View source for Biology:2021 in paleobotany

This article records new [[Biology:Taxon|taxa]] of fossil [[Biology:Plant|plant]]s that are scheduled to be [[Biology:Binomial nomenclature|described]] during the year 2021, as well as other significant discoveries and events related to [[Biology:Paleobotany|paleobotany]] that are scheduled to occur in the year 2021.

==Ferns and fern allies==
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
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''[[Biology:Arthropitys|Arthropitys buritiranensis]]''<ref>{{Cite journal|last1=Neregato |first1=R. |last2=Rößler |first2=R. |last3=Noll |first3=R. |last4=Rohn |first4=R. |year=2021 |title=New petrified calamitaleans from the Permian of the Parnaíba Basin, central-north Brazil, part III, with some concerns regarding anatomical features of Paleozoic petrified sphenophytes |journal=Review of Palaeobotany and Palynology |volume=293 |pages=Article 104499 |doi=10.1016/j.revpalbo.2021.104499 |bibcode=2021RPaPa.29304499N |doi-access=free }}</ref>
|
Sp. nov
|
In press
|
Neregato ''et al.''
|
[[Earth:Permian|Permian]]
|
Motuca Formation
|
{{Flag|Brazil}}
|
A member of [[Biology:Calamitales|Calamitales]].
|
|-
|
''Azolla andreisii''<ref>{{Cite journal|last1=De Benedetti |first1=F. |last2=Zamaloa |first2=M. C. |last3=Gandolfo |first3=M. A. |last4=Cúneo |first4=N. R. |year=2021 |title=Water fern spores (Salviniales) from the Late Cretaceous of Patagonia, Argentina |journal=Review of Palaeobotany and Palynology |volume=290 |pages=Article 104428 |doi=10.1016/j.revpalbo.2021.104428 |bibcode=2021RPaPa.29004428D |s2cid=234815591 |doi-access=free }}</ref>
|
Sp. nov
|
In press
|
De Benedetti ''et al.''
|
[[Earth:Late Cretaceous|Late Cretaceous]] ([[Earth:Maastrichtian|Maastrichtian]])
|
[[Earth:La Colonia Formation|La Colonia Formation]]
|
{{Flag|Argentina}}
|
A species of ''[[Biology:Azolla|Azolla]]''.
|
|-
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''Caulopteris ellipticus''<ref name=RPPPsaroniaceae>{{Cite journal|last1=Wang |first1=S.-J. |last2=Wang |first2=J. |last3=Liu |first3=L. |last4=Hilton |first4=J. |year=2021 |title=Stem diversity of the marattialean tree fern family Psaroniaceae from the earliest Permian Wuda Tuff Flora |journal=Review of Palaeobotany and Palynology |volume=294 |pages=Article 104378 |doi=10.1016/j.revpalbo.2021.104378 |bibcode=2021RPaPa.29404378W |s2cid=234276315 }}</ref>
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Sp. nov
|
In press
|
Wang ''et al.''
|
Early [[Earth:Permian|Permian]]
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Taiyuan Formation
|
{{Flag|China}}
|
A [[Biology:Marattiales|marattialean]] tree fern belonging to the family Psaroniaceae.
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|-
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''Caulopteris neimengensis''<ref name=RPPPsaroniaceae />
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Sp. nov
|
In press
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Wang ''et al.''
|
Early Permian
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Taiyuan Formation
|
{{Flag|China}}
|
A marattialean tree fern belonging to the family Psaroniaceae.
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|-
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''Caulopteris obovatus''<ref name=RPPPsaroniaceae />
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Sp. nov
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In press
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Wang ''et al.''
|
Early Permian
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Taiyuan Formation
|
{{Flag|China}}
|
A marattialean tree fern belonging to the family Psaroniaceae.
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|-
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''Cicatricosisporites pseudograndiosus''<ref name=Passifloriidites />
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Sp. nov
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Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Spores of a member of the genus ''[[Biology:Ceratopteris|Ceratopteris]]''.
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|-
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''Cladarastega''<ref>{{Cite journal|last=Poinar |first=G. |title=A new fern, ''Cladarastega burmanica'' gen. et sp. nov. (Dennstaedtiaceae: Polypodiales) in mid-Cretaceous Burmese amber |year=2021 |journal=Palaeodiversity |volume=14 |issue=1 |pages=153–160 |doi=10.18476/pale.v14.a7 |doi-access=free }}</ref>
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Gen. et sp. nov
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Valid
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Poinar
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Late Cretaceous (Cenomanian)
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Burmese amber
|
{{Flag|Myanmar}}
|
A fern belonging to the family [[Biology:Dennstaedtiaceae|Dennstaedtiaceae]]. Genus includes new species ''C. burmanica''.
|
|-
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''Claytosmunda zhangii''<ref>{{Cite journal|last1=Tian |first1=N. |last2=Wang |first2=Y.-D. |last3=Jiang |first3=Z.-K. |year=2021 |title=A new permineralized osmundaceous rhizome with fungal remains from the Jurassic of western Liaoning, NE China |journal=Review of Palaeobotany and Palynology |volume=290 |pages=Article 104414 |doi=10.1016/j.revpalbo.2021.104414 |bibcode=2021RPaPa.29004414T | issn=0034-6667  |s2cid=233549860 }}</ref>
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Sp. nov
|
In press
|
Tian, Wang & Jiang
|
Late Jurassic
|
[[Earth:Tiaojishan Formation|Tiaojishan Formation]]
|
{{Flag|China}}
|
A fern, a species of ''[[Biology:Claytosmunda|Claytosmunda]]''.
|
|-
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''[[Earth:Dennstaedtia christophelii|Dennstaedtia christophelii]]''<ref name=IJPSPiggetal>{{Cite journal|last1=Pigg |first1=K. B. |last2=DeVore |first2=M. L. |last3=Greenwood |first3=D. R. |last4=Sundue |first4=M. A. |last5=Schwartsburd |first5=P. |last6=Basinger |first6=J. F. |year=2021 |title=Fossil Dennstaedtiaceae and Hymenophyllaceae from the Early Eocene of the Pacific Northwest |journal=International Journal of Plant Sciences |volume=182 |issue=9 |pages=793–807 |doi=10.1086/715633 |s2cid=239036762 }}</ref>
|
Sp. nov
|
Valid
|
Pigg ''et al.''
|
Early Eocene
|
[[Earth:Klondike Mountain Formation|Klondike Mountain Formation]]
|
{{Flag|United States}}<br>({{Flag|Washington}})
|
A fern, a species of ''[[Biology:Dennstaedtia|Dennstaedtia]]''.
|
[[File:Dennstaedtia christophelii holotype SRIC SR 13-004-001 A img1.tif|thumb|center|upright|''Dennstaedtia christophelii'']]
|-
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''Echinosporis densiechinatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Spores of a member of the family [[Biology:Marattiaceae|Marattiaceae]].
|
|-
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''Eoangiopteris congestus''<ref>{{Cite journal|last1=Sun |first1=W. |last2=Li |first2=D. |last3=Zhou |first3=W. |last4=Bek |first4=J. |last5=Liu |first5=L. |last6=Wang |first6=J. |year=2021 |title=''Eoangiopteris congestus'' sp. nov., a marattialean fern from the Lower Permian Taiyuan Formation of Wuda Coalfield, Inner Mongolia |journal=Acta Palaeontologica Sinica |volume=60 |issue=2 |pages=224–242 |doi=10.19800/j.cnki.aps.2020012 |url=http://gswxb.cnjournals.cn/gswxb/article/abstract/20210203 }}</ref>
|
Sp. nov
|
Valid
|
Sun ''et al.''
|
Early [[Earth:Permian|Permian]]
|
Taiyuan Formation
|
{{Flag|China}}
|
A fern belonging to the group [[Biology:Marattiales|Marattiales]].
|
|-
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''Hymenophyllum axsmithii''<ref name=IJPSPiggetal />
|
Sp. nov
|
Valid
|
Pigg ''et al.''
|
Early Eocene
|
Klondike Mountain Formation
|
{{Flag|United States}}<br>({{Flag|Washington}})
|
A fern, a species of ''[[Biology:Hymenophyllum|Hymenophyllum]]''.
|
|-
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''Iberisetum''<ref>{{Cite journal|last1=Correia |first1=P. |last2=Šimůnek |first2=Z. |last3=Sá |first3=A. A. |year=2021 |title=The equisetalean ''Iberisetum wegeneri'' gen. nov., sp. nov. from the Upper Pennsylvanian of Portugal |journal=Historical Biology: An International Journal of Paleobiology |volume=33 |issue=12 |pages=3495–3505 |doi=10.1080/08912963.2021.1874373 |s2cid=234064743 }}</ref>
|
Gen. et sp. nov
|
In press
|
Correia, Šimůnek & Sá
|
[[Earth:Carboniferous|Carboniferous]] (Gzhelian)
|
Douro Basin
|
{{Flag|Portugal}}
|
A member of [[Biology:Equisetales|Equisetales]]. Genus includes new species ''I. wegeneri''.
|
|-
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''Laevigatosporites cultellus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Spores of a member of the family [[Biology:Polypodiaceae|Polypodiaceae]].
|
|-
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''Marsileaceaephyllum ciliatum''<ref>{{Cite journal|last1=Wang |first1=S. |last2=Long |first2=X. |last3=Zhang |first3=H. |last4=Cai |first4=H. |last5=Engel |first5=M. S. |last6=Shi |first6=C. |year=2021 |title=A semi-aquatic fern (Marsileaceae) from the mid-Cretaceous amber of northern Myanmar |journal=Cretaceous Research |volume=133 |pages=Article 105119 |doi=10.1016/j.cretres.2021.105119 |s2cid=245298780 }}</ref>
|
Sp. nov
|
In press
|
Wang ''et al.''
|
[[Earth:Cretaceous|Cretaceous]]
|
[[Earth:Burmese amber|Burmese amber]]
|
{{Flag|Myanmar}}
|
A member of the family [[Biology:Marsileaceae|Marsileaceae]].
|
|-
|
''Nemejcopteris haiwangii''<ref>{{Cite journal|last1=Pšenička |first1=J. |last2=Wang |first2=J. |last3=Bek |first3=J. |last4=Pfefferkorn |first4=H. W. |last5=Opluštil |first5=S. |last6=Zhou |first6=W. |last7=Frojdová |first7=J. |last8=Libertín |first8=M. |year=2021 |title=A zygopterid fern with fertile and vegetative parts in anatomical and compression preservation from the earliest Permian of Inner Mongolia, China |journal=Review of Palaeobotany and Palynology |volume=294 |pages=Article 104382 |doi=10.1016/j.revpalbo.2021.104382 |bibcode=2021RPaPa.29404382P |s2cid=234092362 }}</ref>
|
Sp. nov
|
In press
|
Pšenička ''et al.''
|
Permian ([[Earth:Asselian|Asselian]])
|
Taiyuan Formation
|
{{Flag|China}}
|
A [[Biology:Zygopteridales|zygopterid]] fern.
|
|-
|
''[[Biology:Neocalamites|Neocalamites iranensis]]''<ref>{{Cite journal |last1=Mazaheri-Johari |first1=M. |last2=Kustatscher |first2=E. |last3=Roghi |first3=G. |last4=Ghasemi-Nejad |first4=E. |last5=Gianolla |first5=P. |year=2021 |title=A monotypic stand of ''Neocalamites iranensis'' n. sp. from the Carnian Pluvial Episode (Late Triassic) of the Aghdarband area, NE Iran (Turan Plate) |journal=Rivista Italiana di Paleontologia e Stratigrafia |volume=127 |issue=2 |pages=189–209 |doi=10.13130/2039-4942/15646 }}</ref>
|
Sp. nov
|
Valid
|
Kustatscher, Mazaheri-Johari & Roghi ''in'' Mazaheri-Johari ''et al.''
|
[[Earth:Late Triassic|Late Triassic]] (Carnian)
|
Miakuhi Formation
|
{{Flag|Iran}}
|
A member of the family [[Biology:Equisetaceae|Equisetaceae]].
|
|-
|
''Odontosoria marekgaltieri''<ref>{{Cite journal|last1=Pšenička |first1=J.  |last2=Sakala |first2=J. |last3=Dašková |first3=J. |year=2021 |title=''Odontosoria marekgaltieri'' sp. nov. (Lindsaeaceae), a new fern from the early Miocene of the Czech Republic: First evidence of the genus in the fossil record |journal=Review of Palaeobotany and Palynology |volume=297 |pages=Article 104580 |doi=10.1016/j.revpalbo.2021.104580 }}</ref>
|
Sp. nov
|
In press
|
Pšenička, Sakala & Dašková
|
Early Miocene
|
Most Basin
|
{{Flag|Czech Republic}}
|
A species of ''[[Biology:Odontosoria|Odontosoria]]''.
|
|-
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''Oligosporangiopteris''<ref>{{Cite journal|last1=Votočková Frojdová |first1=J. |last2=Wang |first2=J. |last3=Pšenička |first3=J. |last4=Bek |first4=J. |last5=Opluštil |first5=S. |last6=Libertín |first6=M. |year=2021 |title=A new leptosporangiate fern ''Oligosporangiopteris zhongxiangii'' gen. and sp. nov. from the lowermost Permian of Inner Mongolia, China – morphology, anatomy and reproductive organs |journal=Review of Palaeobotany and Palynology |volume=294 |pages=Article 104479 |doi=10.1016/j.revpalbo.2021.104479 |bibcode=2021RPaPa.29404479V |s2cid=237931032 }}</ref>
|
Gen. et sp. nov
|
In press
|
Votočková Frojdová ''et al.''
|
Early [[Earth:Permian|Permian]]
|
Taiyuan Formation
|
{{Flag|China}}
|
A [[Biology:Leptosporangiate fern|leptosporangiate fern]]. Genus includes new species ''O. zhongxiangii''.
|
|-
|
''Osmunda zhangpuensis''<ref>{{Cite journal|last1=Wang |first1=Z. |last2=Shi |first2=G. |last3=Sun |first3=B. |last4=Dong |first4=C. |last5=Yin |first5=S. |last6=Wu |first6=X. |year=2021 |title=A new species of ''Osmunda'' L. (Osmundaceae) from the middle Miocene of Fujian, Southeast China |journal=Acta Palaeontologica Sinica |volume=60 |issue=3 |pages=Article 2021045 |doi=10.19800/j.cnki.aps.2021045 |url=http://gswxb.cnjournals.cn/gswxb/article/abstract/20210306 }}</ref>
|
Sp. nov
|
|
Wang & Sun ''in'' Wang ''et al.''
|
Miocene
|
Fotan Group
|
{{Flag|China}}
|
A fern, a species of ''[[Biology:Osmunda|Osmunda]]''.
|
|-
|
''Patagoniapteris''<ref>{{Cite journal|last1=Gnaedinger |first1=S. C. |last2=Zavattieri |first2=A. M. |title=A new Late Triassic dipteridacean fern from the Paso Flores Formation, Neuquén Basin, Argentina |year=2021 |journal=Acta Palaeontologica Polonica |volume=66 |issue=4 |pages=885–900 |doi=10.4202/app.00864.2020 |doi-access=free |hdl=11336/171135 |hdl-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Gnaedinger & Zavattieri
|
[[Earth:Late Triassic|Late Triassic]] (Norian–Rhaetian)
|
[[Earth:Paso Flores Formation|Paso Flores Formation]]
|
{{Flag|Argentina}}
|
A member of the family [[Biology:Dipteridaceae|Dipteridaceae]]. Genus includes new species ''P. artabeae''.
|
|-
|
''Pectinangium xuanweiense''<ref>{{Cite journal|last1=Zhou |first1=Y. |last2=Guo |first2=Y. |last3=Pšenička |first3=J. |last4=Bek |first4=J. |last5=Yang |first5=S.-L. |last6=Feng |first6=Z. |year=2021 |title=A new marattialean fern, ''Pectinangium xuanweiense'' sp. nov., from the Lopingian of Southwest China |journal=Review of Palaeobotany and Palynology |volume=295 |pages=Article 104500 |doi=10.1016/j.revpalbo.2021.104500 |bibcode=2021RPaPa.29504500Z |doi-access=free }}</ref>
|
Sp. nov
|
In press
|
Zhou ''et al.''
|
Permian ([[Earth:Lopingian|Lopingian]])
|
|
{{Flag|China}}
|
A fern belonging to the group Marattiales.
|
|-
|
''Polypodiisporites densus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Spores of a member of the family Polypodiaceae.
|
|-
|
''Polypodiisporites fossulatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Spores of a member of the family [[Biology:Lomariopsidaceae|Lomariopsidaceae]].
|
|-
|
''Psilatriletes marginatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Spores of a member of the genus ''[[Biology:Cyathea|Cyathea]]''.
|
|-
|
''Qasimia yunnanica''<ref>{{Cite journal|last1=Guo |first1=Y. |last2=Zhou |first2=Y. |last3=Bek |first3=J. |last4=Yang |first4=S.-L. |last5=Feng |first5=Z. |year=2021 |title=''Qasimia yunnanica'' sp. nov., a marattialean fern with bivalvate synangia from the Lopingian of Southwest China |journal=Review of Palaeobotany and Palynology |volume=293 |pages=Article 104497 |doi=10.1016/j.revpalbo.2021.104497 |bibcode=2021RPaPa.29304497G |doi-access=free }}</ref>
|
Sp. nov
|
In press
|
Guo ''et al.''
|
Permian ([[Earth:Lopingian|Lopingian]])
|
Xuanwei Formation
|
{{Flag|China}}
|
A fern belonging to the group Marattiales.
|
|-
|
''Reticulosporis diversus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Spores of a member of the family Polypodiaceae.
|
|-
|
''Rothwellopteris sanjiaoshuensis''<ref>{{Cite journal|last1=He |first1=X.-Y. |last2=Hilton |first2=J. |last3=Wang |first3=S.-J. |last4=Cheng |first4=X.-S. |year=2021 |title=Exploring the stem to crown group transition in Marattiales: A new species of frond from the late Permian of China with features of the Psaroniaceae and Marattiaceae |journal=Review of Palaeobotany and Palynology |volume=295 |pages=Article 104506 |doi=10.1016/j.revpalbo.2021.104506 |bibcode=2021RPaPa.29504506H }}</ref>
|
Sp. nov
|
In press
|
He ''et al.''
|
Late Permian
|
Xuanwei Formation
|
{{Flag|China}}
|
A fern belonging to the group Marattiales.
|
|-
|
''[[Biology:Sphenophyllum|Sphenophyllum fanwanense]]''<ref>{{Cite journal|last1=Huang |first1=P. |last2=Liu |first2=L. |last3=Liu |first3=L. |last4=Wang |first4=J.-S. |last5=Xue |first5=J.-Z. |title=''Sphenophyllum'' Brongniart (Sphenopsida) from the Upper Devonian of South China |journal=Palaeoworld |year=2022 |volume=31 |issue=3 |pages=402–418 |doi=10.1016/j.palwor.2021.09.007 |s2cid=242991171 }}</ref>
|
Sp. nov
|
In press
|
Huang ''et al.''
|
Late Devonian
|
|
{{Flag|China}}
|
A [[Biology:Sphenophyllales|sphenophyllalean]] [[Biology:Equisetidae|equisetid]]
|
|-
|
''Sphenophyllum parvifolium''<ref>{{Cite journal|last1=Libertín |first1=M. |last2=Bek |first2=J. |last3=Wang |first3=J. |last4=Opluštil |first4=S. |last5=Pšenička |first5=J. |last6=Votočková Frojdová |first6=J. |year=2021 |title=New data about three sphenophylls and their spores from the volcanic tuff of Wuda, Taiyuan Formation, earliest Permian, China |journal=Review of Palaeobotany and Palynology |volume=294 |pages=Article 104484 |doi=10.1016/j.revpalbo.2021.104484 |bibcode=2021RPaPa.29404484L }}</ref>
|
Sp. nov
|
In press
|
Libertín ''et al.''
|
Early Permian
|
Taiyuan Formation
|
{{Flag|China}}
|
A [[Biology:Sphenophyllales|sphenophyllalean]] [[Biology:Equisetidae|equisetid]]
|
|-
|
''Tapelrayen''<ref>{{Cite journal|last1=Machado |first1=M. A. |last2=Vera |first2=E. I. |last3=Passalia |first3=M. G. |last4=Ponce |first4=M. M. |year=2021 |title=Eupolypod ferns with dryopteroid/thelypteroid traits from Arroyo Chacay (Huitrera Formation, Eocene), Río Negro Province, Argentina |journal=Review of Palaeobotany and Palynology |volume=287 |pages=Article 104381 |doi=10.1016/j.revpalbo.2021.104381 |bibcode=2021RPaPa.28704381M |s2cid=233532654 |url=http://rid.unrn.edu.ar/handle/20.500.12049/6970 }}</ref>
|
Gen. et sp. nov
|
In press
|
Machado ''et al.''
|
[[Earth:Eocene|Eocene]]
|
[[Earth:Huitrera Formation|Huitrera Formation]]
|
{{Flag|Argentina}}
|
Fertile remains of a fern comparable with [[Biology:Thelypteridaceae|Thelypteridaceae]] and [[Biology:Dryopteridaceae|Dryopteridaceae]]. Genus includes new species ''T. helgae''.
|
|-
|
''Thyrsopteris cyathindusia''<ref>{{Cite journal|last1=Zhang |first1=H.-R. |last2=Shi |first2=C. |last3=Long |first3=X.-X. |last4=Feng |first4=Q. |last5=Cai |first5=H.-H. |last6=Lü |first6=Y.-T. |last7=Wang |first7=S. |title=A new fossil record of Thyrsopteridaceae (Cyatheales) from the mid-Cretaceous amber of Myanmar |journal=Palaeoworld |year=2022 |volume=31 |issue=3 |pages=478–484 |doi=10.1016/j.palwor.2021.09.002 |s2cid=240519538 }}</ref>
|
Sp. nov
|
In press
|
Zhang ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A tree fern, a species of ''[[Biology:Thyrsopteris|Thyrsopteris]]''.
|
|-
|
''Woodwardia changchangensis''<ref>{{Cite journal|last1=Song |first1=H.-Z. |last2=Naugolnykh |first2=S. V. |last3=Wu |first3=X.-K. |last4=Liu |first4=X.-Y. |last5=Jin |first5=J.-H. |title=Fertile ''Woodwardia'' from the middle Eocene of South China and its implications for palaeogeography and palaeoclimate |journal=Plant Diversity |year=2022 |volume=44 |issue=6 |pages=565–576 |doi=10.1016/j.pld.2021.09.003 |pmid=36540713 |pmc=9751083 |issn=2468-2659 |doi-access=free }}</ref>
|
Sp. nov
|
In press
|
Naugolnykh & Song ''in'' Song ''et al.''
|
Middle Eocene
|
Changchang Formation
|
{{Flag|China}}
|
A fern, a species of ''[[Biology:Woodwardia|Woodwardia]]''.
| 
|-
|}

==Bennettitales==
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''[[Biology:Nilssoniopteris|Nilssoniopteris jogiana]]''<ref name=FEEBlomenkemperetal>{{Cite journal|last1=Blomenkemper |first1=P. |last2=Bäumer |first2=R. |last3=Backer |first3=M. |last4=Abu Hamad |first4=A. |last5=Wang |first5=J. |last6=Kerp |first6=H. |last7=Bomfleur |first7=B. |year=2021 |title=Bennettitalean Leaves From the Permian of Equatorial Pangea—The Early Radiation of an Iconic Mesozoic Gymnosperm Group |journal=Frontiers in Earth Science |volume=9 |pages=Article 652699 |doi=10.3389/feart.2021.652699 |bibcode=2021FrEaS...9..162B |doi-access=free }}</ref>
|
Sp. nov
|
|
Blomenkemper & Abu Hamad ''in'' Blomenkemper ''et al.''
|
[[Earth:Permian|Permian]] ([[Earth:Changhsingian|Changhsingian]])
|
[[Earth:Umm Irna Formation|Umm Irna Formation]]
|
{{Flag|Jordan}}
|
A member of [[Biology:Bennettitales|Bennettitales]].
|
|-
|
''Nilssoniopteris shanxiensis''<ref name=FEEBlomenkemperetal />
|
Sp. nov
|
|
Bäumer, Backer & Wang ''in'' Blomenkemper ''et al.''
|
Permian ([[Earth:Cisuralian|Cisuralian]])
|
Upper Shihhotse Formation
|
{{Flag|China}}
|
A member of Bennettitales.
|
|-
|
''[[Biology:Pterophyllum (plant)|Pterophyllum pottii]]''<ref name=FEEBlomenkemperetal />
|
Sp. nov
|
|
Bomfleur & Kerp ''in'' Blomenkemper ''et al.''
|
Permian (Changhsingian)
|
Umm Irna Formation
|
{{Flag|Jordan}}
|
A member of Bennettitales.
|
|-
|
''[[Biology:Weltrichia|Weltrichia magna]]''<ref>{{Cite journal|last1=Guzmán-Madrid |first1=D. S. |last2=Velasco de León |first2=M. P. |year=2021 |title=''Weltrichia magna'' sp. nov., a new record for the Middle Jurassic of Oaxaca, México |journal=Acta Palaeobotanica |volume=61 |issue=1 |pages=95–106 |doi=10.35535/acpa-2021-0005 |s2cid=237760410 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Guzmán-Madrid & Velasco de León
|
Middle Jurassic (Bajocian)
|
Zorrillo Formation
|
{{Flag|Mexico}}
|
|
|-
|
''[[Biology:Weltrichia|Weltrichia xochitetlii]]''<ref>{{Cite journal|last1=Lozano-Carmona |first1=D. E. |last2=Corro-Ortiz |first2=M. G. |last3=Morales |first3=R. L. |last4=Velasco-de León |first4=M. P. |year=2021 |title=''Weltrichia xochitetlii'' sp. nov. (Bennettitales) from the Middle Jurassic of northwestern Oaxaca, Mexico: First paleobotanical evidence from the Tecomazúchil formation |journal=Journal of South American Earth Sciences |volume=108 |pages=Article 103230 |doi=10.1016/j.jsames.2021.103230|issn=0895-9811 |bibcode=2021JSAES.10803230L |s2cid=234085434 }}</ref>
|
Sp. nov
|
In press
|
Lozano-Carmona ''et al.''
|
Middle Jurassic (Callovian)
|
Tecomazuchil Formation
|
{{Flag|Mexico}}
|
A member of [[Biology:Bennettitales|Bennettitales]].
|
|-
|
''[[Biology:Williamsonia (plant)|Williamsonia sanjuanensis]]''<ref>{{Cite journal|last1=Lozano-Carmona |first1=D. E. |last2=Velasco-de León |first2=M. P. |year=2021 |title=Bennettitales from the Middle Jurassic of northwestern Oaxaca, Mexico: Diversity, sedimentary environments, and phytogeography |journal=Journal of South American Earth Sciences |volume=110 |pages=Article 103404 |doi=10.1016/j.jsames.2021.103404| issn=0895-9811 |bibcode=2021JSAES.11003404L }}</ref>
|
Sp. nov
|
In press
|
Lozano-Carmona & Velasco-de León
|
Middle Jurassic
|
|
{{Flag|Mexico}}
|
|
|-
|}

== Cycadales ==
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Becklesia franconica''<ref>{{Cite journal|last1=Van Konijnenburg-van Cittert |first1=J. H. A. |last2=Pott |first2=C. |last3=Schmeißner |first3=S. |last4=Dütsch |first4=G. |last5=Kustatscher |first5=E. |year=2021 |title=The Rhaetian flora of Wüstenwelsberg, Bavaria, Germany: Description of selected gymnosperms (Ginkgoales, Cycadales, Coniferales) together with an ecological assessment of the locally prevailing vegetation |journal=Review of Palaeobotany and Palynology |volume=288 |pages=Article 104398 |doi=10.1016/j.revpalbo.2021.104398 |bibcode=2021RPaPa.28804398V |s2cid=233918337 |doi-access=free }}</ref>
|
Sp. nov
|
In press
|
Van Konijnenburg-van Cittert ''et al.''
|
[[Earth:Late Triassic|Late Triassic]] (Rhaetian)
|
[[Earth:Exter Formation|Exter Formation]]
|
{{Flag|Germany}}
|
A member of [[Biology:Cycadales|Cycadales]] of uncertain phylogenetic placement.
|
|-
|
''Iratinia''<ref>{{Cite journal|last1=Spiekermann |first1=R. |last2=Jasper |first2=A. |last3=Siegloch |first3=A. M. |last4=Guerra-Sommer |first4=M. |last5=Uhl |first5=D. |year=2021 |title=Not a lycopsid but a cycad-like plant: ''Iratinia australis'' gen. nov. et sp. nov. from the Irati Formation, Kungurian of the Paraná Basin, Brazil |journal=Review of Palaeobotany and Palynology |volume=289 |pages=Article 104415 |doi=10.1016/j.revpalbo.2021.104415 |bibcode=2021RPaPa.28904415S |s2cid=233860955 }}</ref>
|
Gen. et sp. nov
|
In press
|
Spiekermann ''et al.''
|
[[Earth:Permian|Permian]] ([[Earth:Kungurian|Kungurian]])
|
[[Earth:Irati Formation|Irati Formation]]
|
{{Flag|Brazil}}
|
A [[Biology:Cycad|cycad]]-like plant. Genus includes new species ''I. australis''.
|
|-
|}

==Ginkgoales==
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Eretmophyllum hamiensis''<ref>{{Cite journal|last1=Tang |first1=D.-L. |last2=Wang |first2=Z.-E. |last3=Huang |first3=Y.-T. |last4=Ding |first4=H. |last5=Ding |first5=S.-T. |last6=Wu |first6=J.-Y. |title=A new species of ''Eretmophyllum'' (Ginkgoales) from the Middle Jurassic of Turpan-Hami Basin, Xinjiang, China |journal=Palaeoworld |year=2022 |volume=31 |issue=4 |pages=646–657 |doi=10.1016/j.palwor.2021.12.001 |s2cid=245302175 }}</ref>
|
Sp. nov
|
In press
|
Tang ''et al.''
|
Middle Jurassic
|
Xishanyao Formation
|
{{Flag|China}}
|
|
|-
|
''[[Biology:Ginkgoites|Ginkgoites villardeseoanii]]''<ref>{{Cite journal|last1=Andruchow-Colombo |first1=A. |last2=Gandolfo |first2=M. A. |last3=Cúneo |first3=N. R. |last4=Escapa |first4=I. H. |year=2021 |title=''Ginkgoites villardeseoanii'' sp. nov., a ginkgophyte with insect damage from the Upper Cretaceous (Maastrichtian) Lefipán Formation (Chubut, Patagonia, Argentina) |journal=Cretaceous Research |volume=133 |pages=Article 105124 |doi=10.1016/j.cretres.2021.105124 |s2cid=245458671 |doi-access=free }}</ref>
|
Sp. nov
|
In press
|
Andruchow-Colombo ''et al.''
|
[[Earth:Late Cretaceous|Late Cretaceous]] ([[Earth:Maastrichtian|Maastrichtian]])
|
[[Earth:Lefipán Formation|Lefipán Formation]]
|
{{Flag|Argentina}}
|
|
|-
|
''Ginkgoxylon arcticum''<ref>{{Cite journal|last1=Afonin |first1=M. |last2=Gromyko |first2=D. |title=First record of ''Ginkgoxylon'' (Ginkgoales) fossil wood in the Lower Cretaceous of the Arctic region |year=2021 |journal=Cretaceous Research |volume=125 |pages=Article 104868 |doi=10.1016/j.cretres.2021.104868 |bibcode=2021CrRes.12504868A }}</ref>
|
Sp. nov
|
In press
|
Afonin & Gromyko
|
[[Earth:Early Cretaceous|Early Cretaceous]]
|
|
{{Flag|Russia}}<br>({{Flag|Arkhangelsk Oblast}})
|
A member of [[Biology:Ginkgoales|Ginkgoales]] described on the basis of fossil wood.
|
|-
|
''Karkenia irkutensis''<ref>{{Cite journal|last1=Nosova |first1=N. |last2=Crane |first2=P. R. |last3=Shi |first3=G. |year=2021 |title=Ovule-bearing structures of ''Karkenia'' Archangelsky, associated dispersed seeds and ''Sphenobaiera'' leaves from the Middle Jurassic of East Siberia, Russia |journal=Review of Palaeobotany and Palynology |volume=295 |pages=Article 104522 |doi=10.1016/j.revpalbo.2021.104522 |bibcode=2021RPaPa.29504522N |s2cid=239696262 }}</ref>
|
Sp. nov
|
In press
|
Nosova, Crane & Shi
|
Middle Jurassic (Aalenian)
|
Prisayan Formation
|
{{Flag|Russia}}
|
|
|-
|}

==Vladimariales==
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Pseudotorellia doludenkoae''<ref>{{Cite journal|last1=Nosova |first1=N. V. |last2=Kostina |first2=E. I. |last3=Bugdaeva |first3=E. V. |year=2021 |title=''Pseudotorellia'' Florin from the Upper Jurassic–Lower Cretaceous of the Bureya Basin, Russian Far East |journal=Stratigraphy and Geological Correlation |volume=29 |issue=4 |pages=434–449 |doi=10.1134/S0869593821040031 |bibcode=2021SGC....29..434N |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Nosova, Kostina & Bugdaeva
|
Late Jurassic–[[Earth:Early Cretaceous|Early Cretaceous]] (Oxfordian–[[Earth:Berriasian|Berriasian]])
|
Dublikan Formation<br>
Talyndzhan Formation
|
{{Flag|Russia}}<br>({{Flag|Khabarovsk Krai}})
|
|
|-
|''Pseudotorellia irkutensis''<ref name="RPPNosova">{{Cite journal|last=Nosova|first=N.|year=2021|title=Female reproductive structures of ''Umaltolepis'' Krassilov and associated leaves of ''Pseudotorellia'' Florin from the Middle Jurassic of East Siberia, Russia|journal=Review of Palaeobotany and Palynology|volume=289|pages=Article 104412|doi=10.1016/j.revpalbo.2021.104412|bibcode=2021RPaPa.28904412N |s2cid=233790779}}</ref>
|
Sp. nov
|
In press
|
Nosova
|
Middle Jurassic (Aalenian–Bajocian)
|
Prisayan Formation
|
{{Flag|Russia}}
|
A Vladimariales foliage species 
|
|-
|
''[[Biology:Umaltolepis|Umaltolepis irkutensis]]''<ref name=RPPNosova />
|
Sp. nov
|
In press
|
Nosova
|
Middle Jurassic (Aalenian–Bajocian)
|
Prisayan Formation
|
{{Flag|Russia}}
|
A Vladimariales reproducive structure species 
|
|-
|}

== Conifers ==
===Araucariaceae===
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''[[Biology:Agathoxylon|Agathoxylon mendezii]]''<ref>{{Cite journal|last1=Del Fueyo |first1=G. M. |last2=Carrizo |first2=A. |last3=Poiré |first3=D. G. |last4=Lafuente Diaz |first4=M. A. |title=Recurrent volcanic activity recorded in araucarian wood from the Lower Cretaceous Springhill Formation, Patagonia, Argentina: Palaeoenvironmental interpretations |year=2021 |journal=Acta Palaeontologica Polonica |volume=66 |issue=1 |pages=231–253 |doi=10.4202/app.00783.2020 |doi-access=free |hdl=11336/137848 |hdl-access=free }}</ref>
|
Sp. nov
|
Valid
|
Del Fueyo ''et al.''
|
[[Earth:Early Cretaceous|Early Cretaceous]] ([[Earth:Berriasian|Berriasian]]–[[Earth:Valanginian|Valanginian]])
|
Springhill Formation
|
{{Flag|Argentina}}
|
An [[Biology:Araucariaceae|Araucariaceae]] fossil wood.
|
|-
|
''Agathoxylon santanensis''<ref>{{Cite journal|last1=dos Santos |first1=Â. C. S. |last2=Siegloch |first2=A. M. |last3=Guerra-Sommer |first3=M. |last4=Degani-Schmidt |first4=I. |last5=Carvalho |first5=I. S. |year=2021 |title=''Agathoxylon santanensis'' sp. nov. from the Aptian Crato fossil Lagerstätte, Santana Formation, Araripe Basin, Brazil |journal=Journal of South American Earth Sciences |volume=112, Part 2 |pages=Article 103633 |doi=10.1016/j.jsames.2021.103633 |bibcode=2021JSAES.11203633S |s2cid=244110901 }}</ref>
|
Sp. nov
|
In press
|
Dos Santos ''et al.''
|
Early Cretaceous (Aptian)
|
Crato Formation
|
{{Flag|Brazil}}
|
|
|-
|
''Araucaria violetae''<ref>{{Cite journal|last1=Batista |first1=M. E. P. |last2=Loiola |first2=M. I. B. |last3=Soares |first3=A. A. |last4=Mastroberti |first4=A. A. |last5=Sá |first5=A. A. |last6=Nascimento Jr. |first6=D. R. |last7=Silva Filho |first7=W. F. |last8=Kunzmann |first8=L. |year=2021 |title=New Insights into the Evolution of Mucilage Cells in Araucariaceae: ''Araucaria violetae'' sp. nov. from the Early Cretaceous Araripe Basin (Northeast Brazil) |journal=International Journal of Plant Sciences |volume=183 |issue=1 |pages=43–60 |doi=10.1086/717104 |s2cid=239548378 }}</ref>
|
Sp. nov
|
In press
|
Batista ''et al.''
|
[[Earth:Early Cretaceous|Early Cretaceous]]
|
|
{{Flag|Brazil}}
|
A species of ''[[Biology:Araucaria|Araucaria]]''.
|
|-
|}

===Cheirolepidiaceae===
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Brachyoxylon lalongense''<ref>{{Cite journal|last1=Yang |first1=X.-J. |last2=Li |first2=J.-G. |title=A petrified wood ''Brachyoxylon'' from the Lower Cretaceous of Bangoin, Tibet (Xizang), Southwest China |year=2021 |journal=Cretaceous Research |volume=130 |pages=Article 105064 |doi=10.1016/j.cretres.2021.105064 |s2cid=240429516 }}</ref>
|
Sp. nov
|
In press
|
Yang & Li
|
[[Earth:Early Cretaceous|Early Cretaceous]] ([[Earth:Berriasian|Berriasian]]–[[Earth:Barremian|Barremian]])
|
Duoni Formation
|
{{Flag|China}}
|
|
|-
|
''Brachyoxylon patagonicum''<ref>{{Cite journal|last1=Rombola |first1=C. F. |last2=Greppi |first2=C. D. |last3=Pujana |first3=R. R. |last4=García Massini |first4=J. L. |last5=Bellosi |first5=E. S. |last6=Marenssi |first6=S. A. |title=''Brachyoxylon'' fossil woods with traumatic resin canals from the Upper Cretaceous Cerro Fortaleza Formation, southern Patagonia (Santa Cruz Province, Argentina) |year=2021 |journal=Cretaceous Research |volume=130 |pages=Article 105065 |doi=10.1016/j.cretres.2021.105065 |hdl=11336/148807 |s2cid=239597933 |url=https://www.sciencedirect.com/science/article/abs/pii/S019566712100313X |hdl-access=free }}</ref>
|
Sp. nov
|
In press
|
Rombola ''et al.''
|
[[Earth:Late Cretaceous|Late Cretaceous]]
|
[[Earth:Cerro Fortaleza Formation|Cerro Fortaleza Formation]]
|
{{Flag|Argentina}}
|
Fossil wood of a member of the family [[Biology:Cheirolepidiaceae|Cheirolepidiaceae]].
|
|-
|
''Watsoniocladus cunhae''<ref>{{Cite journal|last1=Kvaček |first1=J. |last2=Mendes |first2=M. M. |year=2021 |title=A new Cheirolepidiaceae conifer ''Watsoniocladus cunhae'' sp. nov. from the Early Cretaceous (late Aptian–early Albian) of western Portugal |journal=Review of Palaeobotany and Palynology |volume=295 |pages=Article 104519 |doi=10.1016/j.revpalbo.2021.104519 |bibcode=2021RPaPa.29504519K }}</ref>
|
Sp. nov
|
In press
|
Kvaček & Mendes
|
[[Earth:Early Cretaceous|Early Cretaceous]] ([[Earth:Aptian|Aptian]]–[[Earth:Albian|Albian]])
|
Almargem Formation
|
{{Flag|Portugal}}
|
A member of the family [[Biology:Cheirolepidiaceae|Cheirolepidiaceae]].
|
|-
|}

===Cupressaceae===
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Cupressinoxylon widdringtonioides''<ref>{{Cite journal|last1=de Wit |first1=M. |last2=Bamford |first2=M. |title=Fossil wood from the Upper Cretaceous crater sediments of the Salpeterkop volcano, North West Province, South Africa |year=2021 |journal=South African Journal of Geology |volume=124 |issue=3 |pages=751–760 |doi=10.25131/sajg.124.0028 |bibcode=2021SAJG..124..751D }}</ref>
|
Sp. nov
|
Valid
|
De Wit & Bamford
|
[[Earth:Late Cretaceous|Late Cretaceous]]
|
|
{{Flag|South Africa}}
|
Fossil wood of a member or a relative of the family [[Biology:Cupressaceae|Cupressaceae]].
|
|-
|
''Fokienia tianpingensis''<ref>{{Cite journal|last1=Wu |first1=X. |last2=Zhang |first2=H. |last3=Kodrul |first3=T. M. |last4=Maslova |first4=N. P. |last5=Jiang |first5=S. |last6=Yin |first6=Q. |last7=Quan |first7=C. |last8=Jin |first8=J. |title=First Fossil ''Fokienia'' (Cupressaceae) in South China and Its Palaeogeographic and Palaeoecological Implications |year=2021 |journal=Frontiers in Earth Science |volume=9 |pages=Article 709663 |doi=10.3389/feart.2021.709663 |bibcode=2021FrEaS...9..555W |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Wu & Jin ''in'' Wu ''et al.''
|
Miocene
|
Erzitang Formation
|
{{Flag|China}}
|
A species of ''[[Biology:Fokienia|Fokienia]]''.
|
|-
|
''Nishidastrobus''<ref name=Nishidastrobus>{{Cite journal|last1=Atkinson |first1=B. A. |last2=Contreras |first2=D. L. |last3=Stockey |first3=R. A. |last4=Rothwell |first4=G. W. |title=Ancient diversity and turnover of cunninghamioid conifers (Cupressaceae): two new genera from the Upper Cretaceous of Hokkaido, Japan |year=2021 |journal=Botany |volume=99 |issue=8 |pages=457–473 |doi=10.1139/cjb-2021-0005 |s2cid=237705866 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Atkinson ''et al.''
|
[[Earth:Late Cretaceous|Late Cretaceous]]
|
|
{{Flag|Japan}}
|
A member of the family Cupressaceae belonging to the subfamily [[Biology:Cunninghamioideae|Cunninghamioideae]]. Genus includes new species ''N. japonicum''.
|
|-
|
''Ohanastrobus''<ref name=Nishidastrobus />
|
Gen. et sp. nov
|
Valid
|
Atkinson ''et al.''
|
Late Cretaceous
|
|
{{Flag|Japan}}
|
A member of the family Cupressaceae belonging to the subfamily Cunninghamioideae. Genus includes new species ''O. hokkaidoensis''.
|
|-
|
''Protaxodioxylon metangulense''<ref name=JAESProtaxodioxylon>{{Cite journal|last1=Nhamutole |first1=N. |last2=Bamford |first2=M. |last3=Araújo |first3=R. |year=2021 |title=New species of ''Protaxodioxylon'' (conifer wood) from the Middle Permian of the Metangula Graben (Niassa Province, Mozambique) and their implications |journal=Journal of African Earth Sciences |volume=183 |pages=Article 104323 |doi=10.1016/j.jafrearsci.2021.104323 |bibcode=2021JAfES.18304323N }}</ref>
|
Sp. nov
|
In press
|
Nhamutole, Bamford & Araújo
|
[[Earth:Permian|Permian]]  (late [[Earth:Capitanian|Capitanian]])
|
K5 Formation
|
{{Flag|Mozambique}}
|
A member of the family [[Biology:Cupressaceae|Cupressaceae]].
|
|-
|
''Protaxodioxylon verniersii''<ref name=JAESProtaxodioxylon />
|
Sp. nov
|
In press
|
Nhamutole & Bamford ''in'' Nhamutole, Bamford & Araújo
|
Permian (late Capitanian)
|
K5 Formation
|
{{Flag|Mozambique}}
|
A member of the family Cupressaceae.
|
|-
|
''Thujopsoxylon''<ref name=Thujopsoxylon />
|
Gen. et sp. nov
|
Valid
|
Dolezych, LePage & Williams
|
Oligocene (Chattian)
|
Korlikov Formation
|
{{Flag|Russia}}<br>({{Flag|Tomsk Oblast}})
|
Genus includes new species ''T. schneiderianum''.
|
|-
|}

===Pinaceae===
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Lepidocasus''<ref>{{Cite journal|last1=Herrera |first1=F. |last2=Shi |first2=G. |last3=Bickner |first3=M. A. |last4=Ichinnorov |first4=N. |last5=Leslie |first5=A. B. |last6=Crane |first6=P. R. |last7=Herendeen |first7=P. S. |year=2021 |title=Early Cretaceous abietoid Pinaceae from Mongolia and the history of seed scale shedding |journal=American Journal of Botany |volume=108 |issue=8 |pages=1483–1499 |doi=10.1002/ajb2.1713|issn=0002-9122  |pmid=34458982 |s2cid=239667187 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Herrera ''et al.''
|
[[Earth:Early Cretaceous|Early Cretaceous]] ([[Earth:Aptian|Aptian]]–[[Earth:Albian|Albian]])
|
|
{{Flag|Mongolia}}
|
A member of the family [[Biology:Pinaceae|Pinaceae]]. Genus includes new species ''L. mellonae''.
|
|-
|
''Nothotsuga sinogaia''<ref>{{Cite journal|last1=Ding |first1=S.-T. |last2=Chen |first2=S.-Y. |last3=Ruan |first3=S.-C. |last4=Yang |first4=M. |last5=Han |first5=Y. |last6=Wang |first6=X.-H. |last7=Zhang |first7=T.-H. |last8=Sun |first8=B.-N. |year=2021 |title=First fossil record of ''Nothotsuga'' (Pinaceae) in China: implications for palaeobiogeography and palaeoecology |journal=Historical Biology: An International Journal of Paleobiology |volume=33 |issue=12 |pages=3617–3624 |doi=10.1080/08912963.2021.1881781 |s2cid=233975517 }}</ref>
|
Sp. nov
|
In press
|
Ding ''et al.''
|
Late Miocene
| 
|
{{Flag|China}}
|
A species of ''[[Biology:Nothotsuga|Nothotsuga]]''
|
|-
|
''Piceoxylon nikitinii''<ref name=Thujopsoxylon>{{Cite journal|last1=Dolezych |first1=M. |last2=LePage |first2=B. A. |last3=Williams |first3=C. J. |year=2021 |title=A Chattian-Aquitanian wood flora from the West Siberian Plain: Implications for regional palaeobiogeography |journal=Palaeontographica Abteilung B |volume=302 |issue=1–6 |pages=37–169 |doi=10.1127/palb/2021/0074 |bibcode=2021PalAB.302...37D |s2cid=237651777 }}</ref>
|
Sp. nov
|
Valid
|
Dolezych, LePage & Williams
|
Oligocene (Chattian)
|
Korlikov Formation
|
{{Flag|Russia}}<br>({{Flag|Tomsk Oblast}})
|
A ''[[Biology:Picea|Picea]]'' relative wood.
|
|-
|
''[[Biology:Pinus leptokrempfii|Pinus leptokrempfii]]''<ref>{{Cite journal|last1=Zhang |first1=J.-W. |last2=Wang |first2=L. |last3=D'Rozario |first3=A. |last4=Liang |first4=X.-Q. |last5=Huang |first5=J. |last6=Zhou |first6=Z.-K. |year=2021 |title=''Pinus leptokrempfii'', an Oligocene Relative of the Flat-Needled Pine ''P. krempfii'' (Pinaceae) from China: Implications for Paleogeographic Origin |journal=International Journal of Plant Sciences |volume=182 |issue=5 |pages=389–400 |doi=10.1086/713957 |s2cid=233888471 }}</ref>
|
Sp. nov
|
Valid
|
Zhang ''et al.''
|
Early [[Earth:Oligocene|Oligocene]]
|
|
{{Flag|China}}
|
A [[Biology:Pine|pine]].
|
|-
|
''Pinus nongyaplongensis''<ref>{{Cite journal|last1=Grote |first1=P. J. |last2=Srisuk |first2=P. |year=2021 |title=Fossil ''Pinus'' from the Cenozoic of Thailand |journal=Review of Palaeobotany and Palynology |volume=295 |pages=Article 104501 |doi=10.1016/j.revpalbo.2021.104501 |bibcode=2021RPaPa.29504501G |issn=0034-6667  }}</ref>
|
Sp. nov
|
In press
|
Grote ''in'' Grote & Srisuk
|
Oligocene-early Miocene
|
|
{{Flag|Thailand}}
|
A pine.
|
|-
|
''Pinus weichangensis''<ref>{{Cite journal|last1=Li |first1=Y. |last2=Yi |first2=T.-M. |last3=Grote |first3=P. J. |last4=An |first4=P.-C. |last5=Zhu |first5=Y.-B. |last6=Zhang |first6=Z.-Y. |last7=Li |first7=C.-S. |year=2021 |title=A new species of ''Pinus'' (Pinaceae) from the Miocene of Weichang, Hebei Province, China and its evolutionary significance |journal=Historical Biology: An International Journal of Paleobiology |volume=34 |issue=5 |pages=885–896 |doi=10.1080/08912963.2021.1952197 |s2cid=237692568 }}</ref>
|
Sp. nov
|
In press
|
Li ''et al.''
|
Early Miocene
| 
|
{{Flag|China}}
|
A pine.
|
|-
|
''Schizolepidopsis ediae''<ref>{{Cite journal|last1=Matsunaga |first1=K. K. S. |last2=Herendeen |first2=P. S. |last3=Herrera |first3=F. |last4=Ichinnorov |first4=N. |last5=Crane |first5=P. R. |last6=Shi |first6=G. |year=2021 |title=Ovulate Cones of ''Schizolepidopsis ediae'' sp. nov. Provide Insights into the Evolution of Pinaceae |journal=International Journal of Plant Sciences |volume=182 |issue=6 |pages=490–507 |doi=10.1086/714281 |s2cid=235426888 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Matsunaga ''et al.''
|
[[Earth:Early Cretaceous|Early Cretaceous]]
|
Huolinhe Formation<br>
Tevshiin Govi Formation
|
{{Flag|China}}<br>
{{Flag|Mongolia}}
|
A member or a close relative of the family Pinaceae.
|
|-
|}

===Podocarpaceae===
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Podocarpus yunnanensis''<ref>{{Cite journal|last1=Wu |first1=J. |last2=Chen |first2=H. |last3=Ruan |first3=S. |last4=Yang |first4=M. |last5=Mo |first5=L. |last6=Ji |first6=B. |last7=Zhang |first7=J. |last8=Ding |first8=S. |year=2021 |title=Fossil leaves of ''Podocarpus'' subgenus ''Foliolatus'' (Podocarpaceae) from the Pliocene of southwestern China and biogeographic history of ''Podocarpus'' |journal=Review of Palaeobotany and Palynology |volume=287 |pages=Article 104380 |doi=10.1016/j.revpalbo.2021.104380 |bibcode=2021RPaPa.28704380W |s2cid=234282697 }}</ref>
|
Sp. nov
|
In press
|
Wu ''et al.''
|
Early Pliocene
|
|
{{Flag|China}}
|
A species of ''[[Biology:Podocarpus|Podocarpus]]''.
| 
|-
|
''Protophyllocladoxylon hilarioense''<ref>{{Cite journal|last1=Vallejos Leiz |first1=L. |last2=Crisafulli |first2=A. |last3=Gnaedinger |first3=S. |title=New records of Late Triassic wood from Argentina and their biostratigraphic, paleoclimatic, and paleoecological implications |journal=Acta Palaeontologica Polonica |volume=67 |issue=2 |year=2022 |pages=329–340 |doi=10.4202/app.00939.2021 |doi-access=free }}</ref>
|
Sp. nov
|
In press
|
Vallejos Leiz, Crisafulli & Gnaedinger
|
[[Earth:Late Triassic|Late Triassic]] (Norian–Rhaetian)
|
Hilario Formation
|
{{Flag|Argentina}}
|
A member of the family [[Biology:Podocarpaceae|Podocarpaceae]].
|
|-
|
''Protophyllocladoxylon yiwuense''<ref>{{Cite journal|last1=Gou |first1=X.-D. |last2=Feng |first2=Z. |last3=Wei |first3=H.-B. |last4=Lv |first4=Y. |last5=Yang |first5=S.-L. |year=2021 |title=A new ''Protophyllocladoxylon'' stem from the Xishanyao Formation (Middle Jurassic) in the Santanghu Basin, Xinjiang, Northwest China |journal=Review of Palaeobotany and Palynology |volume=292 |pages=Article 104474 |doi=10.1016/j.revpalbo.2021.104474 |bibcode=2021RPaPa.29204474G |doi-access=free }}</ref>
|
Sp. nov
|
In press
|
Gou & Feng ''in'' Gou ''et al.''
|
Middle Jurassic
|
Xishanyao Formation
|
{{Flag|China}}
|
A conifer of uncertain phylogenetic placement, possibly belonging or related to the family Podocarpaceae.
|
|-
|}

===Other conifers===
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Cargalostrobus''<ref>{{Cite journal|last=Gomankov |first=A. V. |year=2021 |title=On conifer fructifications from the Cargala Mines (the Kazanian of the Southern Fore-Urals) |journal=Lethaea Rossica |volume=23 |pages=21–31 |url=http://paleobot.ru/pdf/02_2021_23.pdf }}</ref>
|
Gen. et sp. nov
|
Valid
|
Gomankov
|
[[Earth:Permian|Permian]]
|
|
{{Flag|Russia}}<br>({{Flag|Orenburg Oblast}})
|
A member of [[Biology:Pinales|Pinales]] belonging to the family Sashiniaceae. Genus includes new species ''C. demetrii''.
|
|-
|
''Megaporoxylon sinensis''<ref>{{Cite journal|last1=Wan |first1=M. |last2=Wang |first2=J. |last3=Shi |first3=T. |last4=Wang |first4=K. |last5=Tang |first5=P. |last6=Wang |first6=J. |year=2021 |title=''Megaporoxylon sinensis'' sp. nov., a new coniferous trunk from the Upper Triassic of northern Bogda Mountains, northwestern China |journal=Review of Palaeobotany and Palynology |volume=295 |pages=Article 104536 |doi=10.1016/j.revpalbo.2021.104536 |bibcode=2021RPaPa.29504536W |s2cid=244206721 }}</ref>
|
Sp. nov
|
In press
|
Wan ''et al.''
|
[[Earth:Late Triassic|Late Triassic]] (Carnian–Norian)
|
Huangshanjie Formation
|
{{Flag|China}}
|
A coniferous trunk.
|
|-
|
''Taxus huolingolensis''<ref>{{Cite journal|last1=Dong |first1=C. |last2=Shi |first2=G. |last3=Herrera |first3=F. |last4=Wang |first4=Y. |last5=Wang |first5=Z. |last6=Zhang |first6=B. |last7=Xu |first7=X. |last8=Herendeen |first8=P. S. |last9=Crane |first9=P. R. |year=2021 |title=Leaves of ''Taxus'' with cuticle micromorphology from the Early Cretaceous of eastern Inner Mongolia, Northeast China |journal=Review of Palaeobotany and Palynology |volume=298 |pages=Article 104588 |doi=10.1016/j.revpalbo.2021.104588 |s2cid=245558315 |doi-access=free }}</ref>
|
Sp. nov
|
In press
|
Dong ''et al.''
|
[[Earth:Early Cretaceous|Early Cretaceous]]
|
Huolinhe Formation
|
{{Flag|China}}
|
A species of ''[[Biology:Taxus|Taxus]]''.
|
|-
|
''Voltzia edithae''<ref>{{Cite journal|last1=Forte |first1=G. |last2=Kustatscher |first2=E. |last3=Van Konijnenburg-van Cittert |first3=J. H. A. |year=2021 |title=Conifer diversity in the Middle Triassic: new data from the Fossillagerstätte Kühwiesenkopf/Monte Prà della Vacca (Pelsonian, Anisian) in the Dolomites (NE Italy) |journal=International Journal of Plant Sciences |volume=182 |issue=6 |pages=445–467 |doi=10.1086/714280 |s2cid=233649930 }}</ref>
|
Sp. nov
|
Valid
|
Forte, Kustatscher & Van Konijnenburg-van Cittert
|
[[Earth:Middle Triassic|Middle Triassic]] (Anisian)
|
|
{{Flag|Italy}}
|
A member of [[Biology:Voltziales|Voltziales]].
|
|-
|
''Xenoxylon utahense''<ref>{{Cite journal|last1=Xie |first1=A. |last2=Gee |first2=C. T. |last3=Bennis |first3=M. B. |last4=Gray |first4=D. |last5=Sprinkel |first5=D. A. |year=2021 |title=A more southerly occurrence of ''Xenoxylon'' in North America: ''X. utahense'' Xie et Gee sp. nov. from the Upper Jurassic Morrison Formation in Utah, USA, and its paleobiogeographic and paleoclimatic significance |journal=Review of Palaeobotany and Palynology |volume=291 |pages=Article 104451 |doi=10.1016/j.revpalbo.2021.104451 |bibcode=2021RPaPa.29104451X |s2cid=236239787 }}</ref>
|
Sp. nov
|
In press
|
Xie & Gee ''in'' Xie ''et al.''
|
Late Jurassic
|
[[Earth:Morrison Formation|Morrison Formation]]
|
{{Flag|United States}}<br>({{Flag|Utah}})
|
Fossil wood of a conifer.
|
|-
|
''Zhuotingoxylon''<ref>{{Cite journal|last1=Wan |first1=M. |last2=Yang |first2=W. |last3=Wang |first3=K. |last4=Liu |first4=L. |last5=Wang |first5=J. |title=''Zhuotingoxylon liaoi'' gen. et sp. nov., a silicified coniferous trunk from the Changhsingian (Permian) of southern Bogda Mountains, northwestern China |year=2021 |journal=Geological Journal |volume=56 |issue=12 |pages=6135–6150 |doi=10.1002/gj.4189 |s2cid=236276022 }}</ref>
|
Gen. et sp. nov
|
In press
|
Wan ''et al.''
|
[[Earth:Permian|Permian]] ([[Earth:Changhsingian|Changhsingian]])
|
Guodikeng Formation
|
{{Flag|China}}
|
A silicified trunk with coniferous affinities. Genus includes new species ''Z. liaoi''.
|
|-
|}

==Flowering plants==
===Basal angiosperms===
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Allonymphaea''<ref name=Allonymphaea>{{Cite journal|last=Doweld |first=A. B. |year=2021 |title=Fossil ''Alloceltidoxylon'', ''Allonymphaea'', ''Arecocaryon'', ''Paralnoxylon'' and ''Paranyssa'' and extant ''Komaroviopsis'', ''Marcanodendron'', and ''Papyrocactus'' (Magnoliophyta), new replacement generic names |journal=Phytotaxa |volume=524 |issue=2 |pages=92–98 |doi=10.11646/phytotaxa.524.2.3 |s2cid=243482734 }}</ref>
|
Nom. nov
|
Valid
|
Doweld
|
Eocene
|
|
{{Flag|Egypt}}
|
A replacement name for ''Thiebaudia'' Chandler (1954).
|
|-
|}

===Unplaced non-eudicots===
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Alcainea''<ref name=Alcainea>{{Cite journal|last1=Sender |first1=L. M. |last2=Doyle |first2=J. A. |last3=Upchurch |first3=G. R. |last4=Endress |first4=P. K. |last5=Villanueva-Amadoz |first5=U. |last6=Diez |first6=J. B. |year=2021 |title=Evidence on vegetative and inflorescence morphology of Chloranthaceae (Angiospermae) from the Early Cretaceous (middle–late Albian) of Spain |journal=Journal of Systematic Palaeontology |volume=18 |issue=24 |pages=2015–2042 |doi=10.1080/14772019.2021.1873434 |s2cid=232116303 |url=https://figshare.com/articles/dataset/Evidence_on_vegetative_and_inflorescence_morphology_of_Chloranthaceae_Angiospermae_from_the_Early_Cretaceous_middle_late_Albian_of_Spain/14138245 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Sender ''et al.''
|
[[Earth:Early Cretaceous|Early Cretaceous]] ([[Earth:Albian|Albian]])
|
[[Earth:Escucha Formation|Escucha Formation]]
|
{{Flag|Spain}}
|
A member of the family [[Biology:Chloranthaceae|Chloranthaceae]]. Genus includes new species ''A. eklundiae''.
|
|-
|
''Todziaphyllum''<ref name=Alcainea />
|
Gen. et sp. nov
|
Valid
|
Sender ''et al.''
|
Early Cretaceous (Albian)
|
Escucha Formation
|
{{Flag|Spain}}
|
A member of the family Chloranthaceae. Genus includes new species ''T. elongatum''.
|
|-
|}

===Magnoliids===
====Canellales====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Aristolochia macginitieana''<ref>{{Cite journal|last1=Freitas |first1=J. |last2=Doweld |first2=A. B. |year=2021 |title=''Aristolochia macginitieana'' (Aristolochiaceae), a replacement name for ''Aristolochia triangularis'' MacGinitie non ''Aristolochia triangularis'' Chamisso |journal=Phytotaxa |volume=500 |issue=1 |pages=59–60 |doi=10.11646/phytotaxa.500.1.11 |s2cid=236589284 }}</ref>
|
Nom. nov
|
Valid
|
Freitas & Doweld
|
Oligocene
|
|
{{Flag|United States}}<br>({{Flag|California}})
|
An ''[[Biology:Aristolochia|Aristolochia]]'' species; a replacement name for ''Aristolochia triangularis'' MacGinitie (1937).
| 
|-
|
''Cryptocaryoxylon grandoleaceum''<ref name=AP611 />
|
Sp. nov
|
Valid
|
Akkemik
|
Middle Miocene
|
Kesmekaya Volcanics
|
{{Flag|Turkey}}
|
A member of the family [[Biology:Lauraceae|Lauraceae]].
|
|-
|
''Laurus elliptica''<ref>{{Cite journal|last1=Winterscheid |first1=H. |last2=Kvaček |first2=Z. |year=2021 |title=Systematic-taxonomic revision of the flora from the late Oligocene Fossillagerstätte Rott near Bonn (Germany). Part 2: Magnoliidae: Basal angiosperms and magnoliids |journal=Palaeontographica Abteilung B |volume=303 |issue=4–6 |pages=119–155 |doi=10.1127/palb/2021/0077 |bibcode=2021PalAB.303..119W |s2cid=244053928 }}</ref>
|
Nom. nov
|
Valid
|
Winterscheid ''in'' Winterscheid & Kvaček
|
Oligocene
|
|
{{Flag|Germany}}
|
A species of ''[[Biology:Laurus|Laurus]]''; a replacement name for ''Laurus obovata'' Weber (1852).
|
|-
|
''Rosarioxylon''<ref>{{Cite journal|last1=Cevallos-Ferriz |first1=S. R. S. |last2=Catharina |first2=A. S. |last3=Kneller |first3=B. |year=2021 |title=Cretaceous Lauraceae wood from El Rosario, Baja California, Mexico |journal=Review of Palaeobotany and Palynology |volume=292 |pages=Article 104478 |doi=10.1016/j.revpalbo.2021.104478 |bibcode=2021RPaPa.29204478C }}</ref>
|
Gen. et sp. nov
|
In press
|
Cevallos-Ferriz, Catharina & Kneller
|
[[Earth:Late Cretaceous|Late Cretaceous]] ([[Earth:Campanian|Campanian]])
|
Rosario Formation
|
{{Flag|Mexico}}
|
A member of the family Lauraceae. Genus includes new species ''R. bajacaliforniensis''.
|
|-
|
''Winteroxylon oleiferum''<ref>{{Cite journal|last1=Brea |first1=M. |last2=Iglesias |first2=A. |last3=Wilf |first3=P. |last4=Moya |first4=E. |last5=Gandolfo |first5=M. A. |year=2021 |title=First South American Record of ''Winteroxylon'', Eocene of Laguna del Hunco (Chubut, Patagonia, Argentina): New Link to Australasia and Malesia |journal=International Journal of Plant Sciences |volume=182 |issue=3 |pages=185–197 |doi=10.1086/712427 |issn=1058-5893 |s2cid=232050459 }}</ref>
|
Sp. nov
|
Valid
|
Brea ''et al.''
|
Early [[Earth:Eocene|Eocene]]
|
[[Earth:Huitrera Formation|Huitrera Formation]]
|
{{Flag|Argentina}}
|
A member of the family [[Biology:Winteraceae|Winteraceae]].
| 
|-
|}

===Monocots===
====Alismatid monocots====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Bognerospadix''<ref>{{Cite journal|last1=Stockey |first1=R. A. |last2=Hoffman |first2=G. L. |last3=Rothwell |first3=G. W. |year=2021 |title=Fossil evidence for Paleocene diversification of Araceae: ''Bognerospadix'' gen. nov. and ''Orontiophyllum grandifolium'' comb. nov. |journal=American Journal of Botany |volume=108 |issue=8 |pages=1417–1440  |doi=10.1002/ajb2.1707 |pmid=34431509 |s2cid=237292226 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Stockey, Hoffman & Rothwell
|
[[Earth:Paleocene|Paleocene]]
|
|
{{Flag|Canada}}<br>({{Flag|Alberta}})
|
A member of the family [[Biology:Araceae|Araceae]]. Genus includes new species ''B. speirsiae''.
|
|-
|}

====Lilioid monocots====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Mirafloris''<ref>{{Cite journal|last=Poinar |first=G. O. |title=A Monocot Flower, ''Mirafloris burmitis'' gen. et sp. nov. (Monocots: Angiospermae), in Burmese Amber |year=2021 |journal=Biosis: Biological Systems |volume=2 |issue=3 |pages=342–348 |doi=10.37819/biosis.002.03.0126 |s2cid=238732295 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Poinar
|
Cretaceous
|
[[Earth:Burmese amber|Burmese amber]]
|
{{Flag|Myanmar}}
|
A member of the family [[Biology:Liliaceae|Liliaceae]]. Genus includes new species ''M. burmitis''.
|
|-
|}
====Commelinid monocots====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Arecipites invaginatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the family [[Biology:Arecaceae|Arecaceae]].
|
|-
|
''Arecocaryon''<ref name=Allonymphaea />
|
Nom. nov
|
Valid
|
Doweld
|
Eocene
|
Messel pit
|
{{Flag|Germany}}
|
A member of the family [[Biology:Arecaceae|Arecaceae]]; a replacement name for ''Friedemannia'' Collinson, Manchester & Wilde ([[Biology:2012 in paleobotany|2012]]).
|
|-
|
''Cyperaceaepollis wesselinghii''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the family [[Biology:Cyperaceae|Cyperaceae]].
|
|-
|
''Eograminis''<ref>{{Cite journal|last1=Poinar |first1=G. |last2=Soreng |first2=R. J. |year=2021 |title=A New Genus and Species of Grass, ''Eograminis balticus'' (Poaceae: Arundinoideae) in Baltic Amber |journal=International Journal of Plant Sciences |volume=182 |issue=9 |pages=808–816 |doi=10.1086/716781 |s2cid=239036777 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Poinar & Soreng
|
[[Earth:Eocene|Eocene]]
|
[[Earth:Baltic amber|Baltic amber]]
|
{{Flag|Russia}}<br>({{Flag|Kaliningrad Oblast}})
|
Grass belonging to the subfamily [[Biology:Arundinoideae|Arundinoideae]]. Genus includes new species ''E. balticus''.
|
|-
|
''Luminidites amazonicus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the family Arecaceae.
|
|-
|
''Orthogonospermum''<ref>{{Cite journal|last1=Smith |first1=S. Y. |last2=Kapgate |first2=D. K. |last3=Robinson |first3=S. |last4=Srivastava |first4=R. |last5=Benedict |first5=J. C. |last6=Manchester |first6=S. R. |year=2021 |title=Fossil fruits and seeds of Zingiberales from the Late Cretaceous–early Cenozoic Deccan Intertrappean Beds of India |journal=International Journal of Plant Sciences |volume=182 |issue=2 |pages=91–108 |doi=10.1086/711474 |s2cid=231875495 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Smith ''et al.''
|
Late Cretaceous ([[Earth:Maastrichtian|Maastrichtian]])
|
[[Earth:Deccan Traps|Deccan]] [[Earth:Intertrappean Beds|Intertrappean Beds]]
|
{{Flag|India}}
|
A member of the family [[Biology:Zingiberaceae|Zingiberaceae]]. Genus includes new species ''O. patanense''.
| 
|-
|
''Sabalites colaniae''<ref>{{Cite journal|last1=Song |first1=A. |last2=Liu |first2=J. |last3=Liang |first3=S.-Q. |last4=Do |first4=T. V. |last5=Nguyen |first5=H. B. |last6=Deng |first6=W.-Y.-D. |last7=Jia |first7=L.-B. |last8=Del Rio |first8=C. |last9=Srivastava |first9=G. |last10=Feng |first10=Z. |last11=Zhou |first11=Z.-K. |last12=Huang |first12=J. |last13=Su |first13=T. |title=Leaf fossils of ''Sabalites'' (Arecaceae) from the Oligocene of northern Vietnam and their paleoclimatic implications |journal=Plant Diversity |year=2022 |volume=44 |issue=4 |pages=406–416 |doi=10.1016/j.pld.2021.08.003 |pmid=35967257 |pmc=9363516 |doi-access=free }}</ref>
|
Sp. nov
|
In press
|
Song, Su, Do & Zhou ''in'' Song ''et al.''
|
[[Earth:Oligocene|Oligocene]]
|
Dong Ho Formation
|
{{Flag|Vietnam}}
|
A member of the family Arecaceae belonging to the subfamily [[Biology:Coryphoideae|Coryphoideae]].
| 
|-
|
''Trichotomosulcites normalis''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the family Arecaceae.
|
|-
|}

=====Commelinid monocot research=====
* A study on the evolutionary history of palms throughout the [[Earth:Cenozoic|Cenozoic]] era, aiming to determine the impact of Cenozoic environmental changes on the diversification and [[Earth:Biogeography|biogeography]] of palms, is published by Lim ''et al.'' (2021).<ref>{{Cite journal|last1=Lim |first1=J. Y. |last2=Huang |first2=H. |last3=Farnsworth |first3=A. |last4=Lunt |first4=D. J. |last5=Baker |first5=W. J. |last6=Morley |first6=R. J. |last7=Kissling |first7=W. D. |last8=Hoorn |first8=C. |year=2021 |title=The Cenozoic history of palms: Global diversification, biogeography and the decline of megathermal forests |journal=Global Ecology and Biogeography |volume=31 |issue=3 |pages=425–439 |doi=10.1111/geb.13436 |hdl=1983/fa5f36f6-5dbe-4109-bcaa-f9838a88e337 |s2cid=245284265 |url=https://research-information.bris.ac.uk/ws/files/308458806/Full_text_PDF_final_published_version_.pdf }}</ref>
* Pollens of member of the family [[Biology:Poaceae|Poaceae]] preserving the same morphological characteristics as that of modern cereal grains are described from a sedimentary core from Lake Acıgöl ([[Place:Turkey|Turkey]]) by Andrieu-Ponel ''et al.'' (2021), who interpret this finding as indicative of the presence of proto-cereals in [[Earth:Anatolia|Anatolia]] since 2.3 million years ago, likely evolving from wild Poaceae as a result of trampling, nitrogen enrichment of soils and browsing by large mammal herds, and evaluate possible benefits from the availability of these proto-cereals for early hominins.<ref>{{Cite journal|last1=Andrieu-Ponel |first1=V. |last2=Rochette |first2=P. |last3=Demory |first3=F. |last4=Alçiçek |first4=H. |last5=Boulbes |first5=N. |last6=Bourlès |first6=D. |last7=Helvacı |first7=C. |last8=Lebatard |first8=A.-E. |last9=Mayda |first9=S. |last10=Michaud |first10=H. |last11=Moigne |first11=A.-M. |last12=Nomade |first12=S. |last13=Perrin |first13=M. |last14=Ponel |first14=P. |last15=Rambeau |first15=C. |last16=Vialet |first16=A. |last17=Gambin |first17=B. |last18=Alçiçek |first18=M. C. |title=Continuous presence of proto-cereals in Anatolia since 2.3 Ma, and their possible co-evolution with large herbivores and hominins |year=2021 |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 8914 |doi=10.1038/s41598-021-86423-8 |pmid=33903602 |pmc=8076274 |bibcode=2021NatSR..11.8914A }}</ref>

===Basal eudicots===
==== Proteales ====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Platanocarpelia''<ref>{{Cite journal|last1=Maslova |first1=N. P. |last2=Kodrul |first2=T. M. |last3=Kachkina |first3=V. V. |title=Leaves of ''Ettingshausenia cuneifolia'' (Bronn) Stiehler (Angiospermae) and Associated Carpels and Stamens from the Turonian of Southern Kazakhstan |year=2021 |journal=Paleontological Journal |volume=55 |issue=10 |pages=1193–1214 |doi=10.1134/S0031030121100063 |s2cid=245540003 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Maslova, Kodrul & Kachkina
|
[[Earth:Late Cretaceous|Late Cretaceous]] ([[Earth:Turonian|Turonian]])
|
|
{{Flag|Kazakhstan}}
|
A member of the family [[Biology:Platanaceae|Platanaceae]]. Genus includes new species ''P. kyzyljarica''.
|
|-
|
''Proteacidites pseudodehaanii''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the family [[Biology:Proteaceae|Proteaceae]].
|
|-
|}
===Superasterids===
====Campanulid euasterids====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Pittosporum ettingshausenii''<ref name=Xenopanax>{{Cite journal|last=Doweld |first=A. B. |year=2021 |title=New names in ''Pittosporum'', extant and fossil (Pittosporaceae) |journal=Phytotaxa |volume=498 |issue=4 |pages=298–300 |doi=10.11646/phytotaxa.498.4.9 |s2cid=235540500 }}</ref>
|
Nom. nov
|
Valid
|
Doweld
|
Miocene
|
|
{{Flag|New Zealand}}
|
A species of ''[[Biology:Pittosporum|Pittosporum]]''; a replacement name for ''Pittosporum elegans'' (Ettingshausen) W.R.B. Oliver (1950).
|
|-
|
''Xenopanax''<ref name=Xenopanax />
|
Gen. et comb. nov
|
Valid
|
Doweld
|
[[Earth:Eocene|Eocene]]
|
|
{{Flag|Russia}}<br>({{Flag|Kamchatka Krai}})
|
A new genus for ''"Pittosporum" beringianum'' Chelebaeva & Akhmetiev (1983).
|
|-
|}

====Lamiid euasterids====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Adina vastanenesis''<ref>{{Cite journal|last1=Shukla |first1=A. |last2=Mehrotra |first2=R. C. |last3=Verma |first3=P. |last4=Chandra |first4=K. |last5=Singh |first5=A. |year=2021 |title="Out-of-India" dispersal for ''Adina'' (tribe Naucleeae; family Rubiaceae): evidence from the early Eocene fossil record from India |journal=Palaeoworld |volume=30 |issue=4 |pages=737–745 |doi=10.1016/j.palwor.2021.01.001 |s2cid=234244901 }}</ref>
|
Sp. nov
|
Valid
|
Shukla ''et al.''
|
Early [[Earth:Eocene|Eocene]]
|
Cambay Shale Formation
|
{{Flag|India}}
|
A species of ''[[Biology:Adina (plant)|Adina]]''.
| 
|-
|
''Dicolpopollis? costatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the genus ''[[Biology:Macoubea|Macoubea]]''.
|
|-
|
''Dolichandra pacei''<ref>{{Cite journal|last1=Franco |first1=M. J. |last2=Brea |first2=M. |last3=Cerdeño |first3=E. |year=2021 |title=First Bignoniaceae liana from the Miocene of South America and its evolutionary significance |journal=American Journal of Botany |volume=108 |issue=9 |pages=1761–1774 |doi=10.1002/ajb2.1736 |pmid=34591314 |s2cid=238230695 |doi-access=free |hdl=11336/184023 |hdl-access=free }}</ref>
|
Sp. nov
|
Valid
|
Franco, Brea & Cerdeño
|
Miocene ([[Earth:Santacrucian|Santacrucian]])
|
Mariño Formation
|
{{Flag|Argentina}}
|
A species of ''[[Biology:Dolichandra|Dolichandra]]''.
|
|-
|
''Fraxinoxylon beypazariense''<ref name=AP611 />
|
Sp. nov
|
Valid
|
Akkemik
|
Early Miocene
|
Hançili Formation
|
{{Flag|Turkey}}
|
A member of the family [[Biology:Oleaceae|Oleaceae]].
|
|-
|
''Fraxinus eoemarginata''<ref>{{Cite journal|last1=Mathewes |first1=R. |last2=Archibald |first2=S. B. |last3=Lundgren |first3=A. |year=2021 |title=Tips and identification of early Eocene ''Fraxinus'' L. samaras from the Quilchena locality, Okanagan Highlands, British Columbia, Canada |journal=Review of Palaeobotany and Palynology |volume=293 |pages=Article 104480 |doi=10.1016/j.revpalbo.2021.104480 |bibcode=2021RPaPa.29304480M }}</ref>
|
Sp. nov
|
In press
|
Mathewes, Archibald & Lundgren
|
Early [[Earth:Eocene|Eocene]]
|
[[Earth:Coldwater Beds|Quilchena site]]
|
{{Flag|Canada}}<br>({{Flag|British Columbia}})
|
A species of ''[[Biology:Fraxinus|Fraxinus]]''.
|
|-
|
''Kapgateophyllum''<ref>{{Cite journal|last=Deshmukh |first=U. B. |year=2021 |title=''Kapgateophyllum'' gen. nov. (Acanthaceae) |journal=Phytotaxa |volume=500 |issue=2 |pages=147–148 |doi=10.11646/phytotaxa.500.2.8 |s2cid=236585198 }}</ref>
|
Nom. nov
|
Valid
|
Deshmukh
|
Late Cretaceous (Maastrichtian) - early Eocene
|
[[Earth:Deccan Traps|Deccan]] [[Earth:Intertrappean Beds|Intertrappean Beds]]
|
{{Flag|India}}
|
A member of the family [[Biology:Acanthaceae|Acanthaceae]]; a replacement name for ''Acanthophyllum'' Ramteke & Kapgate (2014).
|
|-
|
''Ladakhipollenites campbellii''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the genus ''[[Biology:Vitex|Vitex]]''.
|
|-
|
''Lymingtonia splendida''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the family [[Biology:Convolvulaceae|Convolvulaceae]].
|
|-
|
''Maryendressantha''<ref>{{Cite journal|last1=Singh |first1=H. |last2=Judd |first2=W. S. |last3=Samant |first3=B. |last4=Agnihotri |first4=P. |last5=Grimaldi |first5=D. A. |last6=Manchester |first6=S. R. |year=2021 |title=Flowers of Apocynaceae in amber from the early Eocene of India |journal=American Journal of Botany |volume=108 |issue=5 |pages=883–892 |doi=10.1002/ajb2.1651 | issn=0002-9122 |pmid=34018178 |s2cid=235073745 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Singh ''et al.''
|
Early Eocene
|
Cambay amber
|
{{Flag|India}}
|
A member of the family [[Biology:Apocynaceae|Apocynaceae]]. Genus includes new species ''M. succinifera''.
|
|-
|
''Perfotricolpites hexacolpatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the genus ''[[Biology:Merremia|Merremia]]''.
|
|-
|
''Verrustephanoporites intraverrucosus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the family Apocynaceae.
|
|-
|}


====Non euasterids====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Anacolosidites reticulatus''<ref name=RPPHuangetal />
|
Sp. nov
|
In press
|
Morley, Huang & Hoorn ''in'' Huang ''et al.''
|
Middle and late Eocene
|
Yaw Formation
|
|
Pollen probably derived from plants belonging to the genus ''[[Biology:Ptychopetalum|Ptychopetalum]]''.
|
|-
|
''Camptotheca manchesterii''<ref>{{Cite journal|last1=Xie |first1=S.-P. |last2=Zhang |first2=S.-H. |last3=McElwain |first3=J. C. |last4=Zhang |first4=P. |last5=Wang |first5=B. |last6=Zhang |first6=Y. |last7=Yang |first7=Y.-H. |last8=Chen |first8=J.-Y. |year=2021 |title=First occurrence of ''Camptotheca'' fruits from late Miocene of southwestern China |journal=Historical Biology: An International Journal of Paleobiology |volume=33 |issue=12 |pages=3625–3632 |doi=10.1080/08912963.2021.1881782 |s2cid=234020768 }}</ref>
|
Sp. nov
|
In press
|
Xie ''et al.''
|
Late Miocene
|
Bangmai Formation
|
{{Flag|China}}
|
A species of ''[[Biology:Camptotheca|Camptotheca]]''.
| 
|-
|
''Diospyros christensenii''<ref>{{Cite journal|last1=Denk |first1=T. |last2=Bouchal |first2=J. M. |year=2021 |title=Dispersed pollen and calyx remains of ''Diospyros'' (Ebenaceae) from the middle Miocene "Plant beds" of Søby, Denmark |journal=GFF |volume=143 |issue=2–3 |pages=292–304 |doi=10.1080/11035897.2021.1907443 |bibcode=2021GFF...143..292D |s2cid=237648462 |url=http://urn.kb.se/resolve?urn=urn:nbn:se:nrm:diva-4502 }}</ref>
|
Sp. nov
|
Valid
|
Denk & Bouchal
|
Miocene
|
|
{{Flag|Denmark}}
|
A species of ''[[Biology:Diospyros|Diospyros]]''.
|
|-
|
''Halesia mosbruggeri''<ref>{{Cite journal|last=Kvaček |first=Z. |year=2021 |title=''Halesia mosbruggeri'' Kvaček, sp. nov., a new fossil fruit of ''Halesia'' L. (Styracaceae) from the Bohemian Miocene (Czech Republic) |journal=Palaeobiodiversity and Palaeoenvironments |volume=101 |issue=1 |pages=75–78 |doi=10.1007/s12549-020-00463-y |s2cid=232127283 }}</ref>
|
Sp. nov
|
Valid
|
Kvaček
|
Early Miocene
|
Most Basin
|
{{Flag|Czech Republic}}
|
A species of ''[[Biology:Halesia|Halesia]]''.
|
|-
|
''Loranthacites tabatingensis''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the genus ''[[Biology:Struthanthus|Struthanthus]]''.
|
|-
|
''Mecsekispermum''<ref>{{Cite journal|last1=Erdei |first1=B. |last2=Hably |first2=L. |year=2021 |title=Fossil ''Gordonia'' (s.l.)–like (Theaceae) winged seeds from the early Miocene of the Mecsek Mts, W Hungary |journal=Palaeobiodiversity and Palaeoenvironments |volume=101 |issue=1 |pages=59–67 |doi=10.1007/s12549-020-00461-0 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Hably & Erdei
|
Miocene ([[Earth:Burdigalian|Burdigalian]])
|
Feked Formation
|
{{Flag|Hungary}}
|
Possibly a member of the family [[Biology:Theaceae|Theaceae]]. Genus includes new species ''M. gordonioides''.
| 
|-
|
''Miranthus''<ref>{{Cite journal|last1=Friis |first1=E. M. |last2=Crane |first2=P. R. |last3=Pedersen |first3=K. R. |title=Early flowers of primuloid Ericales from the Late Cretaceous of Portugal and their ecological and phytogeographic implications |year=2021 |journal=Fossil Imprint |volume=77 |issue=2 |pages=214–230 |doi=10.37520/fi.2021.016 |s2cid=245553031 |url=http://fi.nm.cz/en/clanek/early-flowers-of-primuloid-ericales-from-the-late-cretaceous-of-portugal-and-their-ecological-and-phytogeographic-implications-2/ |doi-access=free }}</ref>
|
Gen. et 2 sp. nov
|
Valid
|
Friis, Crane
|
[[Earth:Late Cretaceous|Late Cretaceous]] ([[Earth:Campanian|Campanian]]-[[Earth:Maastrichtian|Maastrichtian]])
|
|
{{Flag|Portugal}}
|
A [[Biology:Primulaceae|Primulaceae]] genus. Genus includes new species ''M. elegans'' and ''M. kvacekii''.
|
|-
|
''Nyssa nanningensis''<ref>{{Cite journal|last1=Xu |first1=S.-L. |last2=Kodrul |first2=T. M. |last3=Maslova |first3=N. P. |last4=Song |first4=H.-Z. |last5=Tobias |first5=A. V. |last6=Wu |first6=X.-K. |last7=Quan |first7=C. |last8=Jin |first8=J.-H. |year=2021 |title=First occurrence of ''Nyssa'' endocarps and associated fungi in the Oligocene of South China: palaeogeographical and palaeoecological significance |journal=Papers in Palaeontology |volume=8 |doi=10.1002/spp2.1408 |s2cid=244057027 }}</ref>
|
Sp. nov
|
In press
|
Xu & Jin ''in'' Xu ''et al.''
|
Late Oligocene
|
Yongning Formation
|
{{Flag|China}}
|
A [[Biology:Tupelo|tupelo]].
|
|-
|
''Paranyssa''<ref name=Allonymphaea />
|
Nom. nov
|
Valid
|
Doweld
|
Paleocene
|
|
{{Flag|United States}}<br>({{Flag|Montana}})
|
A member of the family [[Biology:Nyssaceae|Nyssaceae]]; a replacement name for ''Browniea'' Manchester & Hickey (2007).
|
|-
|
''Parsonsidites? minibrenacii''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the family [[Biology:Amaranthaceae|Amaranthaceae]].
|
|-
|
''Ternstroemites klettwitzensis''<ref name=FI771Striegler>{{Cite journal|last=Striegler |first=U. |title=New leaf species from the upper Miocene flora of the leaf-bearing Wischgrund clay (Lower Lusatia, Brandenburg, Germany) |year=2021 |journal=Fossil Imprint |volume=77 |issue=1 |pages=102–110 |doi=10.37520/fi.2021.009 |s2cid=245028668 |url=http://fi.nm.cz/en/clanek/new-leaf-species-from-the-upper-miocene-flora-of-the-leaf-bearing-wischgrund-clay-lower-lusatia-brandenburg-germany-2/ |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Striegler
|
Miocene ([[Biology:Tortonian|Tortonian]])
|
Rauno Formation
|
{{Flag|Germany}}
|
A member of the family Theaceae.
|
|-
|}

===Superrosids===
====Fabids====
=====Fabales=====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Albizia mahuadanrensis''<ref name=HazraetalAlbizia>{{Cite journal|last1=Hazra |first1=T. |last2=Hazra |first2=M. |last3=Spicer |first3=R. A. |last4=Spicer |first4=T. E. V. |last5=Mahato |first5=S. |last6=Bera |first6=S. |last7=Kumar |first7=S. |last8=Khan |first8=M. A. |year=2021 |title=Pliocene ''Albizia'' (Fabaceae) from Jharkhand, eastern India: reappraisal of its biogeography during Cenozoic in Southeast Asia |journal=Palaeoworld |volume=31 |pages=153–168 |doi=10.1016/j.palwor.2021.03.004 |s2cid=233521737 |url=http://oro.open.ac.uk/75930/1/1-s2.0-S1871174X21000263-main.pdf }}</ref>
|
Sp. nov
|
In press
|
Hazra, Hazra & Khan ''in'' Hazra ''et al.''
|
Pliocene
|
Rajdanda Formation
|
{{Flag|India}}
|
A species of ''[[Biology:Albizia|Albizia]]''.
|
|-
|
''Albizia palaeoprocera''<ref name=HazraetalAlbizia />
|
Sp. nov
|
In press
|
Hazra, Hazra & Khan ''in'' Hazra ''et al.''
|
Pliocene
|
Rajdanda Formation
|
{{Flag|India}}
|
A species of ''Albizia''.
|
|-
|
''Cercis zhangpuensis''<ref>{{Cite journal|last1=Wang |first1=Z. |last2=Shi |first2=G. |last3=Sun |first3=B. |last4=Jia |first4=H. |last5=Dong |first5=C. |last6=Yin |first6=S. |last7=Wu |first7=X. |year=2021 |title=A new ''Cercis'' (Leguminosae) from the middle Miocene of Fujian, China |journal=Historical Biology: An International Journal of Paleobiology |volume=34 |pages=94–101 |doi=10.1080/08912963.2021.1900170 |s2cid=233645060 }}</ref>
|
Sp. nov
|
In press
|
Wang ''et al.''
|
Miocene
|
Fotan Group
|
{{Flag|China}}
|
A species of ''[[Biology:Cercis|Cercis]]''.
|
|-
|
''Cladrastis haominiae''<ref>{{Cite journal|last1=Jia |first1=L.-B. |last2=Huang |first2=J. |last3=Su |first3=T. |last4=Spicer |first4=R. A. |last5=Zhang |first5=S.-T. |last6=Li |first6=S.-F. |last7=Pan |first7=B. |last8=Nam |first8=G.-S. |last9=Huang |first9=Y.-J. |last10=Zhou |first10=Z.-K. |year=2021 |title=Fossil infructescence from southwestern China reveals Paleogene establishment of ''Cladrastis'' in Asia |journal=Review of Palaeobotany and Palynology |volume=292 |pages=Article 104456 |doi=10.1016/j.revpalbo.2021.104456 |bibcode=2021RPaPa.29204456J |s2cid=236255120 |url=http://oro.open.ac.uk/76470/1/1-s2.0-S0034666721000804-main.pdf }}</ref>
|
Sp. nov
|
In press
|
Jia ''et al.''
|
[[Earth:Paleogene|Paleogene]]
|
|
{{Flag|China}}
|
A species of ''[[Biology:Cladrastis|Cladrastis]]''.
|
|-
|
''Enterolobiumoxylon''<ref name=Enterolobiumoxylon>{{Cite journal|last1=Pérez-Lara |first1=D. K. |last2=Estrada-Ruiz |first2=E. |last3=Castañeda-Posadas |first3=C. |year=2021 |title=''Kingiodendron'' and ''Enterolobium'' Eocene woods from the El Bosque formation, Chiapas, Mexico |journal=Journal of South American Earth Sciences |volume=111 |pages=Article 103477 |doi=10.1016/j.jsames.2021.103477 |bibcode=2021JSAES.11103477P |s2cid=237819942 }}</ref>
|
Gen. et sp. nov
|
In press
|
Pérez-Lara, Estrada-Ruiz & Castañeda-Posadas
|
[[Earth:Eocene|Eocene]]
|
El Bosque Formation
|
{{Flag|Mexico}}
|
Fossil wood of a member of the family [[Biology:Fabaceae|Fabaceae]]. Genus includes new species ''E. triserial''.
|
|-
|
''Gleditsioxylon fiambalense''<ref>{{Cite journal|last=Baez |first=J. |year=2021 |title=First fossil record in Tambería formation (Neogene) in Bolsón de Fiambalá, Catamarca province, Argentina: Palaeoenvironmental inferences through Leguminosae woods |journal=Journal of South American Earth Sciences |volume=110 |pages=Article 103403 |doi=10.1016/j.jsames.2021.103403 |bibcode=2021JSAES.11003403B }}</ref>
|
Sp. nov
|
In press
|
Baez
|
Miocene
|
Tambería Formation
|
{{Flag|Argentina}}
|
A member of Leguminosae.
|
|-
|
''Kingiodendron mexicanus''<ref name=Enterolobiumoxylon />
|
Sp. nov
|
In press
|
Pérez-Lara, Estrada-Ruiz & Castañeda-Posadas
|
Eocene
|
El Bosque Formation
|
{{Flag|Mexico}}
|
Fossil wood of a member of the family Fabaceae.
|
|-
|
''Ladakhipollenites? pseudocolpiconstrictus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the family [[Biology:Fabaceae|Fabaceae]].
|
|-
|
''Leguminocarpum lottii''<ref>{{Cite journal|last1=Li |first1=X. |last2=Manchester |first2=S. R. |last3=Correa-Narvaez |first3=J. E. |last4=Herendeen |first4=P. S. |year=2021 |title=An Extinct Fruit Species of Fabaceae from the Early Eocene of Northwestern Wyoming, USA |journal=International Journal of Plant Sciences |volume=182 |issue=8 |pages=730–746 |doi=10.1086/715634 |s2cid=237505717 }}</ref>
|
Sp. nov
|
Valid
|
Li & Manchester ''in'' Li ''et al.''
|
Early [[Earth:Eocene|Eocene]]
|
Tepee Trail Formation
|
{{Flag|United States}}<br>({{Flag|Wyoming}})
|
A member of the family [[Biology:Fabaceae|Fabaceae]].
|
|-
|
''Leguminocarpum olmensis''<ref>{{Cite journal|last1=Centeno-González |first1=N. K. |last2=Martínez-Cabrera |first2=H. I. |last3=Porras-Múzquiz |first3=H. |last4=Estrada-Ruiz |first4=E. |title=Late Campanian fossil of a legume fruit supports Mexico as a center of Fabaceae radiation |year=2021 |journal=Communications Biology |volume=4 |issue=1 |pages=Article number 41 |doi=10.1038/s42003-020-01533-9 |pmid=33446929 |pmc=7809014 }}</ref>
|
Sp. nov
|
Valid
|
Centeno-González ''et al.''
|
[[Earth:Late Cretaceous|Late Cretaceous]] ([[Earth:Campanian|Campanian]])
|
[[Earth:Olmos Formation|Olmos Formation]]
|
{{Flag|Mexico}}
|
A member of the family Fabaceae.
| 
|-
|
''Neopapilionia''<ref>{{Cite journal|last1=Hazra |first1=T. |last2=Hazra |first2=M. |last3=Bera |first3=S. |last4=Khan |first4=M. A. |year=2021 |title=First Fossil Legume Flower of Papilionoid Affinity from India |journal=Journal of the Geological Society of India |volume=97 |issue=3 |pages=267–270 |doi=10.1007/s12594-021-1677-3 |issn=0016-7622 |s2cid=232164301 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Hazra, Hazra & Khan ''in'' Hazra ''et al.''
|
Pliocene
|
Rajdanda Formation
|
{{Flag|India}}
|
A member of the family Fabaceae. Genus includes new species ''N. indica''.
| 
|-
|
''Ormosia cyclocarpa''<ref>{{Cite journal|last1=Li |first1=X.-C. |last2=Manchester |first2=S. R. |last3=Xiao |first3=L. |last4=Wang |first4=Q. |last5=Hu |first5=Y. |last6=Sun |first6=B.-N. |year=2021 |title=''Ormosia'' (Fabaceae: Faboideae) from the Miocene of southeastern China support historical expansion of the tropical genus in East Asia |journal=Historical Biology: An International Journal of Paleobiology |volume=33 |issue=12 |pages=3561–3578 |doi=10.1080/08912963.2021.1877700 |s2cid=233806537 }}</ref>
|
Sp. nov
|
In press
|
Li ''et al.''
|
Miocene
|
|
{{Flag|China}}
|
A species of ''[[Biology:Ormosia|Ormosia]]''.
| 
|-
|
''Peltophorum asiatica''<ref>{{Cite journal|last1=Hazra |first1=T. |last2=Hazra |first2=M. |last3=Bera |first3=S. |last4=Khan |first4=M. A. |year=2021 |title=First fossil evidence of marginal winged fruits of ''Peltophorum'' (Caesalpinioideae: Fabaceae) from India |journal=Brittonia |volume=73 |issue=3 |pages=241–250 |doi=10.1007/s12228-021-09679-4 |s2cid=238855320 }}</ref>
|
Sp. nov
|
In press
|
Hazra, Hazra & Khan ''in'' Hazra ''et al.''
|
Pliocene
|
Rajdanda Formation
|
{{Flag|India}}
|
A species of ''[[Biology:Peltophorum|Peltophorum]]''.
|
|-
|
''Polyadopollenites minimus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the genus ''[[Biology:Mimosa|Mimosa]]''.
|
|-
|
''Salpinganthium''<ref>{{Cite journal|last1=Poinar |first1=G. O. |last2=Chambers |first2=K. L. |year=2021 |title=''Salpinganthium hispaniolanum'' gen. et sp. nov. (Fabaceae: Detarieae), a mid-Tertiary flower in Dominican amber |journal=Journal of the Botanical Research Institute of Texas |volume=15 |issue=2 |pages=559–567 |doi=10.17348/jbrit.v15.i2.1161 |s2cid=245099015 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Poinar & Chambers
|
[[Earth:Burdigalian|Burdigalian]]
|
[[Earth:Dominican amber|Dominican amber]]
|
{{Flag|Dominican Republic}}
|
A member of the family [[Biology:Fabaceae|Fabaceae]] belonging to the tribe [[Biology:Detarieae|Detarieae]]. Genus includes new species ''S. hispaniolanum''.
|
|-
|
''Striatopollis crassitectatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the genus ''[[Biology:Macrolobium|Macrolobium]]''.
|
|-
|
''Syncolporites foveolatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the genus ''[[Biology:Swartzia|Swartzia]]''.
|
|-
|}

=====Fagales=====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Eucarpinoxylon kayacikii''<ref name=AP611 />
|
Sp. nov
|
Valid
|
Akkemik
|
Middle Miocene
|
Kesmekaya Volcanics
|
{{Flag|Turkey}}
|
A member of the family [[Biology:Betulaceae|Betulaceae]].
|
|-
|
''Fagus dodgei''<ref name="Posthammer2021">{{cite journal |last1=Wheeler |first1=E. A. |last2=Manchester |first2=S. R. |title=A diverse assemblage of Late Eocene woods from Oregon, western USA |journal=Fossil Imprint |date=2021 |volume=77 |issue=2 |pages=299–329 |doi=10.37520/fi.2021.022|s2cid=246981608 |doi-access=free }}</ref>
|
Sp nov
|
Valid
|
Wheeler & Manchester
|
Late Eocene
|
John Day Formation
|
{{Flag|USA}}<br/>{{Flag|Oregon}}
|
A ''[[Biology:Fagus|Fagus]]'' species wood.
|
|-
|
''Lithocarpoxylon ashwillii''<ref name="Posthammer2021"/>
|
Sp nov
|
Valid
|
Wheeler & Manchester
|
Late Eocene
|
John Day Formation
|
{{Flag|USA}}<br/>{{Flag|Oregon}}
|
A ''Lithocarpoxylon'' species wood.
|
|-
|
''Morella absarokensis''<ref name="Fagaleswood">{{Cite journal|last1=Wheeler |first1=E. A. |last2=Baas |first2=P. |last3=Manchester |first3=S. R. |year=2021 |title=Wood Anatomy of Modern and Fossil Fagales in Relation to Phylogenetic Hypotheses, Familial Classification, and Patterns of Character Evolution |journal=International Journal of Plant Sciences |volume=183 |issue=1 |pages=61–86 |doi=10.1086/717328 |s2cid=244081754 |doi-access=free }}</ref>
|
Comb nov
|
valid
|
(Wheeler, Scott, & Barghoorn) Wheeler & Manchester
|
Middle Eocene 
|
Lamar River Formation
|
{{Flag|USA}}<br/>{{Flag|Wyoming}}
|
Moved from ''Myrica absarokensis'' (1978)
|
|-
|
''Morella scalariformis''<ref name="Fagaleswood"/>
|
Comb nov
|
valid
|
(Kruse) Wheeler, Baas, & Manchester
|
Middle Eocene 
|
Eden Valley Formation
|
{{Flag|USA}}<br/>{{Flag|Wyoming}}
|
Moved from ''Myrica scalariformis'' (1954)
|
|-
|
''Myricamentum''<ref>{{Cite journal|last1=Wilde |first1=V. |last2=Frankenhäuser |first2=H. |last3=Lenz |first3=O. K. |year=2021 |title=A myricaceous male inflorescence with pollen in situ from the middle Eocene of Europe |journal=Palaeobiodiversity and Palaeoenvironments |volume=101 |issue=4 |pages=873–883 |doi=10.1007/s12549-020-00479-4 |s2cid=233417128 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Wilde, Frankenhäuser & Lenz
|
[[Earth:Eocene|Eocene]]
|
Eckfelder Maar
|
{{Flag|Germany}}
|
A [[Biology:Catkin|catkin]]-like male [[Biology:Inflorescence|inflorescence]], probably of [[Biology:Myricaceae|myricaceous]] affinity. Genus includes new species ''M. eckfeldensis''.
|
|-
|
''Ostryoxylon gokceadaense''<ref name=AP611 />
|
Sp. nov
|
Valid
|
Akkemik
|
Middle Miocene
|
Kesmekaya Volcanics
|
{{Flag|Turkey}}
|
A member of the family Betulaceae.
|
|-
|
''Palaeocarpinus borealis''<ref name="Palaeocarpinus">{{cite journal |last1=Correa-Narvaez |first1=J. E. |last2=Manchester |first2=S. R. |title=Distribution and Morphological Diversity of Palaeocarpinus (Betulaceae) from the Paleogene of the Northern Hemisphere. |journal=The Botanical Review |date=2021 |volume=88 |issue=2 |pages=161–203 |doi=10.1007/s12229-021-09258-y|s2cid=237795532 }}</ref>
|
Comb nov
|
Valid
|
(Heer)
|
[[Earth:Paleocene|Paleocene]]
|
[[Earth:Menat Formation|Menat Formation]]
|
{{Flag|France}}
|
A [[Biology:Coryloideae|Coryloideae]] species.<br/>Moved from ''Atriplex borealis'' (1912)
|
|-
|
''Palaeocarpinus parva''<ref name="Palaeocarpinus"/>
|
Sp. nov
|
Valid
|
Manchester & Correa
|
Middle Eocene
|
Clarno Formation
|
{{Flag|USA}}<br/>{{Flag|Oregon}}
|
A [[Biology:Coryloideae|Coryloideae]] species.
|
|-
|
''Palaeocarpinus pteravestigia''<ref name="Palaeocarpinus"/>
|
Sp. nov
|
Valid
|
Correa & Manchester
|
[[Earth:Paleocene|Paleocene]]
|
[[Earth:Fort Union Formation|Fort Union Formation]]<br/>"unnamed lower member"
|
{{Flag|USA}}<br/>{{Flag|Wyoming}}
|
A [[Biology:Coryloideae|Coryloideae]] species.
|
|-
|
''Palaeocarya indica''<ref>{{Cite journal|last1=Hazra |first1=T. |last2=Hazra |first2=M. |last3=Kumar |first3=S. |last4=Mahato |first4=S. |last5=Bera |first5=M. |last6=Bera |first6=S. |last7=Khan |first7=M. A. |year=2021 |title=First fossil evidence of ''Palaeocarya'' (Engelhardioideae: Juglandaceae) from India and its biogeographical implications |journal=Journal of Systematics and Evolution |volume=59 |issue=6 |pages=1307–1320 |doi=10.1111/jse.12736 |s2cid=233897694 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Hazra, Hazra & Khan ''in'' Hazra ''et al.''
|
Pliocene
|
Rajdanda Formation
|
{{Flag|India}}
|
A member of the family [[Biology:Juglandaceae|Juglandaceae]].
| 
|-
|
''Paralnoxylon''<ref name=Allonymphaea />
|
Nom. nov
|
Valid
|
Doweld
|
Paleocene
|
|
{{Flag|United Kingdom}}
|
A member of the family Betulaceae; a replacement name for ''Cantia'' Stopes (1915).
|
|-
|
''Quercoxylon yaltirikii''<ref name=AP611 />
|
Sp. nov
|
Valid
|
Akkemik
|
Early Miocene
|
Hançili Formation
|
{{Flag|Turkey}}
|
A member of the family [[Biology:Fagaceae|Fagaceae]].
|
|-
|}

======Fagalean research======
* A study on wood anatomy in extant and fossil members of Fagales is published by Wheeler, Baas & Manchester who transfer two Eocene species from ''Myrica'' to ''Morella''.<ref name="Fagaleswood"/>

=====Malpighiales=====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Euphorbiotheca deccanensis''<ref>{{Cite journal|last1=Reback |first1=R. G. |last2=Kapgate |first2=D. K. |last3=Wurdack |first3=K. |last4=Manchester |first4=S. R. |year=2021 |title=Fruits of Euphorbiaceae from the Late Cretaceous Deccan Intertrappean Beds of India |journal=International Journal of Plant Sciences |volume=183 |issue=2 |page=000 |doi=10.1086/717691 |s2cid=239507275 }}</ref>
|
Sp. nov
|
In press
|
Reback ''et al.''
|
Late Cretaceous (Maastrichtian)
|
Deccan Intertrappean Beds
|
{{Flag|India}}
|
A member of the family [[Biology:Euphorbiaceae|Euphorbiaceae]].
|
|-
|
''Passiflora appalachiana''<ref>{{Cite journal|last=Hermsen |first=E. J. |year=2021 |title=Review of the fossil record of ''Passiflora'', with a description of new seeds from the Pliocene Gray Fossil Site, Tennessee, U.S.A. |journal=International Journal of Plant Sciences |volume=182 |issue=6 |pages=533–550 |doi=10.1086/714282 |s2cid=233701901 }}</ref>
|
Sp. nov
|
Valid
|
Hermsen
|
Pliocene
|
Gray Fossil Site
|
{{Flag|United States}}<br>({{Flag|Tennessee}})
|
A species of ''[[Biology:Passiflora|Passiflora]]''.
| 
|-
|
''Passifloriidites''<ref name=Passifloriidites>{{Cite journal|last1=D'Apolito |first1=C. |last2=Jaramillo |first2=C. |last3=Harrington |first3=G. |year=2021 |title=Miocene Palynology of the Solimões Formation (Well 1-AS-105-AM), Western Brazilian Amazonia |journal=Smithsonian Contributions to Paleobiology |volume=105 |issue=105 |pages=iv-134 |doi=10.5479/si.16803493 |s2cid=243391850 |url=https://figshare.com/articles/book/Miocene_Palynology_of_the_Solim_es_Formation_Well_1-AS-105-AM_Western_Brazilian_Amazonia/16803493 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the family [[Biology:Passifloraceae|Passifloraceae]]. Genus includes new species ''P. pseudoperculatus''.
|
|-
|
''Populoxylon sebenense''<ref name=AP611 />
|
Sp. nov
|
Valid
|
Akkemik
|
Early Miocene
|
Hançili Formation
|
{{Flag|Turkey}}
|
A member of the family [[Biology:Salicaceae|Salicaceae]].
|
|-
|
''Salicoxylon galatianum''<ref name=AP611 />
|
Sp. nov
|
Valid
|
Akkemik
|
Early Miocene
|
Hançili Formation
|
{{Flag|Turkey}}
|
A member of the family Salicaceae.
|
|-
|
''Verrustephanoporites circularis''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the genus ''[[Biology:Mascagnia|Mascagnia]]''.
|
|-
|}
=====Oxalidales=====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Tropidogyne euthystyla''<ref>{{Cite journal|last1=Poinar |first1=G. O. |last2=Chambers |first2=K. L. |last3=Vega |first3=F. E. |year=2021 |title=''Tropidogyne euthystyla'' sp. nov., a new small-flowered addition to the genus from mid-Cretaceous Myanmar amber |journal=Journal of the Botanical Research Institute of Texas |volume=15 |issue=1 |pages=113–119 |doi=10.17348/jbrit.v15.i1.1053 |s2cid=237700194 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Poinar, Chambers & Vega
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A possible member of [[Biology:Cunoniaceae|Cunoniaceae]].
|
|-
|}
=====Rosales=====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Alloceltidoxylon''<ref name=Allonymphaea />
|
Nom. nov
|
Valid
|
Doweld
|
Eocene
|
Clarno Formation
|
{{Flag|United States}}<br>({{Flag|Oregon}})
|
A flowering plant with possible affinities with [[Biology:Urticales|urticalean rosids]]; a replacement name for ''Scottoxylon'' Wheeler & Manchester (2002).
|
|-
|
''Celtis popsii''<ref name="Posthammer2021"/>
|
Sp nov
|
Valid
|
Wheeler & Manchester
|
Late Eocene
|
John Day Formation
|
{{Flag|USA}}<br/>{{Flag|Oregon}}
|
A ''[[Biology:Celtis|Celtis]]'' species wood.
|
|-
|
''Crataegoxylon sibiricum''<ref name=Thujopsoxylon />
|
Sp. nov
|
Valid
|
Dolezych, LePage & Williams
|
Oligocene ([[Earth:Chattian|Chattian]])
|
Korlikov Formation
|
{{Flag|Russia}}<br>({{Flag|Tomsk Oblast}})
|
|
|-
|
''Morus asiatica''<ref>{{Cite journal|last1=Patel |first1=R. |last2=Hazra |first2=T. |last3=Rana |first3=R. S. |last4=Hazra |first4=M. |last5=Bera |first5=S. |last6=Khan |first6=M. A. |year=2021 |title=First fossil record of mulberry from Asia |journal=Review of Palaeobotany and Palynology |volume=292 |pages=Article 104459 |doi=10.1016/j.revpalbo.2021.104459 |bibcode=2021RPaPa.29204459P }}</ref>
|
Sp. nov
|
In press
|
Patel, Rana & Khan ''in'' Patel ''et al.''
|
Early [[Earth:Eocene|Eocene]]
|
|
{{Flag|India}}
|
A species of ''[[Biology:Morus (plant)|Morus]]''.
|
|-
|
''Prunoidoxylon prunoides''<ref name=AP611 />
|
Sp. nov
|
Valid
|
Akkemik
|
Early Miocene
|
Hançili Formation
|
{{Flag|Turkey}}
|
A member of the family [[Biology:Rosaceae|Rosaceae]].
|
|-
|
''Psilatriporites minimus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the genus ''[[Biology:Celtis|Celtis]]''.
|
|-
|
''Pyracantha pseudococcinea''<ref name=FI771Striegler />
|
Sp. nov
|
Valid
|
Striegler
|
Miocene (Tortonian)
|
Rauno Formation
|
{{Flag|Germany}}
|
A species of ''[[Biology:Pyracantha|Pyracantha]]''.
|
|-
|
''Ulmoxylon kasapligilii''<ref name=AP611 />
|
Sp. nov
|
Valid
|
Akkemik
|
Early Miocene
|
Hançili Formation
|
{{Flag|Turkey}}
|
A member of the family [[Biology:Ulmaceae|Ulmaceae]].
|
|-
|
''Ventilago tibetensis''<ref>{{Cite journal|last1=Del Rio |first1=C. |last2=Wang |first2=T.-X. |last3=Xu |first3=X.-T. |last4=Sabroux |first4=R. |last5=Spicer |first5=T. E. V. |last6=Liu |first6=J. |last7=Chen |first7=P.-R. |last8=Wu |first8=F.-X. |last9=Zhou |first9=Z.-K. |last10=Su |first10=T. |year=2021 |title=''Ventilago'' (Rhamnaceae) Fruit from the Middle Eocene of the Central Tibet, China |journal=International Journal of Plant Sciences |volume=182 |issue=7 |pages=638–648 |doi=10.1086/715507 |s2cid=236410986 }}</ref>
|
Sp. nov
|
Valid
|
Del Rio ''et al.''
|
Middle Eocene
|
|
{{Flag|China}}
|
A species of ''[[Biology:Ventilago|Ventilago]]''.
|
|-
|
''Zelkovoxylon crystalliferum''<ref name=AP611 />
|
Sp. nov
|
Valid
|
Akkemik
|
Early Miocene
|
Hançili Formation
|
{{Flag|Turkey}}
|
A member of the family Ulmaceae.
|
|-
|}

====Malvids====
=====Malvales=====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Bombacacidites hooghiemstrae''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the family [[Biology:Malvaceae|Malvaceae]].
|
|-
|
''Craigia lincangensis''<ref>{{Cite journal|last1=Wang |first1=B. |last2=Zhang |first2=S. |last3=Zhang |first3=P. |last4=Yang |first4=Y. |last5=Chen |first5=J. |last6=Zhang |first6=Y. |last7=Xie |first7=S. |year=2021 |title=A new occurrence of ''Craigia'' (Malvaceae) from the Miocene of Yunnan and its biogeographic significance |journal=Historical Biology: An International Journal of Paleobiology |volume=33 |issue=12 |pages=3402–3412 |doi=10.1080/08912963.2020.1867980 |s2cid=234188949 }}</ref>
|
Sp. nov
|
In press
|
Wang & Xie ''in'' Wang ''et al.''
|
Late Miocene
| 
|
{{Flag|China}}
|
A species of ''[[Biology:Craigia|Craigia]]''
| 
|-
|
''Dipterocapus fotanensis''<ref>{{Cite journal|last1=Chen |first1=J. |last2=Han |first2=L. |last3=Hua |first3=Y. |last4=Wu |first4=G. |last5=Sun |first5=B. |year=2021 |title=Fruits and leaves of ''Dipterocarpus'' from the Miocene of Zhangpu, Fujian, and its geological significance |journal=Arabian Journal of Geosciences |volume=14 |issue=13 |pages=Article 1270 |doi=10.1007/s12517-021-07445-0 |s2cid=235664238 }}</ref>
|
Sp. nov
|
Valid
|
Chen ''et al.''
|
Miocene
| 
|
{{Flag|China}}
|
A species of ''[[Biology:Dipterocarpus|Dipterocarpus]]''
|
|-
|
''Discoidites angulosus''<ref name=RPPHuangetal>{{Cite journal|last1=Huang |first1=H. |last2=Pérez-Pinedo |first2=D. |last3=Morley |first3=R. J. |last4=Dupont-Nivet |first4=G. |last5=Philip |first5=A. |last6=Win |first6=Z. |last7=Aung |first7=D. W. |last8=Licht |first8=A. |last9=Jardine |first9=P. E. |last10=Hoorn |first10=C. |year=2021 |title=At a crossroads: The late Eocene flora of central Myanmar owes its composition to plate collision and tropical climate |journal=Review of Palaeobotany and Palynology |volume=291 |pages=Article 104441 |doi=10.1016/j.revpalbo.2021.104441 |bibcode=2021RPaPa.29104441H |s2cid=235421153 |doi-access=free |hdl=11245.1/6fb3c1f3-e28b-4a90-9d4f-dbe38f07a512 |hdl-access=free }}</ref>
|
Sp. nov
|
In press
|
Huang, Morley & Hoorn ''in'' Huang ''et al.''
|
Late [[Earth:Eocene|Eocene]]
|
Yaw Formation
|
{{Flag|Myanmar}}
|
Pollen probably derived from plants belonging to the genus ''[[Biology:Brownlowia|Brownlowia]]''.
|
|-
|
''Thymelipollis amazonicus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the family [[Biology:Thymelaeaceae|Thymelaeaceae]].
|
|-
|
''Tilia asiatica''<ref>{{Cite journal|last1=Jia |first1=L.B. |last2=Nam |first2=G.S. |last3=Su |first3=T. |last4=Stull |first4=G.W. |last5=Li |first5=S.F. |last6=Huang |first6=Y.J. |last7=Zhou |first7=Z.K. |year=2021 |title=Fossil fruits of ''Firmiana'' and ''Tilia'' from the middle Miocene of South Korea and the efficacy of the Bering land bridge for the migration of mesothermal plants |journal=Plant Diversity |volume=43 |issue=6 |pages=480–491 |doi=10.1016/j.pld.2020.12.006 |pmid=35024517 |pmc=8720707 |doi-access=free }}</ref>
|
Sp. nov
|
In press
|
Jia & Nam ''in'' Jia ''et al.''
|
Middle Miocene
|
Pohang Basin
|
{{Flag|South Korea}}
|
A species of ''[[Biology:Tilia|Tilia]]''
| 
|-
|
''Wataria kvacekii''<ref name="Posthammer2021"/>
|
Sp nov
|
Valid
|
Wheeler & Manchester
|
Late Eocene
|
John Day Formation
|
{{Flag|USA}}<br/>{{Flag|Oregon}}
|
A ''Wataria'' species wood.
|
|-
|}

=====Myrtales=====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Florschuetzia impostora''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen, possibly of a member of the family [[Biology:Lythraceae|Lythraceae]].
|
|-
|
''Lazarocardenasoxylon''<ref>{{Cite journal|last1=Estrada-Ruiz |first1=E. |last2=Martínez-Cabrera |first2=H. I. |last3=García-Hernández |first3=I. P. |year=2021 |title=New dicotyledonous woods from the San Carlos Formation (Upper Cretaceous) in Northern Mexico |journal=IAWA Journal |volume=43 |issue=1–2 |pages=66–79 |doi=10.1163/22941932-bja10079 |s2cid=244902117 }}</ref>
|
Gen. et sp. nov
|
In press
|
Estrada-Ruiz & Martínez-Cabrera ''in'' Estrada-Ruiz, Martínez-Cabrera & García-Hernández
|
[[Earth:Late Cretaceous|Late Cretaceous]]
|
[[Earth:San Carlos Formation|San Carlos Formation]]
|
{{Flag|Mexico}}
|
Possibly a member of the family [[Biology:Myrtaceae|Myrtaceae]]. Genus includes new species ''L. aldamense''.
|
|-
|
''Lythrum portugalliense''<ref>{{Cite journal|last1=Vieira |first1=M. |last2=Zetter |first2=R. |last3=Coiro |first3=M. |last4=Grímsson |first4=F. |year=2021 |title=Pliocene ''Lythrum'' (loosestrife, Lythraceae) pollen from Portugal and the Neogene establishment of European lineages |journal=Review of Palaeobotany and Palynology |volume=296 |pages=Article 104548 |doi=10.1016/j.revpalbo.2021.104548 | issn=0034-6667 |doi-access=free |hdl=10362/144398 |hdl-access=free }}</ref>
|
Sp. nov
|
In press
|
Vieira ''et al.''
|
Pliocene (Piacenzian)
|
|
{{Flag|Portugal}}
|
A species of ''[[Biology:Lythrum|Lythrum]]''.
|
|-
|
''Syzygium guipingensis''<ref>{{Cite journal|last1=Li |first1=Y. |last2=Huang |first2=L. |last3=Quan |first3=C. |last4=Jin |first4=J. |last5=Oskolski |first5=A. A. |year=2021 |title=Fossil wood of ''Syzygium'' from the Miocene of Guangxi, South China: the earliest fossil evidence of the genus in eastern Asia |journal=IAWA Journal |volume=42 |issue=4 |pages=435–441 |doi=10.1163/22941932-bja10069|issn=0928-1541  |s2cid=240550082 }}</ref>
|
Sp. nov
|
In press
|
Li ''et al.''
|
Miocene
|
Erzitang Formation
|
{{Flag|China}}
|
A species of ''[[Biology:Syzygium|Syzygium]]''.
|
|-
|
''Terminalioxylon mozambicense''<ref name=Sorindeioxylon>{{Cite journal|last1=Bamford |first1=M. |last2=Pickford |first2=M. |title=Stratigraphy, chronology and palaeontology of the Tertiary rocks of the Cheringoma Plateau, Mozambique |year=2021 |journal=Fossil Imprint |volume=77 |issue=1 |pages=187–213 |doi=10.37520/fi.2021.014 |s2cid=245074611 |url=http://fi.nm.cz/en/clanek/stratigraphy-chronology-and-palaeontology-of-the-tertiary-rocks-of-the-cheringoma-plateau-mozambique-2/ |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Bamford & Pickford
|
Probably late Eocene
|
|
{{Flag|Mozambique}}
|
A member of the family [[Biology:Combretaceae|Combretaceae]].
|
|-
|
''Trapa sanyingensis''<ref>{{Cite journal|last1=Aung |first1=A. T. |last2=Del Rio |first2=C. |last3=Wang |first3=T.-X. |last4=Liu |first4=J. |last5=Spicer |first5=T. E. V. |last6=Su |first6=T. |year=2021 |title=Fossil fruits and pollen grains of ''Trapa'' from the Upper Pliocene of the Sanying Formation (Yunnan, China) |journal=Review of Palaeobotany and Palynology |volume=293 |pages=Article 104498 |doi=10.1016/j.revpalbo.2021.104498 |bibcode=2021RPaPa.29304498A }}</ref>
|
Sp. nov
|
In press
|
Aung ''et al.''
|
Late Pliocene
|
Sanying Formation
|
{{Flag|China}}
|
A [[Biology:Water caltrop|water caltrop]].
|
|-
|
''Verrutricolporites pusillus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen, possibly of a member of the family Lythraceae.
|
|-
|
''Xystonia''<ref>{{Cite journal|last1=Carvalho |first1=M. |last2=Herrera |first2=F. |last3=Gómez |first3=S. |last4=Martínez |first4=C. |last5=Jaramillo |first5=C. |year=2021 |title=Early records of Melastomataceae from the middle-late Paleocene rainforests of South America conflict with Laurasian origins |journal=International Journal of Plant Sciences |volume=182 |issue=5 |pages=401–412 |doi=10.1086/714053 |s2cid=233910850 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Carvalho ''et al.''
|
[[Earth:Paleocene|Paleocene]]
|
[[Earth:Bogotá Formation|Bogotá Formation]]
|
{{Flag|Colombia}}
|
A member of the family [[Biology:Melastomataceae|Melastomataceae]]. Genus includes new species ''X. simonae''.
| 
|-
|}

=====Sapindales=====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Aceroxylon aceroides''<ref name=AP611 />
|
Sp. nov
|
Valid
|
Akkemik
|
Early Miocene
|
Hançili Formation
|
{{Flag|Turkey}}
|
A member of the family [[Biology:Sapindaceae|Sapindaceae]].
|
|-
|
''Ampelorhiza''<ref>{{Cite journal|last1=Jud |first1=N. A. |last2=Allen |first2=S. E. |last3=Nelson |first3=C. W. |last4=Bastos |first4=C. L. |last5=Chery |first5=J. G. |title=Climbing since the early Miocene: The fossil record of Paullinieae (Sapindaceae) |year=2021 |journal=PLOS ONE |volume=16 |issue=4 |pages=e0248369 |doi=10.1371/journal.pone.0248369 |pmid=33826635 |pmc=8026063 |bibcode=2021PLoSO..1648369J |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Jud ''et al.''
|
Early Miocene
|
[[Earth:Cucaracha Formation|Cucaracha Formation]]
|
{{Flag|Panama}}
|
A member of the family [[Biology:Sapindaceae|Sapindaceae]] belonging to the subfamily [[Biology:Sapindoideae|Sapindoideae]] and the tribe Paullinieae. Genus includes new species ''A. heteroxylon''.
|
|-
|
''Anacardium gassonii''<ref>{{Cite journal|last1=Rodríguez-Reyes |first1=O. |last2=Estrada-Ruiz |first2=E. |last3=Monje Dussán |first3=C. |last4=de Andrade Brito |first4=L. |last5=Terrazas |first5=T. |title=A new Oligocene-Miocene tree from Panama and historical ''Anacardium'' migration patterns |year=2021 |journal=PLOS ONE |volume=16 |issue=6 |pages=e0250721 |doi=10.1371/journal.pone.0250721 |pmid=34077439 |pmc=8171895 |bibcode=2021PLoSO..1650721R |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Rodríguez-Reyes, Estrada-Ruiz & Terrazas ''in'' Rodríguez-Reyes ''et al.''
|
Oligocene-Miocene
|
|
{{Flag|Panama}}
|
A species of ''[[Biology:Anacardium|Anacardium]]''.
|
|-
|
''Atalantioxylon thanobolensis''<ref>{{Cite journal|last1=Soomro |first1=N. |last2=Mangi |first2=J. |last3=Panhwer |first3=M. |last4=Jatoi |first4=G. |last5=Khuhro |first5=S. |last6=Khokhar |first6=Q. |last7=Khan |first7=S. |last8=Mengal |first8=A. |last9=Shaikh |first9=N. |year=2021 |title=Anatomical characteristics of fossil wood collected from the Manchar Formation (Miocene), Thano Bula Khan, Sindh, Pakistan |journal=Italian Botanist |volume=11 |pages=1–8 |doi=10.3897/italianbotanist.11.60344 |s2cid=233274837 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Soomro ''et al.''
|
Miocene
|
Manchar Formation
|
{{Flag|Pakistan}}
|
Fossil wood of a member of the family [[Biology:Rutaceae|Rutaceae]].
| 
|-
|
''Melia santangensis''<ref>{{Cite journal|last1=Liu |first1=W.-Q. |last2=Xu |first2=S.-L. |last3=Fu |first3=Q.-Y. |last4=Quan |first4=C. L. |last5=Jin |first5=J.-H. |title=Late Oligocene ''Melia'' (Meliaceae) from the Nanning Basin of South China and it's biogeographical implication |year=2021 |journal=Journal of Palaeogeography |volume=10 |issue=1 |pages=Article number 16 |doi=10.1186/s42501-021-00097-x |bibcode=2021JPalG..10...16L |s2cid=235690851 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Liu, Xu & Jin ''in'' Liu ''et al.''
|
Late [[Earth:Oligocene|Oligocene]]
|
Yongning Formation
|
{{Flag|China}}
|
A species of ''[[Biology:Melia (plant)|Melia]]''.
|
|-
|
''Pistacia terrazasae''<ref name="Posthammer2021"/>
|
Sp nov
|
Valid
|
Wheeler & Manchester
|
Late Eocene
|
John Day Formation
|
{{Flag|USA}}<br/>{{Flag|Oregon}}
|
A ''[[Biology:Pistacia|Pistacia]]'' species wood.
|
|-
|
''Proteacidites poriscabratus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the family [[Biology:Sapindaceae|Sapindaceae]].
|
|-
|
''Sorindeioxylon''<ref name=Sorindeioxylon />
|
Gen. et sp. nov
|
Valid
|
Bamford & Pickford
|
Probably late Eocene
|
Mazamba Formation
|
{{Flag|Mozambique}}
|
A member of the family [[Biology:Anacardiaceae|Anacardiaceae]]. Genus includes new species ''S. gorongosense''.
|
|-
|
''Syncolporites tenuicolpatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the genus ''[[Biology:Serjania|Serjania]]''.
|
|-
|
''Tetradium nanningense''<ref>{{Cite journal|last1=Huang |first1=L.-L. |last2=Jin |first2=J.-H. |last3=Quan |first3=C. |last4=Oskolski |first4=A. A. |title=Earliest fossil record of the genus ''Tetradium'' (Rutaceae) in Asia: implications for its evolution and palaeoecology |year=2021 |journal=Papers in Palaeontology |volume=7 |issue=4 |pages=2065–2074 |doi=10.1002/spp2.1394 |s2cid=237688959 }}</ref>
|
Sp. nov
|
Valid
|
Huang ''et al.''
|
Late Oligocene
|
Yongning Formation
|
{{Flag|China}}
|
A species of ''[[Biology:Tetradium|Tetradium]]''.
|
|-
|
''Verrutricolporites simplex''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Pollen of a member of the family [[Biology:Simaroubaceae|Simaroubaceae]].
|
|-
|}
====Non eurosid superrosids====
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Hamamelidoxylon crystalliferum''<ref name="Posthammer2021"/>
|
Sp nov
|
Valid
|
Wheeler & Manchester
|
Late Eocene
|
John Day Formation
|
{{Flag|USA}}<br/>{{Flag|Oregon}}
|
A ''Hamamelidoxylon'' species wood.
|
|-
|
''Liquidambar guipingensis''<ref>{{Cite journal|last1=Huang |first1=L.-L. |last2=Jin |first2=J.-H. |last3=Quan |first3=C. |last4=Oskolski |first4=A. A. |year=2021 |title=New occurrences of Altingiaceae fossil woods from the Miocene and upper Pleistocene of South China with phytogeographic implications |journal=Journal of Palaeogeography |volume=10 |issue=4 |pages=482–493 |doi=10.1016/j.jop.2021.11.001 |bibcode=2021JPalG..10..482H |s2cid=252959116 |url=https://www.sciencedirect.com/science/article/pii/S2095383621000043 |doi-access=free }}</ref>
|
Sp. nov
|
In press
|
Huang ''et al.''
|
Miocene
|
Erzitang Formation
|
{{Flag|China}}
|
A species of ''[[Biology:Liquidambar|Liquidambar]]''.
|
|-
|
''Liquidambaroxylon efeae''<ref name=AP611>{{Cite journal|last=Akkemik |first=Ü. |year=2021 |title=A re-examination of the angiosperm wood record from the early and middle Miocene of Turkey, and new species descriptions |journal=Acta Palaeobotanica |volume=61 |issue=1 |pages=42–94 |doi=10.35535/acpa-2021-0004 |s2cid=237741731 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Akkemik
|
Early Miocene
|
Hançili Formation
|
{{Flag|Turkey}}
|
A member of the family [[Biology:Altingiaceae|Altingiaceae]].
|
|-
|
''Obirafructus''<ref>{{Cite journal|last1=Kajita |first1=Y. |last2=Suzuki |first2=M. H. |last3=Nishida |first3=H. |year=2021 |title=A Multicarpellary Apocarpous Gynoecium from the Late Cretaceous (Coniacian–Santonian) of the Upper Yezo Group of Obira, Hokkaido, Japan: ''Obirafructus kokubunii'' gen. & sp. nov. |journal=Acta Phytotaxonomica et Geobotanica |volume=72 |issue=1 |pages=1–21 |doi=10.18942/apg.202009 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Kajita & Nishida ''in'' Kajita, Suzuki & Nishida
|
Late Cretaceous ([[Earth:Coniacian|Coniacian]]–[[Earth:Santonian|Santonian]])
|
Haborogawa Formation
|
{{Flag|Japan}}
|
A member of Saxifragales of uncertain phylogenetic placement. Genus includes new species ''O. kokubunii''.
| 
|-
|
''Paleoaltingia''<ref>{{Cite journal|last1=Lai |first1=Y. |last2=Gandolfo |first2=M. A. |last3=Crepet |first3=W. L. |last4=Nixon |first4=K. C. |year=2021 |title=''Paleoaltingia'' gen. nov., a new genus of Altingiaceae from the Late Cretaceous of New Jersey |journal=American Journal of Botany |volume=108 |issue=3 |pages=461–471 |doi=10.1002/ajb2.1618 |pmid=33660257 |s2cid=232115011 |doi-access=free }}</ref>
|
Gen. nov
|
Valid
|
Lai ''et al.''
|
[[Earth:Late Cretaceous|Late Cretaceous]] ([[Earth:Turonian|Turonian]])
|
|
{{Flag|United States}}<br>({{Flag|New Jersey}})
|
A member of [[Biology:Altingiaceae|Altingiaceae]]. Genus includes ''P. ovum-dinosauri'' and ''P. polyodonta''.
| 
|-
|}

=== Other angiosperms ===
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Araliaephyllum vittenburgii''<ref>{{Cite journal|last1=Golovneva |first1=L. |last2=Bugdaeva |first2=E. |last3=Volynets |first3=E. |last4=Sun |first4=Y. |last5=Zolina |first5=A. |title=Angiosperm diversification in the Early Cretaceous of Primorye, Far East of Russia |year=2021 |journal=Fossil Imprint |volume=77 |issue=2 |pages=231–255 |doi=10.37520/fi.2021.017 |s2cid=245545364 |url=http://fi.nm.cz/clanek/angiosperm-diversification-in-the-early-cretaceous-of-primorye-far-east-of-russia/ |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Golovneva & Volynets ''in'' Golovneva ''et al.''
|
[[Earth:Early Cretaceous|Early Cretaceous]] ([[Earth:Albian|Albian]])
|
Galenki Formation
|
{{Flag|Russia}}<br>({{Flag|Primorsky Krai}})
|
A flowering plant of uncertain phylogenetic placement.
|
|-
|
''Baderadea''<ref>{{Cite journal|last1=Pessoa |first1=E. M. |last2=Ribeiro |first2=A. C. |last3=Jud |first3=N. A. |year=2021 |title=A eudicot leaf from the Lower Cretaceous (Aptian, Araripe Basin) Crato Konservat-Lagerstätte |journal=American Journal of Botany |volume=108 |issue=10 |pages=2055–2065 |doi=10.1002/ajb2.1751 | issn=0002-9122 |url=https://bsapubs.onlinelibrary.wiley.com/doi/10.1002/ajb2.1751 |pmid=34647319 |s2cid=238858698 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Pessoa, Ribeiro & Jud
|
Early Cretaceous (Aptian)
|
[[Earth:Crato Formation|Crato Formation]]
|
{{Flag|Brazil}}
|
A herbaceous [[Biology:Eudicots|eudicot]] similar to some members of [[Biology:Ranunculales|Ranunculales]]. Genus includes new species ''B. pinnatissecta''.
|
|-
|
''Byttneripollis rugulatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Dilcherifructus''<ref>{{Cite journal|last=Wang |first=X. |title=The Currently Earliest Angiosperm Fruit from the Jurassic of North America |year=2021 |journal=Biosis: Biological Systems |volume=2 |issue=4 |pages=416–422 |doi=10.37819/biosis.001.04.0160 |s2cid=245724946 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Wang
|
Middle Jurassic
|
Simón Formation
|
{{Flag|Mexico}}
|
Possibly a fruit of an early flowering plant. Genus includes new species ''D. mexicana''.
|
|-
|
''Farabeipollis deccanensis''<ref>{{Cite journal|last1=Sonkusare |first1=H. |last2=Samant |first2=B. |last3=Mohabey |first3=D. M. |year=2021 |title=Palynoassemblage from intertrappean sediments of Satpura Group, Betul district, Madhya Pradesh: implications in understanding age and palaeoclimate |journal=Journal of the Palaeontological Society of India |volume=66 |issue=1 |pages=35–54 |url=https://www.researchgate.net/publication/353006021 }}</ref>
|
Sp. nov
|
Valid
|
Sonkusare, Samant & Mohabey
|
Late Cretaceous (Maastrichtian)
|
Deccan Intertrappean Beds
|
{{Flag|India}}
|
Pollen of a flowering plant of uncertain affinity.
|
|-
|
''Florigerminis''<ref>{{cite book |last1=Cui |first1=D.-F. |last2=Hou |first2=Y. |last3=Yin |first3=P. |last4=Wang |first4=X. |year=2021 |chapter=A Jurassic flower bud from the Jurassic of China |editor1=S-C. Chang |editor2=D. Zheng |title=Mesozoic Biological Events and Ecosystems in East Asia |journal=Geological Society, London, Special Publications |publisher=The Geological Society of London |doi=10.1144/SP521-2021-122 |s2cid=244737990 }}</ref>
|
Gen. et sp. nov
|
In press
|
Cui ''et al.''
|
Middle-Late Jurassic
|
Jiulongshan Formation
|
{{Flag|China}}
|
A possible flower bud.<br/> The type species is ''F. jurassica''.<br/> First announced online 2021, Final article published [[Biology:2022 in paleobotany|2022]].
|
[[File:Florigerminis F2.A-B.jpg|thumb|center|upright|''Florigerminis jurassica'']]
|-
|
''Gansufructus''<ref>{{cite journal |last1=Du |first1=B. |last2=Zhang |first2=M. |last3=Sun |first3=B. |last4=Li |first4=A. |last5=Zhang |first5=J. |last6=Yan |first6=D. |last7=Xie |first7=S. |last8=Wu |first8=J. |year=2021 |title=An Exceptionally Well-Preserved Herbaceous Eudicot from the Early Cretaceous (late Aptian-early Albian) of Northwest China |journal=National Science Review |volume=8 |issue=12 |pages=nwab084 |doi=10.1093/nsr/nwab084 |pmid=34987839 |pmc=8692937 }}</ref>
|
Gen. et sp. nov
|
In press
|
Du ''in'' Du ''et al.''
|
Early Cretaceous (late [[Earth:Aptian|Aptian]]-early [[Earth:Albian|Albian]])
|
Zhonggou Formation
|
{{Flag|China}}
|
A [[Biology:Eudicots|eudicot]] of uncertain phylogenetic placement. Genus includes new species ''G. saligna''.
| 
|-
|
''Inaperturopollenites tectatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Ladakhipollenites? corvattatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Ladakhipollenites? endoporatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Ladakhipollenites nanus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Ladakhipollenites? sphaericus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Margocolporites bilinearis''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Margocolporites incertus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Multiporopollenites intermedius''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Nigericolpites''<ref>{{Cite journal|last=Hernández |first=J. A. R. |year=2021 |title=''Nigericolpites'': a replacement name for the illegitimate Maastrichtian magnoliopsid pollen genus ''Clavatricolpites'' Hoeken-Klink. (Angiospermae: Magnoliopsida) |journal=Grana |volume=60 |issue=5 |pages=370–371 |doi=10.1080/00173134.2020.1827025|issn=0017-3134  |s2cid=234286062 }}</ref>
|
Nom. nov
|
Valid
|
Hernández
|
Late Cretaceous (Maastrichtian)
|
|
{{Flag|Nigeria}}
|
Pollen of a flowering plant; a replacement name for ''Clavatricolpites'' Hoeken-Klinkenberg (1964).
| 
|-
|
''Psilaperiporites circinatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Psilaperiporites depressus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Psilastephanocolporites ectoporatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Psilastephanocolporites pseudomarinamensis''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Ranunculacidites pontoreticulatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Retibrevitricolpites pseudoretibolus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Retiperiporites retiporatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Retistephanocolpites liberalis''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Retistephanocolporites loxocolpatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Retitrescolpites benjaminensis''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Retitrescolpites brevicolpatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Retitrescolpites grossus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Retitrescolpites kriptoporus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Retitrescolpites marginatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Retitriporites discretus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Retitriporites sifonis''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Rhoipites apertus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Rhoipites? colpiverrucosus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Rhoipites crassinexinicus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Rhoipites crassitectatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Rhoipites grossomurus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Rhoipites guttatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Rhoipites lolongatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Rhoipites protoguttatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Rhoipites pseudocrassopolaris''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Rhoipites pseudopilatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Rhoipites pseudoscabratus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Rhoipites quantulus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Rhoipites vilis''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Tetracolporopollenites nanus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|
''Tetracolporopollenites xatanawensis''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil pollen.
|
|-
|}

== Other plants ==
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Adelocladoxis''<ref>{{Cite journal|last1=Durieux |first1=T. |last2=Lopez |first2=M. A. |last3=Bronson |first3=A. W. |last4=Tomescu |first4=A. M. F. |year=2021 |title=A new phylogeny of the cladoxylopsid plexus: contribution of an early cladoxylopsid from the Lower Devonian (Emsian) of Quebec |journal=American Journal of Botany |volume=108 |issue=10 |pages=2066–2095 |doi=10.1002/ajb2.1752|issn=0002-9122  |pmid=34664712 |s2cid=239028044 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
Valid
|
Durieux ''et al.''
|
[[Earth:Devonian|Devonian]] ([[Earth:Emsian|Emsian]])
|
Battery Point Formation
|
{{Flag|Canada}}<br>({{Flag|Quebec}})
|
A member of [[Biology:Cladoxylopsida|Cladoxylopsida]]. Genus includes new species ''A. praecox''.
|
|-
|
''[[Biology:Baragwanathia|Baragwanathia brevifolioides]]''<ref>{{Cite journal|last1=Kraft |first1=P. |last2=Kvaček |first2=Z. |title=''Baragwanathia brevifolioides'', a nomen novum for ''B. brevifolia'' P.Kraft et Kvaček, 2017 |year=2021 |journal=Fossil Imprint |volume=77 |issue=1 |pages=53–54 |doi=10.37520/fi.2021.006 |s2cid=245034157 |url=http://fi.nm.cz/en/clanek/baragwanathia-brevifolioides-a-nomen-novum-for-b-brevifolia-p-kraft-et-kvacek-2017-2/ |doi-access=free }}</ref>
|
Nom. nov
|
Valid
|
Kraft & Kvaček
|
[[Earth:Silurian|Silurian]] ([[Earth:Pridoli Epoch|Přídolí]])
|
Požáry Formation
|
{{Flag|Czech Republic}}
|
A member of the family [[Biology:Drepanophycaceae|Drepanophycaceae]]; a replacement name for ''Baragwanathia brevifolia'' Kraft & Kvaček (2017).
|
|-
|
''Camarozonosporites fossulatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Spores of a member of the family [[Biology:Lycopodiaceae|Lycopodiaceae]].
|
|-
|
''Cingulatisporites cristatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Spores of a member of the genus ''[[Biology:Selaginella|Selaginella]]''.
|
|-
|
''Cingulatisporites matisiensis''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Spores of a member of the genus ''[[Biology:Phaeoceros|Phaeoceros]]''.
|
|-
|
''Closterium mosbruggeri''<ref>{{Cite journal|last1=Ivanov |first1=D. |last2=Belkinova |first2=D. |year=2021 |title=''Closterium mosbruggeri'' sp. nov.: a new fossil species from the middle Miocene of Northwest Bulgaria |journal=Palaeobiodiversity and Palaeoenvironments |volume=101 |issue=1 |pages=69–74 |doi=10.1007/s12549-020-00476-7 |s2cid=231859320 }}</ref>
|
Sp. nov
|
Valid
|
Ivanov & Belkinova
|
Miocene ([[Earth:Serravallian|Serravallian]])
|
|
{{Flag|Bulgaria}}
|
A green alga, a species of ''[[Biology:Closterium|Closterium]]''.
|
|-
|
''Colpodexylon mergae''<ref name=RPPColpodexylon>{{Cite journal|last1=Harris |first1=C. |last2=Gess |first2=R. W. |last3=Prestianni |first3=C. |last4=Bamford |first4=M. K. |year=2021 |title=A Late Devonian refugium for ''Colpodexylon'' (Lycopsida) at high latitude |journal=Review of Palaeobotany and Palynology |volume=293 |pages=Article 104481 |doi=10.1016/j.revpalbo.2021.104481 |bibcode=2021RPaPa.29304481H |url=https://orbi.uliege.be/bitstream/2268/295475/1/Harris%20et%20al.%202021%20Colpodexylon.pdf }}</ref>
|
Sp. nov
|
In press
|
Harris ''et al.''
|
[[Earth:Devonian|Devonian]] ([[Earth:Famennian|Famennian]])
|
Witpoort Formation
|
{{Flag|South Africa}}
|
A member of Lycopsida.
|
|-
|
''Colpodexylon pullumpedes''<ref name=RPPColpodexylon />
|
Sp. nov
|
In press
|
Harris ''et al.''
|
Devonian (Famennian)
|
Witpoort Formation
|
{{Flag|South Africa}}
|
A member of Lycopsida.
|
|-
|
''Cordaabaxicutis martii''<ref name=SimunekLojka>{{Cite journal|last1=Šimůnek |first1=Z. |last2=Lojka |first2=R. |year=2021 |title=Carboniferous cuticles from the Lubná coal seam (Kladno Formation, Kladno-Rakovník Basin, Czech Republic) |journal=Palaeontographica Abteilung B |volume=303 |issue=4–6 |pages=77–117 |doi=10.1127/palb/2021/0076 |bibcode=2021PalAB.303...77S |s2cid=244561451 }}</ref>
|
Sp. nov
|
Valid
|
Šimůnek & Lojka
|
Carboniferous (Pennsylvanian)
|
Kladno Formation
|
{{Flag|Czech Republic}}
|
[[Biology:Cordaitales|Cordaitalean]] cuticles.
|
|-
|
''Cordaabaxicutis papillosus''<ref name=SimunekLojka />
|
Sp. nov
|
Valid
|
Šimůnek & Lojka
|
Carboniferous (Pennsylvanian)
|
Kladno Formation
|
{{Flag|Czech Republic}}
|
Cordaitalean cuticles.
|
|-
|
''Cordaadaxicutis raristomatus''<ref name=SimunekLojka />
|
Sp. nov
|
Valid
|
Šimůnek & Lojka
|
Carboniferous (Pennsylvanian)
|
Kladno Formation
|
{{Flag|Czech Republic}}
|
Cordaitalean cuticles.
|
|-
|
''Cynodontium luthii''<ref>{{Cite journal|last1=Bippus |first1=A. C. |last2=Rothwell |first2=G. W. |last3=Stockey |first3=R. A. |year=2021 |title=''Cynodontium luthii'' sp. nov.: a permineralized moss gametophyte from the Late Cretaceous of the North Slope of Alaska |journal=American Journal of Botany |volume=108 |issue=3 |pages=495–504 |doi=10.1002/ajb2.1617 |pmid=33650114 |s2cid=232091615 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Bippus, Rothwell & Stockey
|
Late Cretaceous
|
|
{{Flag|United States}}<br>({{Flag|Alaska}})
|
A moss belonging to the family [[Biology:Rhabdoweisiaceae|Rhabdoweisiaceae]], a species of ''[[Biology:Cynodontium|Cynodontium]]''.
|
|-
|
''Dayvaultia''<ref>{{Cite journal|last1=Manchester |first1=S. R. |last2=Zhang |first2=X. |last3=Hotton |first3=C. L. |last4=Wing |first4=S. L. |last5=Crane |first5=P. R. |year=2021 |title=Distinctive quadrangular seed-bearing structures of gnetalean affinity from the Late Jurassic Morrison Formation of Utah, USA |journal=Journal of Systematic Palaeontology |volume=19 |issue=10 |pages=743–760 |doi=10.1080/14772019.2021.1968522 |s2cid=239021014 |url=https://www.tandfonline.com/doi/abs/10.1080/14772019.2021.1968522 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Manchester ''et al.''
|
Late Jurassic
|
[[Earth:Morrison Formation|Morrison Formation]]
|
{{Flag|United States}}<br>({{Flag|Utah}})
|
A seed-bearing structure of [[Biology:Gnetales|gnetalean]] affinity. Genus includes new species ''D. tetragona''.
|
|-
|
''Distefanopolia''<ref>{{Cite journal|last1=Barattolo |first1=F. |last2=Romano |first2=R. |last3=Conrad |first3=M. |title=Evidence of external gametophores in puzzling Late Triassic–Early Jurassic dasycladalean green algae |year=2021 |journal=Acta Palaeontologica Polonica |volume=66 |issue=4 |pages=901–919 |doi=10.4202/app.00883.2021 |doi-access=free }}</ref>
|
Gen. et comb. nov
|
Valid
|
Barattolo, Romano & Conrad
|
[[Earth:Late Triassic|Late Triassic]] and possibly Early Jurassic
|
|
{{Flag|Austria}}<br>
{{Flag|Czech Republic}}<br>
{{Flag|Germany}}<br>
{{Flag|Greece}}<br>
{{Flag|Italy}}<br>
{{Flag|Oman}}<br>
{{Flag|Slovakia}}
|
A green alga belonging to the group [[Biology:Dasycladales|Dasycladales]] and the family Bornetelleae. Genus includes ''"Heteroporella" micropora'' Di Stefano & Senowbari-Daryan (1985), ''"Heteroporella" macropora'' Di Stefano, 1981 ex Di Stefano & Senowbari-Daryan (1985), ''"Chinianella" zanklii'' Ott (1967), ''"Chinianella" crosii'' Ott (1968) and ''"Heteroporella" carpatica'' Bystrický (1967).
|
|-
|
''[[Biology:Distichophytum|Distichophytum mogilatii]]''<ref>{{Cite journal|last=Naugolnykh |first=S. V. |year=2021 |title=A new species of ''Distichophytum'' Mägdefrau, 1938 from the Lower Devonian of Siberia (the Torgashino locality, Krasnojarsk krai, Russia) |journal=Wulfenia |volume=28 |pages=141–150 |url=https://www.researchgate.net/publication/356613631 }}</ref>
|
Sp. nov
|
Valid
|
Naugolnykh
|
[[Earth:Devonian|Devonian]]
|
|
{{Flag|Russia}}<br>({{Flag|Krasnoyarsk Krai}})
|
A member of the family [[Biology:Zosterophyll|Zosterophyll]]aceae.
|
|-
|
''Dragastanella''<ref>{{Cite journal|last1=Barattolo |first1=F. |last2=Bucur |first2=I. I. |last3=Marian |first3=A. V. |year=2021 |title=Deciphering voids in Dasycladales, the case of ''Dragastanella transylvanica'', a new Lower Cretaceous triploporellacean genus and species from Romania |journal=Journal of Paleontology |volume=95 |issue=5 |pages=889–905 |doi=10.1017/jpa.2021.40|bibcode=2021JPal...95..889B | issn=0022-3360 |s2cid=236346831 |doi-access=free }}</ref>
|
Gen. et sp. et comb. nov
|
Valid
|
Barattolo, Bucur & Marian
|
[[Earth:Early Cretaceous|Early Cretaceous]]
|
|
{{Flag|Italy}}<br>
{{Flag|Romania}}<br>
{{Flag|Spain}}
|
A green alga belonging to the group [[Biology:Dasycladales|Dasycladales]]. Genus includes new species ''D. transylvanica'', as well as ''"Zittelina" hispanica'' Masse, Arias & Vilas (1993), ''"Zittelina" massei'' Bucur, Granier & Săsăran (2010) and ''"Triploporella" matesina'' Barattolo (1980).
|
|-
|
''Elandia''<ref name=Elandia>{{Cite journal|last1=Gess |first1=R. W. |last2=Prestianni |first2=C. |year=2021 |title=An early Devonian flora from the Baviaanskloof Formation (Table Mountain Group) of South Africa |journal=Scientific Reports |volume=11 |issue=1 |pages=Article number 11859 |doi=10.1038/s41598-021-90180-z |pmid=34088916 |pmc=8178408 |bibcode=2021NatSR..1111859G }}</ref>
|
Gen. et sp. nov
|
Valid
|
Gess & Prestianni
|
[[Earth:Devonian|Devonian]] ([[Earth:Lochkovian|Lochkovian]]?)
|
Baviaanskloof Formation
|
{{Flag|South Africa}}
|
An early [[Biology:Polysporangiophyte|polysporangiophyte]]. Genus includes new species ''E. itshoba''.
|
|-
|
''Flabellopteris''<ref>{{Cite journal|last1=Gess |first1=R. W. |last2=Prestianni |first2=C. |year=2021 |title=''Flabellopteris lococannensis'' gen. et sp. nov.: A new fern-like plant from the Famennian of South Africa |journal=Review of Palaeobotany and Palynology |volume=297 |pages=Article 104585 |doi=10.1016/j.revpalbo.2021.104585 |s2cid=245209478 |url=https://orbi.uliege.be/bitstream/2268/302493/1/1-s2.0-S0034666721002098-main.pdf }}</ref>
|
Gen. et sp. nov
|
In press
|
Gess & Prestianni
|
Devonian ([[Earth:Famennian|Famennian]])
|
Witpoort Formation
|
{{Flag|South Africa}}
|
A fern-like plant of uncertain affinities. Genus includes new species ''F. lococannensis''.
|
|-
|
''Foraminisporis connexus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Spores of a member of the genus ''[[Biology:Anthoceros|Anthoceros]]''.
|
|-
|
''Frullania kachinensis''<ref>{{Cite journal|last1=Li |first1=Y. |last2=Wang |first2=Y.-D. |last3=Feldberg |first3=K. |last4=Wang |first4=Q. |last5=Yang |first5=X.-J. |year=2021 |title=A new leafy liverwort of ''Frullania'' (Frullaniaceae, Porellales) from the mid-Cretaceous Kachin amber, Myanmar |journal=Geological Journal |volume=56 |issue=10 |pages=5046–5057 |doi=10.1002/gj.4222 |s2cid=238795680 }}</ref>
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A liverwort, a species of ''[[Biology:Frullania|Frullania]]''.
|
|-
|
''Frullania palaeoafricana''<ref name=Boujoetalamber>{{Cite journal|last1=Bouju |first1=V. |last2=Feldberg |first2=K. |last3=Kaasalainen |first3=U. |last4=Schäfer-Verwimp |first4=A. |last5=Hedenäs |first5=L. |last6=Buck |first6=W. R. |last7=Wang |first7=B. |last8=Perrichot |first8=V. |last9=Schmidt |first9=A. R. |title=Miocene Ethiopian amber: a new source of fossil cryptogams |journal=Journal of Systematics and Evolution |year=2022 |volume=60 |issue=4 |pages=932–954 |doi=10.1111/jse.12796 |s2cid=236586190 |url=https://hal-insu.archives-ouvertes.fr/insu-03231697/file/BOUJU-2021.pdf }}</ref>
|
Sp. nov
|
In press
|
Bouju ''et al.''
|
Miocene
|
Ethiopian amber
|
{{Flag|Ethiopia}}
|
A liverwort, a species of ''Frullania''.
|
|-
|
''Frullania shewanensis''<ref name=Boujoetalamber />
|
Sp. nov
|
In press
|
Bouju ''et al.''
|
Miocene
|
Ethiopian amber
|
{{Flag|Ethiopia}}
|
A liverwort, a species of ''Frullania''.
|
|-
|
''Gilboaphyton fuyunensis''<ref>{{Cite journal|last1=Liu |first1=B.-C. |last2=Bai |first2=J. |last3=Wang |first3=Y. |last4=Yang |first4=N. |last5=Xu |first5=H.-H. |year=2021 |title=On the discovery of ''Gilboaphyton'' (Lycopsida) from the Upper Devonian of East Junggar, Xinjiang, and its global distribution |journal=Review of Palaeobotany and Palynology |volume=292 |pages=Article 104473 |doi=10.1016/j.revpalbo.2021.104473 |bibcode=2021RPaPa.29204473L }}</ref>
|
Sp. nov
|
In press
|
Liu ''et al.''
|
Late [[Earth:Devonian|Devonian]]
|
Kaxiweng Formation
|
{{Flag|China}}
|
A member of [[Biology:Protolepidodendrales|Protolepidodendrales]].
|
|-
|
''Guazia''<ref>{{Cite journal|last1=Wang |first1=D.-M. |last2=Liu |first2=L. |last3=Zhou |first3=Y. |last4=Qin |first4=M. |last5=Meng |first5=M.-C. |last6=Guo |first6=Y. |last7=Xue |first7=J.-Z. |year=2021 |title=''Guazia'', the earliest ovule without cupule but with unique integumentary lobes |journal=National Science Review |volume=9 |issue=4 |pages=nwab196 |doi=10.1093/nsr/nwab196 |pmid=35386924 |pmc=8982201 }}</ref>
|
Gen. et sp. nov
|
In press
|
Wang ''et al.''
|
Late Devonian
|
Wutong Formation
|
{{Flag|China}}
|
A seed plant of uncertain phylogenetic placement. Genus includes new species ''G. dongzhiensis''.
|
|-
|
''Ixostrobus daohugouensis''<ref>{{Cite journal|last1=Na |first1=Y. |last2=Sun |first2=C. |last3=Wang |first3=H. |last4=Huang |first4=T. |last5=Bevitt |first5=J. |last6=Li |first6=Y. |last7=Li |first7=T. |last8=Zhao |first8=Y. |last9=Li |first9=N. |year=2021 |title=Application of neutron tomography in studying new material of ''Ixostrobus'' Raciborski from the Middle Jurassic of Inner Mongolia, China |journal=Geological Journal |volume=56 |issue=9 |pages=4618–4626 |doi=10.1002/gj.4196 |s2cid=237869542 }}</ref>
|
Sp. nov
|
In press
|
Na & Sun ''in'' Na ''et al.''
|
Middle Jurassic
|
|
{{Flag|China}}
|
A member of [[Biology:Czekanowskiales|Czekanowskiales]].
|
|-
|
''Kenrickia''<ref>{{Cite journal|last1=Toledo |first1=S. |last2=Bippus |first2=A. C. |last3=Atkinson |first3=B. A. |last4=Bronson |first4=A. W. |last5=Tomescu |first5=A. M. F. |year=2021 |title=Taxon sampling and alternative hypotheses of relationships in the euphyllophyte plexus that gave rise to seed plants: insights from an Early Devonian radiatopsid |journal=New Phytologist |volume=232 |issue=2 |pages=914–927 |doi=10.1111/nph.17511 |pmid=34031894 |s2cid=235199240 |doi-access=free }}</ref>
|
Gen. et sp. nov
|
In press
|
Toledo ''et al.''
|
[[Earth:Devonian|Devonian]] ([[Earth:Emsian|Emsian]])
|
Battery Point Formation
|
{{Flag|Canada}}
|
An early [[Biology:Euphyllophyte|euphyllophyte]] belonging to the group Radiatopses. Genus includes new species ''K. bivena''.
|
|-
|
''Krommia''<ref name=Elandia />
|
Gen. et sp. nov
|
Valid
|
Gess & Prestianni
|
Devonian (Lochkovian?)
|
Baviaanskloof Formation
|
{{Flag|Brazil}}<br>
{{Flag|South Africa}}
|
An early polysporangiophyte. Genus includes new species ''K. parvapila''.
|
|-
|
''Lejeunea abyssinicoides''<ref name=Boujoetalamber />
|
Sp. nov
|
In press
|
Bouju ''et al.''
|
Miocene
|
Ethiopian amber
|
{{Flag|Ethiopia}}
|
A liverwort, a species of ''[[Biology:Lejeunea|Lejeunea]]''.
|
|-
|
''Lycaugea''<ref>{{Cite journal|last1=Meyer-Berthaud |first1=B. |last2=Decombeix |first2=A.-L. |last3=Blanchard |first3=R. |year=2021 |title=''Lycaugea edieae'' gen. et sp. nov., a Late Devonian lycopsid from New South Wales, Australia |journal=International Journal of Plant Sciences |volume=182 |issue=6 |pages=418–429 |doi=10.1086/714350 |s2cid=233666757 |url=https://hal.inrae.fr/hal-03231982/file/2021%20Meyer%20Berthaud%20et%20al%20Lycaugea%20.pdf }}</ref>
|
Gen. et sp. nov
|
Valid
|
Meyer-Berthaud, Decombeix & Blanchard
|
[[Earth:Devonian|Devonian]] ([[Earth:Famennian|Famennian]])
|
|
{{Flag|Australia}}
|
A [[Biology:Lycopsid|lycopsid]]. Genus includes new species ''L. edieae''.
|
|-
|
''Lycopodiumsporites amazonicus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Spores of a member of the family Lycopodiaceae.
|
|-
|
''Melvillipteris sonidia''<ref>{{Cite journal|last1=Bai |first1=L. |last2=Huang |first2=P. |last3=Yang |first3=N. |last4=Ju |first4=W. |last5=Liu |first5=J. |last6=Basinger |first6=J. F. |last7=Xu |first7=H. |last8=Xue |first8=J. |year=2021 |title=A new Late Devonian flora from Sonid Zuoqi, Inner Mongolia, northeastern China |journal=Journal of Paleontology |volume=96 |issue=2 |pages=462–484 |doi=10.1017/jpa.2021.93 |s2cid=239867911 |url=https://zenodo.org/record/5301469 }}</ref>
|
Sp. nov
|
In press
|
Bai ''et al.''
|
[[Earth:Devonian|Devonian]] (probably [[Earth:Famennian|Famennian]])
|
|
{{Flag|China}}
|
A member of [[Biology:Rhacophytales|Rhacophytales]].
|
|-
|
''Mesochara adobensis''<ref>{{Cite journal|last1=De Sosa Tomas |first1=A. |last2=Martín-Closas |first2=C. |last3=Vallati |first3=P. |last4=Krause |first4=M. |title=Early Cretaceous ''Mesochara''-rich assemblages from central Patagonia, Argentina, predate the origin of homogenous Charoidean floras by about 30 million years |year=2021 |journal=Cretaceous Research |volume=129 |pages=Article 105017 |doi=10.1016/j.cretres.2021.105017|issn=0195-6671 |s2cid=239707025 }}</ref>
|
Sp. nov
|
In press
|
De Sosa Tomas ''et al.''
|
[[Earth:Early Cretaceous|Early Cretaceous]]
|
[[Earth:Los Adobes Formation|Los Adobes Formation]]
|
{{Flag|Argentina}}
|
A member of [[Biology:Charophyta|Charophyta]].
|
|-
|
''Mesochara dobrogeica''<ref>{{Cite journal|last1=Sanjuan |first1=J. |last2=Vicente |first2=A. |last3=Pérez-Cano |first3=J. |last4=Stoica |first4=M. |last5=Martín-Closas |first5=C. |year=2021 |title=Early Cretaceous charophytes from south Dobrogea (Romania). Biostratigraphy and palaeobiogeography |journal=Cretaceous Research |volume=122 |pages=Article 104762 |doi=10.1016/j.cretres.2021.104762 |bibcode=2021CrRes.12204762S |hdl=2445/181959 |s2cid=234160551 |doi-access=free |hdl-access=free }}</ref>
|
Sp. nov
|
In press
|
Sanjuan ''et al.''
|
[[Earth:Early Cretaceous|Early Cretaceous]] ([[Earth:Berriasian|Berriasian]])
|
|
{{Flag|Romania}}
|
A member of Charophyta.
| 
|-
|
''Mixoxylon''<ref>{{Cite journal|last1=Chernomorets |first1=O. |last2=Sakala |first2=J. |year=2021 |title=''Mixoxylon australe'' gen. et sp. nov., a unique homoxylous wood with non-angiosperm affinity from the Lower Cretaceous of Antarctica (Albian, James Ross Island) |journal=Antarctic Science |volume=33 |issue=5 |pages=493–501 |doi=10.1017/S0954102021000389 |bibcode=2021AntSc..33..493C |s2cid=243834316 }}</ref>
|
Gen. et sp. nov
|
In press
|
Chernomorets & Sakala
|
[[Earth:Early Cretaceous|Early Cretaceous]] ([[Earth:Albian|Albian]])
|
Whisky Bay Formation
|
[[Earth:Antarctica|Antarctica]]
|
A homoxylous wood of uncertain systematic affinities. Genus includes new species ''M. australe''.
|
|-
|
''Mtshaelo''<ref name=Elandia />
|
Gen. et sp. nov
|
Valid
|
Gess & Prestianni
|
Devonian (Lochkovian?)
|
Baviaanskloof Formation
|
{{Flag|South Africa}}
|
An early polysporangiophyte. Genus includes new species ''M. kougaensis''.
|
|-
|
''Omniastrobus''<ref>{{Cite journal|last1=Bonacorsi |first1=N. K. |last2=Gensel |first2=P. G. |last3=Hueber |first3=F. M. |last4=Leslie |first4=A. B. |year=2021 |title=''Omniastrobus'' gen. nov., an Emsian plant with implications for the evolution of heterospory in the Early Devonian |journal=International Journal of Plant Sciences |volume=182 |issue=3 |pages=198–209 |doi=10.1086/712356 |s2cid=232050463 }}</ref>
|
Gen. et sp. nov
|
Valid
|
Bonacorsi ''et al.''
|
[[Earth:Devonian|Devonian]] ([[Earth:Emsian|Emsian]])
|
Campbellton Formation
|
{{Flag|Canada}}
|
A [[Biology:Lycophyte|lycophyte]]. Genus includes new species ''O. dawsonii''.
|
|-
|
''Palaeonitella trifurcata''<ref>{{Cite journal|last1=Martín-Closas |first1=C. |last2=Segura-Altés |first2=R. |last3=Pérez-Cano |first3=J. |last4=Bover-Arnal |first4=T. |last5=Sanjuan |first5=J. |year=2021 |title=''Palaeonitella trifurcata'' n. sp., a cortoid-building charophyte from the Lower Cretaceous of Catalonia |journal=Review of Palaeobotany and Palynology |volume=295 |pages=Article 104523 |doi=10.1016/j.revpalbo.2021.104523 |bibcode=2021RPaPa.29504523M |doi-access=free |hdl=2445/181500 |hdl-access=free }}</ref>
|
Sp. nov
|
Valid
|
Martín-Closas ''et al.''
|
[[Earth:Early Cretaceous|Early Cretaceous]] ([[Earth:Barremian|Barremian]]–[[Earth:Aptian|Aptian]])
|
|
{{Flag|Spain}}
|
A member of Charophyta belonging to the family [[Biology:Characeae|Characeae]].
|
|-
|
''Paratingia wuhaia''<ref>{{Cite journal|last1=Wang |first1=J. |last2=Hilton |first2=J. |last3=Pfefferkorn |first3=H. W. |last4=Wang |first4=S. |last5=Zhang |first5=Y. |last6=Bek |first6=J. |last7=Pšenička |first7=J. |last8=Seyfullah |first8=L. J. |last9=Dilcher |first9=D. |year=2021 |title=Ancient noeggerathialean reveals the seed plant sister group diversified alongside the primary seed plant radiation |journal=Proceedings of the National Academy of Sciences of the United States of America |volume=118 |issue=11 |pages=e2013442118 |doi=10.1073/pnas.2013442118 | issn=0027-8424 |pmid=33836571 |pmc=7980368 |bibcode=2021PNAS..11813442W |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
Wang ''et al.''
|
Permian ([[Earth:Asselian|Asselian]])
|
Taiyuan Formation
|
{{Flag|China}}
|
A [[Biology:Progymnosperm|progymnosperm]] belonging to the group [[Biology:Noeggerathiales|Noeggerathiales]] and the family Tingiostachyaceae.
|
|-
|
''Permotheca? musaformis''<ref>{{Cite journal |last1=Foraponova |first1=T. S. |last2=Karasev |first2=E. V. |title=Systematic problems of pollen organs of the genus ''Permotheca'' Zalessky from the Permian of Subangara |year=2021 |journal=Paleontological Journal |volume=55 |issue=6 |pages=691–706 |doi=10.1134/S0031030121060058 |s2cid=245010226 |url=https://www.elibrary.ru/item.asp?id=46621802 }}</ref>
|
Sp. nov
|
Valid
|
Foraponova & Karasev
|
Permian
|
|
{{Flag|Russia}}
|
A [[Biology:Pteridospermatophyta|pteridosperm]].
|
|-
|
''Radula heinrichsii''<ref>{{Cite journal|last1=Feldberg |first1=K. |last2=Schäfer-Verwimp |first2=A. |last3=Renner |first3=M. A. M. |last4=von Konrat |first4=M. |last5=Bechteler |first5=J. |last6=Müller |first6=P. |last7=Wang |first7=Y.-D. |last8=Schneider |first8=H. |last9=Schmidt |first9=A. R. |title=Liverworts from Cretaceous amber |year=2021 |journal=Cretaceous Research |volume=128 |pages=Article 104987 |doi=10.1016/j.cretres.2021.104987 |bibcode=2021CrRes.12804987F |s2cid=238782100 }}</ref>
|
Sp. nov
|
In press
|
Feldberg ''et al.''
|
Cretaceous
|
Burmese amber
|
{{Flag|Myanmar}}
|
A liverwort, a species of ''[[Biology:Radula (plant)|Radula]]''.
|
|-
|
''Rehamnia''<ref>{{Cite journal|last1=Oukassou |first1=M. |last2=Naugolnykh |first2=S. V. |year=2021 |title=''Rehamnia'' (Lycopodiophyta), a new enigmatic Late Devonian plant from Morocco |journal=Journal of African Earth Sciences |volume=182 |pages=Article 104274 |doi=10.1016/j.jafrearsci.2021.104274 |bibcode=2021JAfES.18204274O }}</ref>
|
Gen. et sp. nov
|
In press
|
Oukassou & Naugolnykh
|
Late [[Earth:Devonian|Devonian]]
|
|
{{Flag|Morocco}}
|
A member of [[Biology:Lycopodiophyta|Lycopodiophyta]] of uncertain phylogenetic placement. Genus includes new species ''R. michardis''.
|
|-
|
''Ricciopsis baojishanensis''<ref>{{Cite journal|last1=Han |first1=L. |last2=Yang |first2=T. |last3=Wang |first3=H.-J. |last4=Cai |first4=J.-H. |last5=Liang |first5=W.-Y. |last6=Bao |first6=L. |last7=Chen |first7=H.-Y. |last8=Zhang |first8=L. |last9=Li |first9=W.-J. |last10=Yan |first10=D.-F. |year=2021 |title=Liverwort fossils from the Late Triassic of Baiyin, Gansu, and their geological significance |journal=Palaeoworld |volume=in press |doi=10.1016/j.palwor.2021.09.009 |s2cid=242525785 }}</ref>
|
Sp. nov
|
In press
|
Han & Yan ''in'' Han ''et al.''
|
[[Earth:Late Triassic|Late Triassic]]
|
Nanying'er Formation
|
{{Flag|China}}
|
A liverwort.
|
|-
|
''[[Biology:Salopella|Salopella laidae]]''<ref>{{cite journal |last1=McSweeney |first1=F. R. |last2=Shimeta |first2=J. |last3=Buckeridge |first3=J. S. |year=2021 |title=Early land plants from the Lower Devonian of central Victoria, Australia, including a new species of ''Salopella'' |journal=Memoirs of Museum Victoria |volume=80 |pages=193–205 |doi=10.24199/j.mmv.2021.80.11 |s2cid=242026265 |doi-access=free }}</ref>
|
Sp. nov
|
Valid
|
McSweeney, Shimeta & Buckeridge
|
[[Earth:Devonian|Devonian]] ([[Earth:Pragian|Pragian]])
|
Yea Formation
|
{{Flag|Australia}}
|
An early land plant of uncertain affinities.
|
|-
|
''Skyttegaardia''<ref>{{Cite journal|last1=Friis |first1=E. M. |last2=Crane |first2=P. R. |last3=Pedersen |first3=K. R. |year=2021 |title=Microsporangiophores from the Early Cretaceous (Berriasian) of Bornholm, Denmark, with comments on a pre-angiosperm xerophytic flora |journal=Review of Palaeobotany and Palynology |volume=293 |pages=Article 104487 |doi=10.1016/j.revpalbo.2021.104487 |bibcode=2021RPaPa.29304487F }}</ref>
|
Gen. et sp. nov
|
In press
|
Friis, Crane & Pedersen
|
[[Earth:Early Cretaceous|Early Cretaceous]] ([[Earth:Berriasian|Berriasian]])
|
|
{{Flag|Denmark}}
|
A plant of uncertain phylogenetic placement, possibly close to cycads. Genus includes new species ''S. galtieri''.
|
|-
|
''Thysananthus aethiopicus''<ref name=Boujoetalamber />
|
Sp. nov
|
In press
|
Bouju ''et al.''
|
Miocene
|
Ethiopian amber
|
{{Flag|Ethiopia}}
|
A liverwort belonging to the family [[Biology:Lejeuneaceae|Lejeuneaceae]].
|
|-
|
''Velascoa''<ref>{{Cite journal|last1=Flores Barragan |first1=M. A. |last2=Velasco de León |first2=M. P. |last3=Ortega Chavez |first3=E. |year=2021 |title=New genus for megaphyllous leaves from the Upper Paleozoic of Mexico ''Velascoa pueblensis'' gen. nov |journal=Journal of South American Earth Sciences |volume=110 |pages=Article 103408 |doi=10.1016/j.jsames.2021.103408 |bibcode=2021JSAES.11003408F }}</ref>
|
Gen. et sp. nov
|
Junior homonym
|
Flores Barragan, Velasco de León & Ortega Chavez
|
[[Earth:Permian|Permian]]
|
Matzitzi Formation
|
{{Flag|Mexico}}
|
Fossil leaves of a plant of uncertain phylogenetic placement, with a morphology similar to [[Biology:Ginkgophyta|Ginkgophyta]]. Genus includes new species ''V. pueblensis''. The generic name is preoccupied by ''[[Biology:Velascoa|Velascoa]]'' Calderón & Rzedowski (1997).
|
|-
|
''Vitinellopsis''<ref>{{Cite journal|last1=Vachard |first1=D. |last2=Bucur |first2=I. I. |last3=Munnecke |first3=A. |year=2021 |title=''Vitinellopsis'' nov. gen., a new calcareous alga (Chlorophyta, Bryopsidales) from the Silurian of Gotland (Sweden), and the tribe Vitinelleae nov. nom |journal=Geobios |volume=70 |pages=75–85 |doi=10.1016/j.geobios.2021.10.001 |s2cid=244766381 }}</ref>
|
Gen. et sp. nov
|
In press
|
Vachard, Bucur & Munnecke
|
[[Earth:Silurian|Silurian]]
|
|
{{Flag|Sweden}}
|
A green alga belonging to the group [[Biology:Bryopsidales|Bryopsidales]]. Genus includes new species ''V. gotlandica''.
|
|-
|
''[[Biology:Zosterophyllum|Zosterophyllum confertum]]''<ref>{{Cite journal|last1=Gossmann |first1=R. |last2=Poschmann |first2=M. J. |last3=Giesen |first3=P. |last4=Schultka |first4=S. |title=A stratigraphically significant new zosterophyllopsid from the Rhenish Lower Devonian (W Germany) |journal=Palaeobiodiversity and Palaeoenvironments |year=2022 |volume=102 |issue=3 |pages=503–519 |doi=10.1007/s12549-021-00509-9 |s2cid=238479823 |url=https://link.springer.com/article/10.1007/s12549-021-00509-9 }}</ref>
|
Sp. nov
|
In press
|
Gossmann ''et al.''
|
Early [[Earth:Devonian|Devonian]]
|
|
{{Flag|Germany}}
|
A [[Biology:Zosterophyll|zosterophyll]].
|
|-
|}

== Palynology ==
{| class="wikitable sortable" align="center" width="100%"
|-
!Name
!Novelty
!Status
!Authors
!Age
!Type locality
!Location
!Notes
!Images
|-
|
''Camarozonosporites trilobatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
A spore of [[Biology:Lycopodiales|lycopodialean]] affinity.
|
|-
|
''Echinatisporis infantulus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil spores.
|
|-
|
''Erlansonisporites duwaensis''<ref name=Luntaispora>{{Cite book|last1=Li |first1=W.B. |last2=Batten |first2=D. J. |last3=Li |first3=J.G. |last4=Peng |first4=J.G. |year=2021 |title=Mesozoic Megaspores and Palynomorphs from Tarim Basin, Northwest China  |journal=Palaeontologia Sinica |volume=202 |pages=1–250 |isbn=978-7030696526 }}</ref>
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Erlansonisporites exquisitus''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Erlansonisporites perbellus''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Erlansonisporites textilis''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Ginkgomonocolpites''<ref>{{Cite journal|last=Hernández |first=J. A. R. |year=2021 |title=A replacement name for the poorly known illegitimate Paleogene genus ''Psilamonocolpites'' Y.K.Mathur (Ginkgoales) |journal=Phytotaxa |volume=500 |issue=1 |pages=57–58 |doi=10.11646/phytotaxa.500.1.10 |s2cid=236595043 }}</ref>
|
Nom. nov
|
Valid
|
Hernández
|
Paleogene
|
|
{{Flag|India}}
|
A gymnosperm pollen; a replacement name for ''Psilamonocolpites'' Mathur (1966).
|
|-
|
''Hamulatisporis bareanus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil spores.
|
|-
|
''Henrisporites longibaculiformis''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Horstisporites comitus''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Horstisporites denticulatus''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Horstisporites subtilis''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Horstisporites tarimensis''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Hughesisporites reticulatus''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Hughesisporites unicus''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Ischyosporites dubius''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
A spore of [[Biology:Dicksoniaceae|dicksoniaceous]] affinity.
|
|-
|
''Ischyosporites granulatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil spores.
|
|-
|
''Laevigatosporites indigestus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil spores.
|
|-
|
''Luntaispora''<ref name=Luntaispora />
|
Gen. et sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
Genus includes new species ''L. laevigata''.
|
|-
|
''Microfoveolatosporis simplex''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil spores.
|
|-
|
''Minerisporites tarimensis''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Minerisporites triangularis''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Narkisporites conicus''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Narkisporites densibaculatus''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Narkisporites densiconicus''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Narkisporites tarimensis''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Neoraistrickia dubia''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil spores.
|
|-
|
''Noniasporites triassicus''<ref>{{cite book |last1=Ghosh |first1=A. K. |last2=Chatterjee |first2=R. |last3=Pramanik |first3=S. |last4=Kar |first4=R. |year=2021 |chapter=Radiation of Flora in the Early Triassic Succeeding the End Permian Crisis: Evidences from the Gondwana Supergroup of Peninsular India |editor1=S. Banerjee |editor2=S. Sarkar |title=Mesozoic Stratigraphy of India |series=Society of Earth Scientists Series |pages=87–113 |publisher=Springer |doi=10.1007/978-3-030-71370-6_3 |isbn=978-3-030-71369-0 |s2cid=243201506 }}</ref>
|
Sp. nov
|
Valid
|
Ghosh ''et al.''
|
[[Earth:Early Triassic|Early Triassic]]
|
Panchet Formation
|
{{Flag|India}}
|
A [[Biology:Megaspore|megaspore]].
|
|-
|
''Otynisporites tarimensis''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Polypodiisporites discretus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil spores.
|
|-
|
''Psilatriletes delicatus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil spores.
|
|-
|
''Punctatosporites latrubessei''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil spores.
|
|-
|
''Rotverrusporites amazonicus''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil spores.
|
|-
|
''Stellibacutriletes''<ref name=Luntaispora />
|
Gen. et 4 sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
Genus includes new species ''S. capillaris'', ''S. gracilis'', ''S. rarus'' and ''S. solidus''.
|
|-
|
''Striatriletes inconspicuus''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Tarimispora''<ref name=Luntaispora />
|
Gen. et 2 sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
Genus includes new species ''T. auriculata'' and ''T. perfecta''.
|
|-
|
''Tricristatispora trilobata''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Tricristatispora yingmailensis''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Trileites plicatilis''<ref name=Luntaispora />
|
Sp. nov
|
Valid
|
Li ''et al.''
|
Mesozoic
|
|
{{Flag|China}}
|
|
|-
|
''Verrucatotriletes pseudovirueloides''<ref name=Passifloriidites />
|
Sp. nov
|
Valid
|
D'Apolito, Jaramillo & Harrington
|
Miocene
|
Solimões Formation
|
{{Flag|Brazil}}
|
Fossil spores.
|
|-
|}

===Palynological research===
* Strother & Foster (2021) describe an assemblage of fossil spores from the [[Earth:Ordovician|Ordovician]] ([[Earth:Tremadocian|Tremadocian]]) of {{wipe|Australia}}, representing a morphology that was intermediate morphology between confirmed land plant spores and earlier forms of uncertain phylogenetic placement, and evaluate the implications of these fossils for the knowledge of the evolution of land plants from their algal ancestors.<ref>{{Cite journal|last1=Strother |first1=P. K. |last2=Foster |first2=C. |year=2021 |title=A fossil record of land plant origins from charophyte algae |journal=Science |volume=373 |issue=6556 |pages=792–796 |doi=10.1126/science.abj2927 |pmid=34385396 |bibcode=2021Sci...373..792S |s2cid=236991210 }}</ref>
* A study on the fossil pollen record from [[Earth:New Zealand|New Zealand]], dating from 100 million years ago to the present, is published by Prebble ''et al.'' (2021), who report evidence indicating that [[Earth:Cretaceous|Cretaceous]] diversification was closely followed by an increase in flowering plants frequency, but their maximum frequency did not occur until the [[Earth:Eocene|Eocene]].<ref>{{cite journal |last1=Prebble |first1=J. G. |last2=Kennedy |first2=E. M. |last3=Reichgelt |first3=T. |last4=Clowes |first4=C. |last5=Womack |first5=T. |last6=Mildenhall |first6=D. C. |last7=Raine |first7=J. I. |last8=Crouch |first8=E. M. |year=2021 |title=A 100 million year composite pollen record from New Zealand shows maximum angiosperm abundance delayed until Eocene |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=566 |pages=Article 110207 |doi=10.1016/j.palaeo.2020.110207 |bibcode=2021PPP...56610207P |s2cid=233656482 }}</ref>
* A study on changes of abundance in spores and pollen record from the Danish Basin, and on their implications for the knowledge of the impact of the [[Biology:Triassic–Jurassic extinction event|Triassic–Jurassic extinction event]] on land plants, is published by Lindström (2021).<ref>{{cite journal |last=Lindström |first=S. |year=2021 |title=Two-phased Mass Rarity and Extinction in Land Plants During the End-Triassic Climate Crisis |journal=Frontiers in Earth Science |volume=9 |pages=Article 780343 |doi=10.3389/feart.2021.780343 |bibcode=2021FrEaS...9.1079L |doi-access=free }}</ref>
* A study on the vegetation history in the southwestern Balkans, as indicated by pollen from the sedimentary record in the Lake Ohrid extending to 1.36 million years ago, is published by Donders ''et al.'' (2021).<ref>{{Cite journal|last1=Donders |first1=T. |last2=Panagiotopoulos |first2=K. |last3=Koutsodendris |first3=A. |last4=Bertini |first4=A. |last5=Mercuri |first5=A. M. |last6=Masi |first6=A. |last7=Combourieu-Nebout |first7=N. |last8=Joannin |first8=S. |last9=Kouli |first9=K. |last10=Kousis |first10=I. |last11=Peyron |first11=O. |last12=Torri |first12=P. |last13=Florenzano |first13=A. |last14=Francke |first14=A. |last15=Wagner |first15=B. |last16=Sadori |first16=L. |year=2021 |title=1.36 million years of Mediterranean forest refugium dynamics in response to glacial–interglacial cycle strength |journal=Proceedings of the National Academy of Sciences of the United States of America |volume=118 |issue=34 |pages=e2026111118 |doi=10.1073/pnas.2026111118 |pmid=34400496 |pmc=8403972 |bibcode=2021PNAS..11826111D |doi-access=free }}</ref>

== Research ==
* A study on changes of the morphological complexity of reproductive structures of land plants throughout their evolutionary history, based on data from fossil and extant land plants, is published by Leslie, Simpson & Mander (2021).<ref>{{Cite journal|last1=Leslie |first1=A. B. |last2=Simpson |first2=C. |last3=Mander |first3=L. |year=2021 |title=Reproductive innovations and pulsed rise in plant complexity |journal=Science |volume=373 |issue=6561 |pages=1368–1372 |doi=10.1126/science.abi6984 |pmid=34529461 |bibcode=2021Sci...373.1368L |s2cid=237547326 |url=http://oro.open.ac.uk/79109/1/Leslie%20et%20al%20accepted%20manuscript.pdf }}</ref>
* Revision of [[Earth:Silurian|Silurian]] ([[Earth:Wenlock epoch|Wenlock]] to [[Earth:Pridoli epoch|Přídolí]]) assemblages of [[Biology:Polysporangiophyte|polysporangiophyte]]s with dispersed spores and [[Biology:Cryptospore|cryptospore]]s, aiming to determine the relationship between Silurian plant evolution and climate changes linked with perturbations of the global carbon cycle, is published by Pšenička ''et al.'' (2021).<ref>{{Cite journal|last1=Pšenička |first1=J. |last2=Bek |first2=J. |last3=Frýda |first3=J. |last4=Žárský |first4=V. |last5=Uhlířová |first5=M. |last6=Štorch |first6=P. |year=2021 |title=Dynamics of Silurian Plants as Response to Climate Changes |journal=Life |volume=11 |issue=9 |pages=Article 906 |doi=10.3390/life11090906 |pmid=34575055 |pmc=8470493 |bibcode=2021Life...11..906P |doi-access=free }}</ref>
* Reconstruction of the structure and development of the rooting system of ''[[Biology:Asteroxylon|Asteroxylon]] mackiei'' is presented by Hetherington ''et al.'' (2021).<ref>{{Cite journal|last1=Hetherington |first1=A. J. |last2=Bridson |first2=S. L. |last3=Jones |first3=A. L. |last4=Hass |first4=H. |last5=Kerp |first5=H. |last6=Dolan |first6=L. |year=2021 |title=An evidence-based 3D reconstruction of ''Asteroxylon mackiei'', the most complex plant preserved from the Rhynie chert |journal=eLife |volume=10 |pages=e69447 |doi=10.7554/eLife.69447 |pmid=34425940 |pmc=8384418 |doi-access=free }}</ref>
* A study on factors influencing the extent of arboreal vegetation during the [[Earth:Late Paleozoic icehouse|Late Paleozoic icehouse]] is published by Matthaeus ''et al.'' (2021), who interpret their findings as indicating that [[Earth:Pangaea|Pangaea]] could have supported widespread arboreal plant growth and forest cover based on leaf water constraints, but the forest extent was restricted because of impact of freezing on plants, and estimate that contracting forest cover increased net global [[Earth:Surface runoff|surface runoff]] by up to 6.1%.<ref>{{Cite journal|last1=Matthaeus |first1=W. J. |last2=Macarewich |first2=S. I. |last3=Richey |first3=J. D. |last4=Wilson |first4=J. P. |last5=McElwain |first5=J. C. |last6=Montañez |first6=I. P. |last7=DiMichele |first7=W. A. |last8=Hren |first8=M. T. |last9=Poulsen |first9=C. J. |last10=White |first10=J. D. |year=2021 |title=Freeze tolerance influenced forest cover and hydrology during the Pennsylvanian |journal=Proceedings of the National Academy of Sciences of the United States of America |volume=118 |issue=42 |pages=e2025227118 |doi=10.1073/pnas.2025227118 |pmid=34635589 |pmc=8594576 |bibcode=2021PNAS..11825227M |doi-access=free }}</ref>
* Description of the reproductive organs of the lycopsids from the Upper Devonian Wutong Formation (China), and a study on the ability of the [[Biology:Sporophyll|sporophyll]] units for wind dispersal, is published by Zhou ''et al.'' (2021), who name new [[Biology:Form classification|form species]] ''Lepidophylloides longshanensis'' and ''Lepidophylloides changxingensis''.<ref>{{Cite journal|last1=Zhou |first1=Y. |last2=Wang |first2=D.-M. |last3=Liu |first3=L. |last4=Huang |first4=P. |year=2021 |title=The morphometric of lycopsid sporophylls and the evaluation of their dispersal potential: an example from the Upper Devonian of Zhejiang Province, China |journal=BMC Ecology and Evolution |volume=21 |issue=1 |pages=Article number 198 |doi=10.1186/s12862-021-01933-3 |pmid=34732141 |pmc=8565055 |doi-access=free }}</ref>
* An exceptionally well preserved ''[[Biology:Brasilodendron|Brasilodendron]]''-like lycopsid forest containing over 150 upright stumps is described from an early [[Earth:Permian|Permian]] postglacial landscape of western [[Earth:Gondwana|Gondwana]] (Paraná Basin, {{wipe|Brazil}}) by Mottin ''et al.'' (2021).<ref>{{Cite journal|last1=Mottin |first1=T. E. |last2=Iannuzzi |first2=R. |last3=Vesely |first3=F. F. |last4=Montañez |first4=I. P. |last5=Griffis |first5=N. |last6=Canata |first6=R. E. |last7=Barão |first7=L. M. |last8=da Silveira |first8=D. M. |last9=Garcia |first9=A. M. |year=2021 |title=A glimpse of a Gondwanan postglacial fossil forest |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=588 |pages=Article 110814 |doi=10.1016/j.palaeo.2021.110814 |s2cid=245582665 }}</ref>
* A study on the anatomy of ''[[Biology:Stigmaria|Stigmaria]] asiatica'' is published by Chen ''et al.'' (2021).<ref>{{Cite journal|last1=Chen |first1=B.-Y. |last2=Wan |first2=M.-L. |last3=Zhou |first3=W.-M. |last4=Wang |first4=S.-J. |last5=Wang |first5=J. |year=2021 |title=Anatomy of ''Stigmaria asiatica'' Jongmans et Gothan from the Asselian (lowermost Permian) of Wuda Coalfield, Inner Mongolia, North China |journal=Palaeoworld |volume=31 |issue=2 |pages=311–323 |doi=10.1016/j.palwor.2021.05.001| issn=1871-174X |s2cid=236366665 }}</ref>
* Stump casts of ''[[Biology:Sigillaria|Sigillaria]]'', preserving traces of internal anatomy, are described from the earliest Permian Wuda Tuff (China) by D'Antonio ''et al.'' (2021).<ref>{{Cite journal|last1=D'Antonio |first1=M. P. |last2=Boyce |first2=C. K. |last3=Zhou |first3=W.-M. |last4=Pfefferkorn |first4=H. W. |last5=Wang |first5=J. |title=Primary tissues dominated ground-level trunk diameter in ''Sigillaria'': evidence from the Wuda Tuff, Inner Mongolia |year=2021 |journal=Journal of the Geological Society |volume=179 |issue=2 |doi=10.1144/jgs2021-021 |s2cid=238701484 }}</ref>
* A study aiming to determine probable causes of the world-wide proliferation of members of [[Biology:Isoetales|Isoetales]], particularly ''[[Biology:Pleuromeia|Pleuromeia]]'', during and in the aftermath of the [[Earth:Permian–Triassic extinction event|Permian–Triassic extinction event]], and evaluating the implications of this proliferation for the knowledge of environmental stresses during and in the aftermath of this extinction event, is published by Looy, van Konijnenburg-van Cittert & Duijnstee (2021).<ref>{{Cite journal|last1=Looy |first1=C. V. |last2=van Konijnenburg-van Cittert |first2=J. H. A. |last3=Duijnstee |first3=I. A. P. |year=2021 |title=Proliferation of Isoëtalean Lycophytes During the Permo-Triassic Biotic Crises: A Proxy for the State of the Terrestrial Biosphere |journal=Frontiers in Earth Science |volume=9 |pages=Article 615370 |doi=10.3389/feart.2021.615370 |bibcode=2021FrEaS...9...55L |doi-access=free }}</ref>
* New fossil material of ''Saportaea salisburioides'', providing new information on leaf morphology and growth of this plant, is described from the Permian [[Earth:Umm Irna Formation|Umm Irna Formation]] (Jordan) by Kerp ''et al.'' (2021), who interpret their findings as indicating that ''Saportaea grandifolia'' and ''Baiera virginiana'' were [[Biology:Synonym (taxonomy)|synonyms]] of ''S. salisburioides'', and possibly indicating that the [[Biology:Fructification|fructification]] belonging to the genus ''Nystroemia'' is a part of ''Saportaea''.<ref>{{Cite journal|last1=Kerp |first1=H. |last2=Blomenkemper |first2=P. |last3=Hamad |first3=A. A. |last4=Bomfleur |first4=B. |year=2021 |title=''Saportaea'' Fontaine et White 1880 – An enigmatic, long-ranging, widely distributed but rare type of late Palaeozoic and early Mesozoic foliage |journal=Review of Palaeobotany and Palynology |volume=296 |pages=Article 104542 |doi=10.1016/j.revpalbo.2021.104542 |s2cid=244595344 }}</ref>
* Description of ''[[Biology:Geinitzia (plant)|Geinitzia]] reichenbachii'' from its gross morphology to the cellular scale, and a study on the likely ecology of this conifer, is published by Moreau ''et al.'' (2021).<ref>{{Cite journal|last1=Moreau |first1=J.-D. |last2=Philippe |first2=M. |last3=Néraudeau |first3=D. |last4=Dépré |first4=E. |last5=Le Couls |first5=M. |last6=Fernandez |first6=V. |last7=Beurel |first7=S. |year=2021 |title=Paleohistology of the Cretaceous resin-producing conifer ''Geinitzia reichenbachii'' using X-ray synchrotron microtomography |journal=American Journal of Botany |volume=108 |issue=9 |pages=1745–1760 |doi=10.1002/ajb2.1722 |pmid=34495546 |s2cid=237441544 |doi-access=free }}</ref>
* A study on the evolutionary history of the family [[Biology:Cycadaceae|Cycadaceae]], based on genomic data and fossil record, is published by Liu ''et al.'' (2021).<ref>{{Cite journal|last1=Liu |first1=J. |last2=Lindstrom |first2=A. J. |last3=Marler |first3=T. E. |last4=Gong |first4=X. |year=2021 |title=Not that young: combining plastid phylogenomic, plate tectonic and fossil evidence indicates a Palaeogene diversification of Cycadaceae |journal=Annals of Botany |volume=129 |issue=2 |pages=217–230 |doi=10.1093/aob/mcab118 |pmid=34520529 |pmc=8796677 |doi-access=free }}</ref>
* Well-preserved recurved [[Biology:Calybium and cupule|cupules]] of seed plants are described from the Lower Cretaceous of {{wipe|China}} by Shi ''et al.'' (2021), who interpret the structure of these cupules as consistent with the recurved form and development of the second integument in the [[Biology:Glossary of botanical terms#bitegmic|bitegmic]] anatropous [[Biology:Ovule|ovule]]s of flowering plants, and evaluate the implications of these fossils for the knowledge of the origin of the flowering plants.<ref>{{Cite journal|last1=Shi |first1=G. |last2=Herrera |first2=F. |last3=Herendeen |first3=P. S. |last4=Clark |first4=E. G. |last5=Crane |first5=P. R. |year=2021 |title=Mesozoic cupules and the origin of the angiosperm second integument |journal=Nature |volume=594 |issue=7862 |pages=223–226 |doi=10.1038/s41586-021-03598-w |pmid=34040260 |bibcode=2021Natur.594..223S |s2cid=235217720 }}</ref>
* Taxonomically diverse flora from the Seafood Salad locality, found ~65 m below the Cretaceous-Paleogene boundary in the [[Earth:Hell Creek Formation|Hell Creek Formation]] (Montana, {{wipe|United States}}), is described by Wilson, Wilson Mantilla & Strӧmberg (2021), who study the affinities of plants of this locality and compare them with other Late Cretaceous floras of the Western Interior.<ref>{{Cite journal|last1=Wilson |first1=P. K. |last2=Wilson Mantilla |first2=G. P. |last3=Strӧmberg |first3=C. A. E. |year=2021 |title=Seafood Salad: A diverse latest Cretaceous flora from eastern Montana |journal=Cretaceous Research |volume=121 |pages=Article 104734 |doi=10.1016/j.cretres.2020.104734 |bibcode=2021CrRes.12104734W |s2cid=233543523 }}</ref>
* A study on the timing of the origin of the flowering plants, based on data from fossil record and from the diversity of extant members of this group, is published by Silvestro ''et al.'' (2021), who interpret their findings as indicating that several flowering plant families originated in the [[Earth:Jurassic|Jurassic]].<ref>{{Cite journal|last1=Silvestro |first1=D. |last2=Bacon |first2=C. D. |last3=Dong |first3=W. |last4=Zhang |first4=Q. |last5=Donoghue |first5=P. C. J. |last6=Antonelli |first6=A. |last7=Xing |first7=Y. |year=2021 |title=Fossil data support a pre-Cretaceous origin of flowering plants |journal=Nature Ecology & Evolution |volume=5 |issue=4 |pages=449–457 |doi=10.1038/s41559-020-01387-8 |pmid=33510432 |hdl=1983/11314812-19f6-4cbc-84a2-28791cb16d53 |s2cid=231763334 |url=https://research-information.bris.ac.uk/ws/files/264149147/Silvestro_et_al_AF.pdf }}</ref>
* A study on the diversity of insect damage types in fossil plants from the Cretaceous ([[Earth:Albian|Albian]] to [[Earth:Cenomanian|Cenomanian]]) Dakota Formation (United States), evaluating their implications for the knowledge of the early evolution of angiosperm florivory and associated pollination, is published by Xiao ''et al.'' (2021).<ref>{{cite journal |last1=Xiao |first1=L. |last2=Labandeira |first2=C. |last3=Dilcher |first3=D. |last4=Ren |first4=D. |year=2021 |title=Florivory of Early Cretaceous flowers by functionally diverse insects: implications for early angiosperm pollination |journal=Proceedings of the Royal Society B: Biological Sciences |volume=288 |issue=1953 |pages=Article ID 20210320 |doi=10.1098/rspb.2021.0320 |pmid=34132112 |pmc=8207559 }}</ref>
* New fossil material of ''Callianthus dilae'' is described from the Lower Cretaceous [[Earth:Yixian Formation|Yixian Formation]] (China) by Wang ''et al.'' (2021), who reconstruct the whole plant of ''Callianthus'', interpreting it as an aquatic flowering plant.<ref>{{Cite journal|last1=Wang |first1=X. |last2=Shih |first2=C. |last3=Liu |first3=Z.-J. |last4=Lin |first4=L. |last5=Singh |first5=K. J. |year=2021 |title=Reconstructing the ''Callianthus'' plant – an early aquatic angiosperm from the Lower Cretaceous of China |journal=Cretaceous Research |volume=128 |pages=Article 104983 |doi=10.1016/j.cretres.2021.104983|bibcode=2021CrRes.12804983W |issn=0195-6671 |doi-access=free }}</ref>
* A study on the anatomy of the [[Biology:Epidermis (botany)|epidermal]] features of the floating leaves of ''Quereuxia angulata'' from the Upper Cretaceous Yong'ancun Formation (China) is published by Liang ''et al.'' (2021).<ref>{{Cite journal|last1=Liang |first1=F. |last2=Tian |first2=N. |last3=Sun |first3=W. |last4=Wu |first4=Q. |last5=Liu |first5=B. |last6=Wang |first6=H. |year=2021 |title=Epidermal features of the floating leaves of ''Quereuxia angulata'' (Newberry) Krištofovič, an aquatic angiosperm from the Upper Cretaceous of Northeast China |journal=Cretaceous Research |volume=125 |pages=Article 104835 |doi=10.1016/j.cretres.2021.104835 |bibcode=2021CrRes.12504835L }}</ref>
* A study on plant extinction and ecological change in tropical forests resulting from the [[Earth:Cretaceous–Paleogene extinction event|Cretaceous–Paleogene extinction event]], based on data from fossil pollen and leaves from Colombia, is published by Carvalho ''et al.'' (2021), who report evidence indicative of a long interval of low plant diversity in the [[Biology:Neotropical realm|Neotropics]] after the end-Cretaceous extinction, and the emergence of forests with a structure resembling modern Neotropical [[Earth:Rainforest|rainforest]]s, with a closed [[Biology:Canopy|canopy]] and multistratal structure dominated by flowering plants, during the [[Earth:Paleocene|Paleocene]].<ref>{{Cite journal|last1=Carvalho |first1=M. R |last2=Jaramillo |first2=C. |last3=de la Parra |first3=F. |last4=Caballero-Rodríguez |first4=D. |last5=Herrera |first5=F. |last6=Wing |first6=S. |last7=Turner |first7=B. L. |last8=D'Apolito |first8=C. |last9=Romero-Báez |first9=M. |last10=Narváez |first10=P. |last11=Martínez |first11=C. |last12=Gutierrez |first12=M. |last13=Labandeira |first13=C. |last14=Bayona |first14=G. |last15=Rueda |first15=M. |last16=Paez-Reyes |first16=M. |last17=Cárdenas |first17=D. |last18=Duque |first18=Á. |last19=Crowley |first19=J. L. |last20=Santos |first20=C. |last21=Silvestro |first21=D. |year=2021 |title=Extinction at the end-Cretaceous and the origin of modern Neotropical rainforests |journal=Science |volume=372 |issue=6537 |pages=63–68 |doi=10.1126/science.abf1969 |pmid=33795451 |bibcode=2021Sci...372...63C |s2cid=232484243 }}</ref>
* A study on the impact of the mid-Eocene greenhouse warming event on floras from southernmost South America is published by Fernández ''et al.'' (2021).<ref>{{Cite journal|last1=Fernández |first1=D. A. |last2=Palazzesi |first2=L. |last3=González Estebenet |first3=M. S. |last4=Tellería |first4=M. C. |last5=Barreda |first5=V. D. |title=Impact of mid Eocene greenhouse warming on America's southernmost floras |year=2021 |journal=Communications Biology |volume=4 |issue=1 |pages=Article number 176 |doi=10.1038/s42003-021-01701-5 |pmid=33564110 |pmc=7873257 }}</ref>
* Evidence from middle Eocene-middle Miocene [[Earth:Tuff|tuff]]aceous deposits of central and northern Patagonia, indicating that soils, vegetation, insects and mammal herbivores began to record diverse traits related to the presence of grasslands with mosaic vegetation since middle Eocene, is presented by Bellosi ''et al.'' (2021).<ref>{{Cite journal|last1=Bellosi |first1=E. |last2=Genise |first2=J. F. |last3=Zucol |first3=A. |last4=Bond |first4=M. |last5=Kramarz |first5=A. |last6=Sánchez |first6=M. V. |last7=Krause |first7=J. M. |title=Diverse evidence for grasslands since the Eocene in Patagonia |year=2021 |journal=Journal of South American Earth Sciences |volume=108 |pages=Article 103357 |doi=10.1016/j.jsames.2021.103357 |bibcode=2021JSAES.10803357B |s2cid=235567426 }}</ref>
* A study on Middle Miocene microfloral assemblages from ten localities in the Madrid Basin ({{wipe|Spain}}), providing evidence of prevalence of open habitats with grass-dominated, savannah-like vegetation under a warm and semi-arid climatic regime in the Iberian Peninsula in the Middle Miocene, is published by Casas-Gallego ''et al.'' (2021).<ref>{{Cite journal|last1=Casas-Gallego |first1=M. |last2=Postigo-Mijarra |first2=J. M. |last3=Rivas-Carballo |first3=M. R. |last4=Valle-Hernández |first4=M. F. |last5=Morín-de Pablos |first5=J. |last6=Barrón |first6=E. |title=Early evidence of continental aridity and open-habitat grasslands in Europe as revealed by the Middle Miocene microflora of the Madrid Basin |year=2021 |journal=Palaeogeography, Palaeoclimatology, Palaeoecology |volume=581 |pages=Article 110603 |doi=10.1016/j.palaeo.2021.110603 |bibcode=2021PPP...58110603C |s2cid=238847370 }}</ref>
* Crump ''et al.'' (2021) present a record of vegetation from the Last Interglacial based on ancient DNA from lake sediment from the Baffin Island ({{wipe|Canada}}), and report evidence of major ecosystem changes in the Arctic in response to warmth, including a ~400&nbsp;km northward range shift of dwarf birch relative to today.<ref>{{Cite journal|last1=Crump |first1=S. E. |last2=Fréchette |first2=B. |last3=Power |first3=M. |last4=Cutler |first4=S. |last5=de Wet |first5=G. |last6=Raynolds |first6=M. K. |last7=Raberg |first7=J. H. |last8=Briner |first8=J. P. |last9=Thomas |first9=E. K. |last10=Sepúlveda |first10=J. |last11=Shapiro |first11=B. |last12=Bunce |first12=M. |last13=Miller |first13=G. H. |year=2021 |title=Ancient plant DNA reveals High Arctic greening during the Last Interglacial |journal=Proceedings of the National Academy of Sciences of the United States of America |volume=118 |issue=13 |pages=e2019069118 |doi=10.1073/pnas.2019069118 |pmid=33723011 |pmc=8020792 |bibcode=2021PNAS..11819069C |doi-access=free }}</ref>

==Deaths==
*Alan Graham (1934–2021), passed away on 8 July 2021.  Graham earned his PhD in 1962 under the guidance of Chester A. Arnold, and was noted for a career studying the Cenozoic paleobotany of the Caribbean and Central America. <ref name="GrahamObit">{{cite journal |last1=Jaramillo |first1=C. |last2=Jarzen |first2=D. M. |title=Alan Keith Graham (1934–2021) |journal=Palynology |date=2022 |volume=46 |issue=1 |pages=1–4 |doi=10.1080/01916122.2021.1971121 |bibcode=2022Paly...4671121J |s2cid=238700678 |url=https://www.tandfonline.com/doi/full/10.1080/01916122.2021.1971121|doi-access=free }}</ref>

== References ==
{{reflist|30em}}

[[Category:Paleobotany]]

{{Sourceattribution|2021 in paleobotany}}