View source for Biology:Larix laricina

{{Short description|Species of larch native to North America}}
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{{Speciesbox
| image = Larix laricina.jpg
| image_caption = Tamarack larch in fall colors, with black spruce in the background
| status = LC
| status_system = IUCN3.1
| status_ref = <ref name="iucn status 19 November 2021">{{cite iucn |author=Farjon, A. |date=2013 |title=''Larix laricina'' |volume=2013 |page=e.T42313A2971618 |doi=10.2305/IUCN.UK.2013-1.RLTS.T42313A2971618.en |access-date=19 November 2021}}</ref>
| genus = Larix
| species = laricina
| authority = (Du Roi) K. Koch
| range_map = Larix laricina range map 3.png
| range_map_caption = Natural range of ''Larix laricina''
| synonyms_ref = <ref>{{ThePlantList |authority= (Du Roi) K.Koch}}</ref>
| synonyms =
*''Abies microcarpa'' <small>(Lamb.) Lindl.</small>
*''Larix alaskensis'' <small>W.Wight</small>
*''Larix americana'' <small>Michx.</small>
*''Larix fraseri'' <small>Curtis ex Gordon</small>
*''Larix intermedia'' <small>(Du Roi) Lodd. ex J.Forbes</small>
*''Larix microcarpa'' <small>(Lamb.) J.Forbes</small>
*''Larix rubra'' <small>Steud.</small>
*''Larix tenuifolia'' <small>Salisb.</small>
*''Pinus intermedia'' <small>Du Roi</small>
*''Pinus laricina'' <small>Du Roi</small>
*''Pinus microcarpa'' <small>Lamb.</small>
}}

'''''Larix laricina''''', commonly known as the '''tamarack''',<ref name=GD>{{Gymnosperm Database |family=Pinaceae |genus=Larix |species=laricina |access-date = 2011-05-28}}</ref> '''hackmatack''',<ref name=GD /> '''eastern larch''',<ref name=GD /> '''black larch''',<ref name=GD /> '''red larch''',<ref name=GD /> or '''American larch''',<ref name=GD /> is a species of [[Biology:Larch|larch]] native to {{wipe|Canada}}, from eastern [[Earth:Yukon|Yukon]] and Inuvik, [[Earth:Northwest Territories|Northwest Territories]] east to Newfoundland, and also south into the upper northeastern United States from Minnesota to Cranesville Swamp, West Virginia; there is also an isolated population in central [[Earth:Alaska|Alaska]].<ref>{{BONAP|ref |genus=Larix |species=laricina |state=1}}</ref><!-- The population is isolated. The distribution is disjunct.-->

== Description ==
''Larix laricina'' is a small to medium-size boreal [[Biology:Deciduous|deciduous]] [[Biology:Conifer|conifer]] [[Earth:Tree|tree]] reaching {{convert|15|-|23|m|abbr=on|ft}} tall, with a trunk up to {{convert|60|cm|in|abbr=on}} diameter.<ref name="FEIS3">{{FEIS|genus=Larix|species=laricina|type=tree|last=Uchytil|first=Ronald J.|year=1991}}</ref> The bark of mature trees are reddish, the young trees are gray with smooth bark.<ref name=":02">{{eFloras|1|233500743|Larix laricina|family=Pinaceae|first=William H.|last=Parker}}</ref> The [[Biology:Leaf|leaves]] are needle-like, {{convert|2.5|cm|in||abbr=on|frac=4|}} short, light blue-green, turning bright yellow before they fall in the autumn, leaving the shoots bare until the next spring.<ref name="FEIS3" /> The needles are produced in clusters on long woody spur shoots.<ref name="FEIS3" /> The [[Conifer cone|cones]] are the smallest of any larch, only {{convert|1|-|2.3|cm|abbr=on|frac=16|in}} long, with 12-25 seed scales; they are bright red, turning brown and opening to release the [[Biology:Seed|seeds]] when mature, 4 to 6 months after [[Biology:Pollination|pollination]].<ref name=":02" />

Key characteristics:<ref>{{cite book |last1=Barnes |first1=Burton V. |url=https://archive.org/details/michigantreesgui00barn |title=Michigan Trees |last2=Wagner Jr. |first2=Warren H. |date=September 15, 1981 |publisher=[[Organization:University of Michigan Press|University of Michigan Press]] |isbn=978-0-472-08018-2 |url-access=registration}}</ref>

* The needles are normally borne on a short shoot in groups of 10–20 needles.
* The larch is deciduous and the needles turn yellow in autumn.
* The seed cones are small, less than {{convert|2|cm|in|abbr=on|frac=4}} long, with lustrous brown scales.
* Larch are commonly found in swamps, fens, bogs, and other low-land areas.

== Distribution and ecology ==
[[File:Larix laricina foliagecones.jpg|left|thumb|Tamarack larch foliage and cones in August. The lighter brown cones are from the current season; the darker brown cones are mature cones from previous seasons.]]
Tamaracks are very cold tolerant, able to survive temperatures down to at least {{convert|-62|C|F|0}}, and commonly occurs at the Arctic [[Earth:Tree line|tree line]] at the edge of the [[Earth:Tundra|tundra]]. Trees in these severe climatic conditions are smaller than farther south, often only {{convert|3|m|ft|round=5|abbr=on}} tall. They can tolerate a wide range of soil conditions but grow most commonly in [[Earth:Swamp|swamp]]s, [[Earth:Bog|bog]]s, or [[Earth:Muskeg|muskeg]]s, in wet to moist organic soils such as [[Biology:Sphagnum|sphagnum]], [[Earth:Peat|peat]], and woody peat.<ref name="GD" /><ref name=":02" /> <ref name="FEIS3" />They are also found on mineral soils that range from heavy clay to coarse sand; thus texture does not seem to be limiting. Although tamarack can grow well on [[Earth:Calcareous|calcareous]] soils, it is not abundant on the [[Earth:Limestone|limestone]] areas of eastern Ontario.<ref name="FEIS3" />

[[File:2015-08-20 17 12 36 Tamarack seedling in a sphagnum bog adjacent to Taborton Road (Rensselaer County Route 42) in Sand Lake, New York.jpg|thumb|upright|Tamarack sapling in a sphagnum bog]]
Tamarack is generally the first forest tree to grow on filled-lake bogs.<ref name="FEIS3" /><ref name=":2" /> In the lake states, tamarack may appear first in the [[Biology:Cyperaceae|sedge]] mat, sphagnum moss, or not until the bog shrub stage. Farther north, it is the pioneer tree in the bog shrub stage. Tamarack is fairly well adapted to reproduce successfully on burns, so it is one of the common [[Earth:Pioneer species|pioneers]] on sites in the boreal forest immediately after a fire.<ref name=":1">{{cite web|url=http://globaltwitcher.auderis.se/artspec_information.asp?thingid=44751 |title=Black Spruce |first=C. Michael |last=Hogan |work=GlobalTwitcher.com |date=November 24, 2008 |access-date=2011-05-28 |url-status=dead |archive-url=https://web.archive.org/web/20111005174426/http://globaltwitcher.auderis.se/artspec_information.asp?thingid=44751 |archive-date=October 5, 2011 }}</ref>

The central [[Earth:Alaska|Alaska]]n population, separated from the eastern [[Earth:Yukon|Yukon]] populations by a gap of about {{convert|700|km|mi}}, is treated as a distinct variety ''Larix laricina'' var. ''alaskensis'' by some botanists, though others argue that it is not sufficiently distinct to be distinguished.<ref name="FEIS3" />

=== Damaging agents ===
Tamaracks are easily susceptible to fires, as they have shallow roots and thin bark.<ref name="FEIS3" /> The tamarack's shallow root system also leaves the susceptible to being knocked over by high-speed winds. It has also been discovered that abnormally high water levels often kill tamarack stands.<ref name=":2" /> Flooding, mainly caused by beaver dams and newly constructed roads, can kill off stands and damage adventitious roots.<ref name="FEIS3" /><ref name=":2" />

Tamaracks are targeted by many species of insects. One of the most prominent damaging insect is larch sawfly, which is non-native.<ref name="FEIS3" /><ref name=":2" /> They cause damage across their range and cause defoliation which can kill the trees within 6 to 9 years.<ref name="FEIS3" /><ref name=":2" /> To lessen the problem parasites have been imported to kill the larch sawflies in parts of Minnesota and Manitoba.<ref name=":2" /> Another serious defoliator is the larch casebearer (''[[Biology:Coleophora laricella|Coleophora laricella]])''.  All tamaracks are susceptible to being killed by the larch casebearer, however recently the outbreaks of larch casebearer have been less severe.<ref name=":2" />

There are some other insects that can harm Tamaracks, including [[Biology:Spruce budworm|spruce budworm]] (''[[Biology:Choristoneura fumiferana|Choristoneura fumiferana]]''), the larch-bud moth (''[[Biology:Zeiraphera improbana|Zeiraphera improbana]]''), the spruce spider mite (''[[Biology:Oligonychus ununguis|Oligonychus ununguis]]'') the larch-shoot moth ''([[Biology:Argyresthia laricella|Argyresthia laricella]]),'' and the eastern larch beetle (''[[Biology:Dendroctonus simplex|Dendroctonus simplex]]'').<ref name=":2" /> Healthy trees are left mostly unaffected by eastern larch beetles.<ref name=":2" /> Defoliation by the larch casebearer makes infestation of the eastern larch beetle more likely.<ref name="McKee2">{{cite thesis |last=McKee |first=Fraser |date=2015 |title=Biology and population dynamics of the eastern larch beetle, Dendroctonus simplex LeConte, and its interactions with eastern larch (tamarack), Larix laricina. |url=https://conservancy.umn.edu/handle/11299/175514 |type=PhD |chapter=1.2 Factors associated with the increased tree-killing activity of eastern larch beetles |publisher=University of Minnesota}}</ref><ref name=":2" />

Only one of the many pathogens that affect Tamarack cause diseases serious enough to have an economic impact on its culture, the ''[[Biology:Lachnellula willkommii|Lachnellula willkommii]]'' fungus. It is a relatively new pathogen in Canada, first recorded in 1980 and originating in Europe. The fungus cause large [[Biology:Canker|cankers]] to form and a disease known as [[Biology:Larch#Diseases|larch canker]] which is particularly harmful to the tamarack larch, killing both young and mature trees.<ref>[http://www.exoticpests.gc.ca/es-details/disease/1000096 European larch canker] Natural Resources Canada</ref> Rust is the only common foliage disease amongst Tamaracks, and cause minimal damage to the trees.<ref name=":2" /> The needle-cast fungus ''(Hypodermella laricis)'' is also cause for concern in Tamaracks.<ref name=":2" />

=== Associated forest cover ===
[[File:Tamarack.jpg|left|thumb|upright|Young tree with fall color]]
Tamarack forms extensive pure stands in the boreal region of {{wipe|Canada}} and in northern Minnesota.<ref name="FEIS3" /> In the rest of its United States range and in the Maritime Provinces, tamarack is found locally in both pure and mixed stands.<ref name="FEIS3" />

Black spruce (''Picea mariana'') is usually tamarack's main associate in mixed stands on all sites. Other commonly associated overgrowth species include balsam fir (''Abies balsamea''), white spruce (''Picea glauca''), and quaking aspen (''Populus tremuloides'') in the boreal region.<ref name="FEIS3" /> In the better organic soil sites in the northern forest region, the most common associates are the northern white-cedar (''Thuja occidentalis''), balsam fir, [[Biology:Fraxinus nigra|black ash]] (''Fraxinus nigra''), and red maple (''Acer rubrum'').<ref name="FEIS3" /> In [[Earth:Alaska|Alaska]], quaking aspen and tamarack are almost never found together.<ref name="FEIS3" /> Additional common associates are American elm (''Ulmus americana''), balsam poplar (''Populus balsamifera''), [[Biology:Jack pine|jack pine]] (''Pinus banksiana''), paper birch (''Betula papyrifera''), Kenai birch (''B. papyrifera'' var. ''kenaica''), and yellow birch (''B. alleghaniensis'').<ref name="FEIS3" />

There are a vast number of shrubs associated with Tamarack due to their range, some of the common ones are dwarf and swamp birch (''Betula glandulosa'' and ''Betula pumila''), [[Biology:Willow|willows]] (''Salix'' spp.), speckled alder (''Alnus rugosa''), and red-osier dogwood (''Cornus stolonifera'') bog Labrador tea (''Ledum groenlandicum''), bog-rosemary (''Andromeda glaucophylla''), leather leaf (''Chamaedaphne calyculata''), blueberries and huckleberries (''Vaccinium spp.)'' and small cranberry (''Vaccinium oxycoccos'').<ref name="FEIS3" /><ref name=":2" /> Characteristically the herbaceous cover includes [[Biology:Carex|sedges]] ''(Carex'' spp.), cottongrass ''(Eriophorum'' spp.), three-leaved false Solomonseal (''Maianthemum trifolium''), marsh cinquefoil (''Potentilla palustris''), [[Biology:Caltha palustris|marsh-marigold]] (''Caltha palustris''), and bogbean (''Menyanthes trifoliata'').<ref name=":2" />

== Uses ==
[[File:Larix laricina bonsai by Nick Lenz.jpg|thumb|upright|''Larix laricina'' bonsai]]
The [[Biology:Wood|wood]] is tough and durable, but also flexible in thin strips, and was used by the Algonquian people for making [[Engineering:Snowshoe|snowshoe]]s and other products where toughness was required. The natural crooks located in the stumps and roots are also preferred for creating [[Engineering:Knee (construction)|knees]] in wooden boats.<ref name=":2" /> Currently, the wood is used principally for pulpwood, but also for posts, poles, rough lumber, and fuelwood; it is not a major commercial timber species.<ref name=":2" /> Tamarack wood is also used as kickboards in horse stables.<ref name="FEIS3" /> Older log homes built in the 19th century sometimes incorporated tamarack along with other species like red or white oak. The hewn logs have a coarse grainy surface texture.

It is also grown as an ornamental tree in gardens in cold regions. Several dwarf cultivars have been created that are available commercially.<ref>{{cite web |title=Larix laricina |url=http://www.hort.uconn.edu/plants/l/larlar/larlar1.html |publisher=University of Connecticut |quote=Dwarf forms include: 'Blue Sparkler', with bluish foliage; 'Deborah Waxman', which reaches 4' in time; 'Lanark', which grows very low and wide; and 'Newport Beauty', a tiny form probably never exceeding 2' tall and wide. |access-date=2012-11-15 |archive-url=https://web.archive.org/web/20130315184441/http://www.hort.uconn.edu/plants/l/larlar/larlar1.html |archive-date=2013-03-15 |url-status=dead }}</ref><ref>{{cite encyclopedia |title=Larix Laricina: Cultivar List |url=http://coniferworld.com/listing.php?x=l&a=Larix&b=laricina |publisher=Royal Horticultural Society |encyclopedia=Encyclopedia of Conifers}}</ref> Tamarack is commonly used for [[Biology:Bonsai|bonsai]].<ref>{{cite book |title=The Art of Natural Bonsai: Replicating Nature's Beauty |first=David |last=Joyce |publisher=Sterling Publishing Company |year=2006 |isbn=978-1-4027-3524-0 |page=[https://archive.org/details/artofnaturalbons0000joyc/page/154 154] |quote=As bonsai, they are my favorite genus because of their speed of growth, hardiness, ease of wiring and shaping, and, most of all, for their beautiful foliage color in spring and autumn. |url=https://archive.org/details/artofnaturalbons0000joyc|url-access=registration }}</ref>

Tamarack poles were used in corduroy roads because of their resistance to rot. Tamarack posts were used before 1917 in Alberta to mark the northeast corner of sections surveyed within townships. They were used by the surveyors because at that time the very rot-resistant wood was readily available in the bush and was light to carry.{{citation needed|date=November 2012}} Their rot resistance was also why they were often used in early water distribution systems.

The aboriginal peoples of Canada's northwest regions used the inner bark as a poultice to treat cuts, infected wounds, [[Medicine:Frostbite|frostbite]], boils and hemorrhoids. The outer bark and roots are also said to have been used with another plant as a treatment for [[Medicine:Arthritis|arthritis]], cold and general aches and pains.<ref>{{cite book |title=Aboriginal Plant Use in Canada's Northwest Boreal Forest |first=Robin James |last=Marles |publisher=Canadian Forest Service |year=2009 |isbn=978-0-660-19869-9 |page=<!--missing-->}}</ref>

Wildlife use the tree for food and nesting. Porcupines eat the inner bark, snowshoe hares feeds on tamarack seedlings, and red squirrels eat the seeds.<ref name=":2">{{Silvics |volume=1 |genus=Larix |species=laricina |first=William F. |last=Johnston |access-date=2015-12-04}}</ref> Birds that frequent tamaracks during the summer include the white-throated sparrow, song sparrow, veery, common yellowthroat, and Nashville warbler.<ref>Dawson, Deanna K. 1979. Bird communities associated with succession and management of lowland conifer forests. ''In'' Management of north central and northeastern forests for nongame birds: workshop proceedings, 1979. p. 120-131. USDA Forest Service, General Technical Report NC-51. North Central Forest Experiment Station, St. Paul, MN.</ref><ref name=":2" />

== Reaction to competition ==
Tamarack is very intolerant of shade. <ref name=":2" />Although it can tolerate some shade during the first several years, it must become dominant to survive.<ref name=":2" /> When mixed with other species, it must be in the over story.<ref name="FEIS3" /> The tree is a good self-pruner, and boles of 25- to 30-year-old trees may be clear for one-half or two-thirds their length.<ref name="FEIS3" /><ref name=":2" />

Because tamarack is very shade-intolerant, it does not become established in its own shade.<ref name=":2" /> Consequently, the more tolerant black spruce eventually succeeds tamarack on poor bog sites, whereas northern white-cedar, balsam fir, and swamp hardwoods succeed tamarack on good swamp sites.<ref name="FEIS3" /><ref name=":2" /> Recurring [[Biology:Sawfly|sawfly]] outbreaks throughout the range of tamarack have probably sped the usual succession to black spruce or other associates.<ref name=":2" />

Various tests on planting and natural reproduction indicate that competing vegetation hinders tamarack establishment.<ref name=":2" />

The shade-intolerance of tamarack dictates the use of even-aged management.<ref name="FEIS3" /><ref name=":2" /> Some adaptation of clear cutting or seed-tree cutting is generally considered the best silvicultural system because tamarack seeds apparently germinate better in the open, and the seedlings require practically full light to survive and grow well.<ref name=":2" /> Tamarack is also usually wind-firm enough for the seed-tree system to succeed. Satisfactory reestablishment of tamarack, however, often requires some kind of site preparation, such as slash disposal and [[Biology:Herbicide|herbicide]] spraying.<ref name=":2" />

== Names ==

The names ''tamarack'' and ''hackmatack'' appear to derive from [[Social:Algonquian languages|Algonquian]] but have undergone contamination with the word ''tacamahac'', from [[Social:Nahuatl|Nahuatl]], so the precise words that underlie them are unclear.<ref>{{cite web|title=hackmatack, n., Etymology|url=https://www.oed.com/dictionary/hackmatack_n?tab=etymology#2163959|website=Oxford English Dictionary|date=July 2023|access-date=24 September 2023}}</ref> The word ''akemantak'' meaning "wood used for snowshoes" has been cited{{by whom|date=September 2023}} as a name for the species, but the Proto-Algonquian ''*a·kema·xkwa'' this appears to represent was the name for the [[Biology:Fraxinus americana|white ash]].<ref>{{cite journal|last1=Siebert|first1=Frank T.|date=1967|title=The original home of the Proto-Algonquian people|url=https://ojs.library.carleton.ca/index.php/ALGQP/article/view/427|journal=Algonquian Papers-Archive|volume=1|access-date=24 September 2023}}</ref>

== Gallery ==
<gallery mode="packed" heights="200">
File:Larix laricina female cone.jpg|Young female cone
File:Larix laricina old cones.jpg|Old seed cones
</gallery>

== See also ==
* [[Engineering:Larch ball|Larch ball]]

== References ==
{{Reflist}}
* https://srs.fs.usda.gov/pubs/misc/ag_654/volume_1/larix/laricina.htm

== External links ==
{{Commons|Larix laricina}}
* [https://arboretum.harvard.edu/plants/image-search/?keyword=Larix+laricina&submit=Search ''Larix laricina'' images at the Arnold Arboretum of Harvard University Plant Image Database]
* Enzenbacher, Tiffany. [https://arboretum.harvard.edu/stories/plant-collecting-in-the-wisconsin-wilds/ "Plant Collecting in the Wisconsin Wilds"]. Arnold Arboretum of Harvard University website, 30 August 2017. Accessed 12 May 2020.
* [http://www.rook.org/earl/bwca/nature/trees/larixlar.html Earl J. S. Rook, Boundary Waters Compendium, Flora, Fauna, Earth, and Sky, The Natural History of the Northwoods, Trees of the Northwoods, ''Larix laricina'']

{{Taxonbar|from=Q734085}}

[[Category:Larix|laricina]]

{{Sourceattribution|Larix laricina}}