratus''<ref name=SPP89 />
|
Sp. nov
| 
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation<br>
Jubah Formation
|
{{Flag|Australia}}<br>
{{Flag|Saudi Arabia}}<br>
{{Flag|Spain}}<ref name=PPAskewWellman>{{Cite journal|author1=Alexander J. Askew |author2=Charles H. Wellman |year=2019 |title=An endemic flora of dispersed spores from the Middle Devonian of Iberia |journal=Papers in Palaeontology |volume=5 |issue=3 |pages=415–459 |doi=10.1002/spp2.1245 |bibcode=2019PPal....5..415A |doi-access=free }}</ref>
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Retusotriletes celatus''<ref name=SPP89 />
|
Sp. nov
|
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Samarisporites tunisiensis''<ref name=SPP89 />
|
Sp. nov
|
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Ouan Kasa Formation
|
{{Flag|Tunisia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Verrucosisporites nafudensis''<ref name=SPP89 />
|
Sp. nov
|
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Verrucosisporites onustus''<ref name=SPP89 />
|
Sp. nov
|
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Verrucosisporites stictus''<ref name=SPP89 />
|
Sp. nov
|
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation<br>
Ouan Kasa Formation
|
{{Flag|Saudi Arabia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Zonotriletes brevivelatus''<ref name=SPP89 />
|
Sp. nov
|
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Awaynat Wanin I Formation<br>
Jauf Formation<br>
Jubah Formation<br>
Ouan Kasa Formation
|
{{Flag|Algeria}}<br>
{{Flag|Libya}}<br>
{{Flag|Saudi Arabia}}<br>
{{Flag|Tunisia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Zonotriletes rotundus''<ref name=SPP89 />
|
Sp. nov
|
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Awaynat Wanin I Formation<br>
Awaynat Wanin II Formation<br>
Ouan Kasa Formation
|
{{Flag|Brazil}}<br>
{{Flag|Tunisia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|
''Zonotriletes venatus''<ref name=SPP89 />
|
Sp. nov
|
Valid
|
Breuer & Steemans
|
[[Earth:Devonian|Devonian]]
|
Jauf Formation<br>
Ouan Kasa Formation
|
{{Flag|Algeria}}<br>
{{Flag|Brazil}}<br>
{{Flag|Saudi Arabia}}<br>
{{Flag|Tunisia}}
|
A [[Biology:Spore#Anatomy|trilete spore]].
|
|-
|}

==References==
{{Reflist|30em}}

[[Category:Paleobotany]]

{{Sourceattribution|2013 in paleobotany}}