View source for Biology:Timeline of dromaeosaurid research

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[[File:Deinonychus (Raptor Prey Restraint).jpg|thumb|right|upright|350px|Artistic restoration of ''Deinonychus'' preying on ''Zephyrosaurus'']]
This '''timeline of dromaeosaurid research''' is a chronological listing of events in the history of [[Biology:Paleontology|paleontology]] focused on the dromaeosaurids, a group of sickle-clawed, [[Biology:Bird|bird]]-like theropod dinosaurs including animals like ''Velociraptor''. Since the Native Americans of Montana used the sediments of the [[Earth:Cloverly Formation|Cloverly Formation]] to produce pigments, they may have encountered remains of the dromaeosaurid ''Deinonychus'' hundreds of years before these fossils came to the attention of formally trained scientists.<ref name="mayor-crow-272-273" />


After Ostrom's initial research on ''Deinonychus'', evidence continued to mount for a close evolutionary relationship between dromaeosaurids and birds.<ref name="norell-makovicky-systevo-206" /> The dromaeosaurid ''Sinornithosaurus milennii'', described in 1999 by Xu, Wang, and Wu, is a notable example as the fine-grained Chinese [[Earth:Limestone|limestone]] from which it was collected preserved its [[Biology:Feathered dinosaur|life covering of feathers]].<ref name="norell-makovicky-intro-196" /> Discoveries of feathered dromaeosaurids continued into the 2000s. Xu, Zhou, and Wang named the new genus ''[[Biology:Microraptor|Microraptor]]'' in 2000.<ref name="norell-makovicky-table-198" /> Three years later, Xu and others would report a new species in this genus that exhibited a bizarre "four winged" body plan with long pennaceous flight feathers on both its front and hind limbs.<ref name="xu-etal-inpassim-335-340" />

==Prescientific==
[[File:Cloverly Fm.jpg|thumb|right|upright|An outcrop of the Cloverly Formation]]
* The Crow people and other Native American groups inhabiting Montana used to use rocks from the [[Earth:Cloverly Formation|Cloverly Formation]] to make red pigments. Since the red pigments are richest in the same layers of the formation that preserve dinosaur fossils, it is likely that Native Americans encountered ''Deinonychus'' fossils long before scientifically trained paleontologists.<ref name="mayor-crow-272-273" />

==19th century==

===1880s===
'''<span style="color:#14145C;">1887</span>'''{{anchor|1887}}

* Harry Govier Seeley described the new species ''Ornithodesmus cluniculus''.<ref name="norell-makovicky-table-199" />

==20th century==
[[File:Dromaeosaurus skeleton.JPG|thumb|right|upright|Skeletal reconstruction of ''Dromaeosaurus'']]

===1910s===
'''<span style="color:#14145C;">1914</span>'''{{anchor|1914}}
* [[Biography:Barnum Brown|Barnum Brown]] collected a partial skull and foot in [[Earth:Late Cretaceous|Late Cretaceous]] rocks that would later serve as the type specimen of ''Dromaeosaurus albertensis''.<ref name="norell-makovicky-intro-196" />

===1920s===
[[File:Velociraptor mongoliensis AMNH 6515.jpg|thumb|right|upright|The type skull of ''Velociraptor mongoliensis'']]
'''<span style="color:#14145C;">1922</span>'''{{anchor|1922}}
* Matthew and Brown described the new genus and species ''Dromaeosaurus albertensis'' and named the [[Biology:Dromaeosauridae|Dromaeosauridae]].<ref name="norell-makovicky-table-198" />

'''<span style="color:#14145C;">1924</span>'''{{anchor|1924}}
* Henry Fairfield Osborn described the new genus and species ''Velociraptor mongoliensis''.<ref name="norell-makovicky-table-199" />

'''<span style="color:#14145C;">1926</span>'''{{anchor|1926}}
* Matthew and Brown first observed that the braincase of dromaeosaurids were large for reptiles of their body size.<ref name="norell-makovicky-paleoeco-209" />

===1960s===
[[File:Deinonychus ewilloughby.png|thumb|right|upright|Artistic restoration of ''Deinonychus'']]
'''<span style="color:#14145C;">1969</span>'''{{anchor|1969}}
* John Ostrom described the new genus and species ''Deinonychus antirrhopus''.<ref name="norell-makovicky-table-198" /> This description provided the [[Scientific literature|scientific literature]] with its first detailed information about the anatomy of the dromaeosaurid body, as previous work focused on the skull.<ref name="norell-makovicky-intro-196" /> He proposed that ''Deinonychus'' killed its prey with the enlarged, sharply curved claws on the second toe of its feet.<ref name="norell-makovicky-paleoeco-209" />
* Ostrom further described the anatomy of ''Deinonychus antirrhopus'', providing further details about the anatomy of its body.<ref name="norell-makovicky-intro-196" /> Ostrom proposed that ''Deinonychus'' only walked on two of its toes.<ref name="norell-makovicky-paleoeco-209" />

===1970s===
[[File:Fighting dinosaurs (1).jpg|thumb|right|The "Fighting Dinosaurs" specimen of ''Velociraptor mongoliensis'' and ''Protoceratops andrewsi'']]
'''<span style="color:#14145C;">1972</span>'''{{anchor|1972}}
* Zofia Kielan-Jaworowska and Rinchen Barsbold reported the associated remains of a ''Velociraptor'' and ''[[Biology:Protoceratops|Protoceratops]]'' apparently killed and preserved while fighting.<ref name="norell-makovicky-paleoeco-209" />

'''<span style="color:#14145C;">1973</span>'''{{anchor|1973}}
* Ostrom first discussed the apparent close evolutionary relationship between ''Deinonychus antirrhopus'' and [[Biology:Bird|bird]]s.<ref name="norell-makovicky-systevo-206" />

'''<span style="color:#14145C;">1975</span>'''{{anchor|1975}}
* Ostrom published further arguments building a case for a close evolutionary relationship between dromaeosaurids and birds.<ref name="norell-makovicky-systevo-206" />

[[File:Saurornitholestes digging Burrows wahweap.jpg|thumb|right|Artist's restoration of ''Saurornitholestes'' feeding on a mammal]]
'''<span style="color:#14145C;">1976</span>'''{{anchor|1976}}
* Ostrom published further arguments building a case for a close evolutionary relationship between dromaeosaurids and birds.<ref name="norell-makovicky-systevo-206" />

'''<span style="color:#14145C;">1978</span>'''{{anchor|1978}}
* Sues described the new genus and species ''Saurornitholestes langstoni''.<ref name="norell-makovicky-table-198" />

'''<span style="color:#14145C;">1979</span>'''{{anchor|1979}}
* Powell described the new genus and species ''Unquillosaurus ceibalii''.<ref name="norell-makovicky-table-199" />

===1980s===
[[File:Adasaurus Restoration.jpg|thumb|right|upright|Artistic restoration of ''Adasaurus mongoliensis'']]
'''<span style="color:#14145C;">1981</span>'''{{anchor|1981}}
* Jensen described the new genus and species ''Palaeopteryx thomsoni''.<ref name="norell-makovicky-table-199" />

'''<span style="color:#14145C;">1982</span>'''{{anchor|1982}}
* Osmolska described the new genus and species ''Hulsanpes perlei''.<ref name="norell-makovicky-table-199" />

'''<span style="color:#14145C;">1983</span>'''{{anchor|1983}}
[[File:Archaeopteryx NT.jpg|thumb|right|upright|''Archaeopteryx'']]
* Rinchen Barsbold divided the [[Biology:Dromaeosauridae|Dromaeosauridae]] into two subfamilies; the Dromaeosaurinae and Velociraptorinae.<ref name="norell-makovicky-systevo-206" />
* Barsbold described the new genus and species ''Adasaurus mongoliensis''.<ref name="norell-makovicky-table-198" /> Unlike most dromaeosaurs, famous for the "killing claws" on their second toe, the second toe claw of ''A. mongoliensis'' was relatively small and not sharply curved. It is similar to those of troodontids.<ref name="norell-makovicky-paleoeco-209" />

'''<span style="color:#14145C;">1985</span>'''{{anchor|1985}}
* Ostrom published further arguments building a case for a close evolutionary relationship between dromaeosaurids and birds.<ref name="norell-makovicky-systevo-206" />

'''<span style="color:#14145C;">1986</span>'''{{anchor|1986}}
* Gauthier found strong support for Ostrom's hypothesis that dromaeosaurids were closely related to birds in "a pioneering and thorough analysis".<ref name="norell-makovicky-systevo-206" />

===1990s===
[[File:Utahraptor scale.png|thumb|right|upright|Illustration of a ''Utahraptor'' with a human to scale]]
'''<span style="color:#14145C;">1990</span>'''{{anchor|1990}}
* Ostrom published further arguments building a case for a close evolutionary relationship between dromaeosaurids and birds.<ref name="norell-makovicky-systevo-206" />

'''<span style="color:#14145C;">1993</span>'''{{anchor|1993}}
* Kirkland, Burge, and Gaston described the new genus and species ''Utahraptor ostrommaysi''.<ref name="norell-makovicky-table-198" />
* Osmolska challenged Kielan-Jaworowska and Barsbold's 1972 interpretation of an associated skeleton of ''Velociraptor'' and ''Protoceratops'' as being of two animals killed and preserved while locked in mortal combat. She reinterpreted the association as a ''Velociraptor'' that had been killed while scavenging an already dead ''Protoceratops''.<ref name="norell-makovicky-paleoeco-209" />
[[File:Tenontosaurus BW.jpg|thumb|right|upright|''Tenontosaurus'' may have been the prey of ''Deinonychus'']]
'''<span style="color:#14145C;">1994</span>'''{{anchor|1994}}
* Ostrom speculated that ''Deinonychus'' hunted in packs to subdue prey larger than itself based on an association between several ''Deinonychus'' skeletons and a specimen of the [[Biology:Ornithopod|ornithopod]] ''[[Biology:Tenontosaurus|Tenontosaurus]]''.<ref name="norell-makovicky-paleoeco-209" />

'''<span style="color:#14145C;">1995</span>'''{{anchor|1995}}
* Currie followed Barsbold's division of [[Biology:Dromaeosauridae|Dromaeosauridae]] into Dromaeosaurinae and Velociraptorinae. However, he proposed differing definitions for these subfamilies and argued that they had different members than were included under Barsbold's classification scheme.<ref name="norell-makovicky-systevo-206" />
* Maxwell and Ostrom reported on the association of ''Deinonychus'' teeth and ''Tenontosaurus'' remains and argued that these provided further support for the hypothesis that ''Deinonychus'' was a pack hunter.<ref name="norell-makovicky-paleoeco-209" />
* Unwin and others followed the fighting-to-the-death interpretation of the fighting dinosaurs specimen.<ref name="norell-makovicky-paleoeco-209" />
[[File:Unenlagia comahuensis.jpg|thumb|right|upright|Artist's restoration of ''Unenlagia'']]
'''<span style="color:#14145C;">1997</span>'''{{anchor|1997}}

*Novas and Puerta described the new genus and species ''Unenlagia comahuensis''.<ref name="norell-makovicky-table-198" />
* Chatterjee argued that dromaeosaurids lived in trees based on the anatomy of their hands and the backward orientation of the pubic bones in their pelvis.<ref name="norell-makovicky-paleoeco-209" />
* Reid examined histological sections of the bones of ''Saurornitholestes'' to see whether or not they were consistent with the idea that dromaeosaurids were warm blooded. However, the results of his study were inconclusive.<ref name="norell-makovicky-paleoeco-209" />
* Ruben and others attempted to reconstruct the way air would flow through the snouts of dromaeosaurids during respiration to see whether or not it was consistent with the airflow patterns of modern cold or warm-blooded animals. They found that the probable course of airflow through the snout of a living, breathing dromaeosaurid would have been more consistent with that of a cold blooded animal. However, the anatomical reconstructions the researchers used to draw these conclusions have been criticized for positioning the choana too far forward and for excluding the animals' secondary palate.<ref name="norell-makovicky-paleoeco-209" />

'''<span style="color:#14145C;">1998</span>'''{{anchor|1998}}
* Le Leouff and Buffetaut described the new genus and species ''Variraptor mechinorum''.<ref name="norell-makovicky-table-199" />
* Novas described the new genus and species ''Megaraptor namunhuaiquii''.<ref name="norell-makovicky-table-199" />
[[File:Sinornithosaurus IVPP V12811.jpg|thumb|right|upright|Feathered type specimen of ''Sinornithosaurus'']]
* D. L. Brinkman and others reinterpreted the association between several ''Deinonychus'' specimens and a ''Tenontosaurus'' as preserving the aftermath of an ancient feeding frenzy rather than as evidence of pack hunting behavior in ''Deinonychus'' as argued by Ostrom.<ref name="norell-makovicky-paleoeco-209" />

'''<span style="color:#14145C;">1999</span>'''{{anchor|1999}}
* Clark and others argued that it was impossible to know whether or not ''Velociraptor'' was a predator of ''Protoceratops'' based on the famous "fighting dinosaurs" specimen, since this was only one specimen.<ref name="norell-makovicky-paleoeco-209" />
* Altangerel Perle, Norell and Clark described the new genus and species ''Achillobator giganticus''.<ref name="norell-makovicky-table-198" />
* Xu, Wang, and Wu described the new genus and species ''Sinornithosaurus millenii''.<ref name="norell-makovicky-table-198" /> The specimen actually preserved the remains of the [[Biology:Feathered dinosaur|feathers]] that covered the animal in life. The presence of a feathery covering is consistent with the idea that dromaeosaurids were warm-blooded.<ref name="norell-makovicky-paleoeco-209" /> They also performed a cladistic analysis of the [[Biology:Dromaeosauridae|Dromaeosauridae]], finding more support for dividing the family into Dromaeosaurinae and Velociraptorinae. The researchers found both groups to be monophyletic clades, with ''Sinornithosaurus milenii'' the sister group to both, and therefore the most primitive known dromaeosaurid.<ref name="norell-makovicky-systevo-207" />

==21st century==
=== 2000s ===

'''<span style="color:#14145C;">2000</span>'''{{anchor|2000}}
* Burnham and others described the new genus and species ''Bambiraptor feinbergorum''.<ref name="norell-makovicky-table-198" />
* Xu, Zhou and Wang described the new genus and species ''Microraptor zhaoianus''.<ref name="norell-makovicky-table-198" />
* Allain and Taquet described the new genus and species ''Pyroraptor olympius''.<ref name="norell-makovicky-table-199" />

[[File:Pyroraptor fossil 01.png|thumb|right|upright|Known skeletal remains of ''[[Biology:Pyroraptor|Pyroraptor]]'']]
[[File:Dans l'ombre des dinosaures - Bambiraptor jeune - 04.jpg|thumb|right|upright|Skeletal reconstruction of ''Bambiraptor'']]
[[File:Microraptor gui holotype.png|thumb|right|upright|Fossil of ''Microraptor'']]
'''<span style="color:#14145C;">2001</span>'''{{anchor|2001}}

* A. R. Jacobsen published a description of a dentary referred to ''[[Biology:Saurornitholestes|Saurornitholestes]]'' with tooth marks.<ref name="saur-bite-abs-58" /> The specimen was preserved in the [[Earth:Dinosaur Park Formation|Dinosaur Park Formation]].<ref name="saur-bite-intro-59" /> Although a specific identification cannot be made, the shape of the preserved serration marks implicate a juvenile individual of one of the formation's tyrannosaurids, like ''[[Biology:Gorgosaurus|Gorgosaurus]]'', ''[[Biology:Daspletosaurus|Daspletosaurus]]'', or ''[[Biology:Aublysodon|Aublysodon]]''.<ref name="saur-bite-discuss-61" /> All of the marks on the jawbone seem to have been left by the same animal because the serration marks all share the same morphology.<ref name="saur-bite-discuss-60" />
* Kevin Padian, Ji Qiang and Ji Shu-an published a review of known feathered dinosaurs and their implications for the origin of flight.<ref name="feather-review-abs-117" /> The authors observed that many aspects of the distribution of feather homologues in the dinosaur family tree met the expectations of earlier phylogenetic hypotheses, including a gradual transition from primitive filaments in ''Sinosauropteryx'' to the shared filaments and "rudimentary" true feathers in ''Caudipteryx'' and ''Protarchaeopteryx'', to flight feathers in ''Archaeopteryx''.<ref name="feather-review-conc-131-132" /> However, they also noted, only filamentous protofeathers had been observed in dromaeosaurs like ''[[Biology:Sinornithosaurus|Sinornithosaurus]]'', when ''Caudipteryx''-style true feathers would be expected due to the family's phylogenetic position.<ref name="feather-review-conc-132" /> Also noted were the lack of known integumentary structures among troodontids.<ref name="feather-review-conc-132" /> The authors proposed that either science's understanding of coelurosaur evolution is faulty or feathers evolved multiple times.<ref name="feather-review-conc-132" />
<blockquote>
{| class="wikitable collapsible collapsed"
|-
! Cladogram of feathered dinosaurs from Padian et al. 2001
|-
|
{{cladogram|title=Feathered Dinosaurs
|caption= Taxa marked with a P were known to bear plumulaceous or pennaceous feathers at the time of the study. Taxa marked with F were known to bear simple [[Biology:Protein filament|filament]]ous integumentary structures.
|clades=
{{clade |style=font-size:100%; width:40em;
|label1= Coelurosaurs
|1={{clade
|label1=
|1=
{{clade
|label1=
|1={{clade
|label1= [[Biology:Maniraptora|Maniraptora]]
|1={{clade
|label1=
|1={{clade
|label1= [[Biology:Aves|Aves]]
|1={{clade
|label1=
|1={{clade
|label1=
|1={{clade
|label1=
|1={{clade
|label1=
|1={{clade
|label1=
|1={{clade
|label1=
|1= Neornithes (P)
|2= ''[[Biology:Ichthyornis|Ichthyornis]]''
}}

|2= ''[[Biology:Hesperornis|Hesperornis]]''
}}

|2= ''[[Biology:Patagopteryx|Patagopteryx]]''
}}

|2= [[Biology:Enantiornithes|Enantiornithes]] (P)
}}

|2= ''[[Biology:Confuciusornis|Confuciusornis]]'' (P)
}}

|2= ''[[Biology:Archaeopteryx|Archaeopteryx]]'' (P)
}}

|2= {{clade
|label1=
|1= Dromaeosaurs (F)
|2= Troodontids
}}

}}

|2={{clade
|label1=
|1={{clade
|label1=
|1={{clade
|label1=
|1= ''[[Biology:Oviraptor|Oviraptor]]''
|2= ''[[Biology:Caudipteryx|Caudipteryx]]'' (F)
}}

|2= Therizinosaurs (F)
}}

|2= ''[[Biology:Protarchaeopteryx|Protarchaeopteryx]]'' (F)
}}

}}

|2= Ornithomimids
}}

|2= ''[[Biology:Tyrannosaurus|Tyrannosaurus]]''
}}
|2={{clade
|label1=
|1= ''[[Biology:Sinosauropteryx|Sinosauropteryx]]'' (F)
|2= ''[[Biology:Compsognathus|Compsognathus]]''
}}
}}

}}
}}
|}
</blockquote>
'''<span style="color:#14145C;">2002</span>'''{{anchor|2002}}
* Hwang and others performed a cladistic analysis of the Dromaeosauridae.<ref name="norell-makovicky-systevo-207" />
* Xu and others performed a cladistic analysis of the Dromaeosauridae.<ref name="norell-makovicky-systevo-207" />
[[File:Dromaeosauroides.jpg|thumb|right|upright|Artist's restoration of ''Dromaeosauroides'']]

* Czerkas and others described the new genus and species ''Cryptovolans pauli''.<ref name="czerkas-etal-abs-96" />

'''<span style="color:#14145C;">2003</span>'''{{anchor|2003}}
* Christiansen and Bonde described the new genus and species ''Dromaeosauroides bornholmensis''.<ref name="christiansen-bonde-abs-287" />

'''<span style="color:#14145C;">2004</span>'''{{anchor|2004}}
* Currie and Varricchio described the new genus and species ''Atrociraptor marshalli''.<ref name="currie-varricchio-abs-112" />
[[File:BuitreraptorROM.JPG|thumb|right|upright|Skeletal mount of ''Buitreraptor'']]
* Xu and Wang described the new genus and species ''Graciliraptor lujiatunensis''.<ref name="xu-wang-abs-11" />

'''<span style="color:#14145C;">2005</span>'''{{anchor|2005}}
* Makovicky, Apesteguía and Agnolin described the new genus and species ''Buitreraptor gonzalezorum''.<ref name="makovicky-apestaguia-agnolin-abs-1007" />
* Novas and Pol described the new genus and species ''Neuquenraptor argentinus''.<ref name="novas-pol-abs-858" />

'''<span style="color:#14145C;">2006</span>'''{{anchor|2006}}
* Norell and others described the new genus and species ''Tsaagan mangas''.<ref name="norell-etal-abs-1" />

'''<span style="color:#14145C;">2007</span>'''{{anchor|2007}}
* Lü and others described the new genus and species ''Luanchuanraptor henanensis''.<ref name="lu-etal-abs-777" />
[[File:Austroraptor BW.jpg|thumb|right|upright|Artist's restoration of ''Austroraptor'']]
* Turner and others described the new genus and species ''Mahakala omnogovae''.<ref name="turner-etal-abs-1378" />
* A. S. Turner, S. H. Hwang, and M. A. Norell described the new genus and species ''Shanag ashile''.<ref name="turner-hwang-norell-abs-1" />

'''<span style="color:#14145C;">2008</span>'''{{anchor|2008}}
* Novas et al. described the new genus and species ''Austroraptor cabazai''.<ref name="novas-etal-abs-1101" />

'''<span style="color:#14145C;">2009</span>'''{{anchor|2009}}
* Longrich and Currie described the new genus and species ''Hesperonychus elizabethae''.<ref name="longrich-currie-abs-5002" />

===2010s===
'''<span style="color:#14145C;">2010</span>'''{{anchor|2010}}
* Xu and others described the new genus and species ''Linheraptor exquisitus''.<ref name="xu-etal-abs-1" />
* Zheng and others described the new genus and species ''Tianyuraptor ostromi''.<ref name="zheng-etal-abs-211" />

'''<span style="color:#14145C;">2011</span>'''{{anchor|2011}}
* Porfiri, Calvo and Santos described the new genus and species ''Pamparaptor micros''.<ref name="porfiri-calvo-santos-abs-109" />

'''<span style="color:#14145C;">2012</span>'''{{anchor|2012}}
* Gong and others described the new species ''Microraptor hanqingi''.<ref name="gong-etal-abs-1" />
[[File:Changyuraptor.jpg|thumb|right|upright|Artist's restoration of ''Changyuraptor'']]
* Senter and others described the new genus and species ''Yurgovuchia doellingi''.<ref name="senter-etal-abs-58" />

'''<span style="color:#14145C;">2013</span>'''{{anchor|2013}}
* D. C. Evans, D. W. Larson, and P. J. Currie described the new genus and species ''Acheroraptor temertyorum''.<ref name="evans-larson-currie-abs-1041" />

'''<span style="color:#14145C;">2014</span>'''{{anchor|2014}}
* Han and others described the new genus and species ''Changyuraptor yangi''.<ref name="han-etal-abs" />

'''<span style="color:#14145C;">2015</span>'''{{anchor|2015}}
* Jasinski described the new species ''Saurornitholestes sullivani''.<ref name="jasinski-abs" />
* Lü and Brusatte described the new genus and species ''Zhenyuanlong suni''.<ref name="lu-brusatte-abs" />
* DePalma et al. described the new genus and species ''Dakotaraptor steini''.<ref name="depalma-etal-2015-inpassim" />

<blockquote>
{| class="wikitable collapsible collapsed"
|-
! Cladogram of dromaeosaurids from Lü and Brusatte 2015
|-
|{{clade| style=font-size:85%;line-height:85%
|label1=[[Biology:Dromaeosauridae|Dromaeosauridae]]
|1={{clade
   |1=''Mahakala''
   |2={{clade
      |1=''[[Biology:Unenlagia|Unenlagia]]''
      |2=''[[Biology:Austroraptor|Austroraptor]]''
      |3=''[[Biology:Buitreraptor|Buitreraptor]]''
      |4=''[[Biology:Rahonavis|Rahonavis]]''
      |5={{clade
         |1=''[[Biology:Graciliraptor|Graciliraptor]]''
         |2=''[[Biology:Changyuraptor|Changyuraptor]]''
         |3=''[[Biology:Microraptor|Microraptor]]''
         |4=''[[Biology:Sinornithosaurus|Sinornithosaurus]]''
         |5=''[[Biology:Tianyuraptor|Tianyuraptor]]''
         |6='''''Zhenyuanlong'''''
         |7={{clade
            |label1=Eudromaeosauria    
            |1={{clade
               |1=''[[Biology:Bambiraptor|Bambiraptor]]''
               |2=''[[Biology:Saurornitholestes|Saurornitholestes]]''
               |3=''Tsaagan''
               |4=''[[Biology:Adasaurus|Adasaurus]]''
               |5=''Deinonychus''
               |6={{clade
                  |1=''[[Biology:Balaur bondoc|Balaur]]''
                  |2=''Velociraptor''
               |label7=Dromaeosaurinae
               |7={{clade
                  |1=''Achillobator''
                  |2=''[[Biology:Utahraptor|Utahraptor]]''
                  |3=''Atrociraptor''
                  |4=''[[Biology:Dromaeosaurus|Dromaeosaurus]]'' }} }} }} }} }} }} }} }}
|}
</blockquote>
* Bell and Currie described the new genus and species ''Boreonykus certekorum''.<ref name="bell-currie-inpassim" />

'''<span style="color:#14145C;">2017</span>'''{{anchor|2017}}
* Xu and others described the new genus and species ''Zhongjianosaurus yangi''.
* Cau and others described the new genus and species ''Halszkaraptor escuilliei''.
'''<span style="color:#14145C;">2019</span>'''{{anchor|2019}}

* An [[Biology:Ungual|ungual]] [[Biology:Phalanx bone|phalanx]] of a [[Biology:Dromaeosauridae|dromaeosaurid]] theropod is described from the Blagoveshchensk area ({{wipe|Russia}}) by Bolotskii, Bolotskii & Sorokin (2019).<ref>{{Cite journal|author1=I. Yu. Bolotskii |author2=Yu. L. Bolotskii |author3=A. P. Sorokin |year=2019 |title=The first find of an ungual phalanx of a dromaeosaurid dinosaur (Dinosauria: Dromaeosauridae) from the Blagoveshchensk area of Late Cretaceous dinosaurs (Amur Region, Russia) |journal=Doklady Earth Sciences |volume=484 |issue=1 |pages=18–20 |doi=10.1134/S1028334X19010100 |bibcode=2019DokES.484...18B |s2cid=134803475 }}</ref>

==See also==
* History of paleontology
* [[Earth:Timeline of paleontology|Timeline of paleontology]]

==Footnotes==
{{Reflist|30em|refs=
<!--
<ref name="makovicky-norell-intro-184">"Introduction", Makovicky and Norell (2004); page 184.</ref>
<ref name="makovicky-norell-table-185">"Table 9.1: Troodontidae", Makovicky and Norell (2004); page 185.</ref>
<ref name="makovicky-norell-table-186">"Table 9.1: Troodontidae", Makovicky and Norell (2004); page 186.</ref>
<ref name="makovicky-norell-systevo-193">"Systematics and Evolution", Makovicky and Norell (2004); page 193.</ref>
<ref name="makovicky-norell-systevo-193-194">"Systematics and Evolution", Makovicky and Norell (2004); pages 193–194.</ref>
<ref name="makovicky-norell-systevo-194">"Systematics and Evolution", Makovicky and Norell (2004); page 194.</ref>
<ref name="makovicky-norell-paleobio-194">"Paleobiology", Makovicky and Norell (2004); page 194.</ref>
<ref name="makovicky-norell-biogeo-194">"Biogeography", Makovicky and Norell (2004); page 194.</ref>
<ref name="makovicky-norell-paleobio-195">"Paleobiology", Makovicky and Norell (2004); page 195.</ref> -->

<ref name="norell-makovicky-intro-196">Norell and Makovicky (2004); "Introduction", page 196.</ref>
<ref name="norell-makovicky-table-198">Norell and Makovicky (2004); "Table 10.1: Dromaeosauridae", page 198.</ref>
<ref name="norell-makovicky-table-199">Norell and Makovicky (2004); "Table 10.1: Dromaeosauridae", page 199.</ref>
<ref name="norell-makovicky-systevo-206">Norell and Makovicky (2004); "Systematics and Evolution", page 206.</ref>
<ref name="norell-makovicky-systevo-207">Norell and Makovicky (2004); "Systematics and Evolution", page 207.</ref>
<ref name="norell-makovicky-paleoeco-209">Norell and Makovicky (2004); "Paleoecology", page 209.</ref>

<ref name="saur-bite-abs-58">Jacobsen (2001); "Abstract," page 58.</ref>
<ref name="saur-bite-intro-59">Jacobsen (2001); "Introduction," page 59.</ref>
<ref name="saur-bite-discuss-61">Jacobsen (2001); "Discussion," page 61.</ref>
<ref name="saur-bite-discuss-60">Jacobsen (2001); "Discussion," page 60.</ref>

<ref name="feather-review-abs-117">Padian, Ji, and Ji (2001); "Abstract," page 117.</ref>
<ref name="feather-review-conc-131-132">Padian, Ji, and Ji (2001); "Conclusions," pages 131–132.</ref>
<ref name="feather-review-conc-132">Padian, Ji, and Ji (2001); "Conclusions," page 132.</ref>

<ref name="mayor-crow-272-273">Mayor (2005); "Crow Fossil Collections," pages 272–273.</ref>

<ref name="bigsky-history-53-54">Horner (2001); "History of Dinosaur Collecting in Montana," pages 53–54.</ref>

<ref name="ostrom-aff-147-148">Ostrom (1969); "6. Affinities of ''Deinonychus''," pages 147–148.</ref>

<ref name="xu-etal-inpassim-335-340">Xu et al. (2003); ''in passim'' pages 335–340.</ref>

<ref name="czerkas-etal-abs-96">Czerkas et al. (2002); "Abstract," page 96.</ref>

<ref name="christiansen-bonde-abs-287">Christiansen and Bonde (2003); "Abstract," page 287.</ref>

<ref name="currie-varricchio-abs-112">Currie and Varricchio (2004); "Abstract," page 112.</ref>
<ref name="xu-wang-abs-11">Xu and Wang (2004); "Abstract," page 11.</ref>

<ref name="makovicky-apestaguia-agnolin-abs-1007">Makovicky, Apesteguia and Agnolin (2005); "Abstract," page 1007.</ref>
<ref name="novas-pol-abs-858">Novas and Pol (2005); "Abstract," page 858.</ref>

<ref name="norell-etal-abs-1">Norell et al. (2006); "Abstract," page 1.</ref>

<ref name="lu-etal-abs-777">Lü et al. (2007); "Abstract," page 777.</ref>
<ref name="turner-etal-abs-1378">Turner et al. (2007); "Abstract," page 1378.</ref>
<ref name="turner-hwang-norell-abs-1">Turner, Hwang, and Norell (2007); "Abstract," page 1.</ref>

<ref name="novas-etal-abs-1101">Novas et al. (2008); "Abstract," page 1101.</ref>

<ref name="longrich-currie-abs-5002">Longrich and Currie (2009); "Abstract," page 5002.</ref>

<!-- <ref name="csiki-etal-abs-15357">Csiki et al. (2010); "Abstract," page 15357.</ref> -->
<ref name="xu-etal-abs-1">Xu et al. (2010); "Abstract," page 1.</ref>
<ref name="zheng-etal-abs-211">Zheng et al. (2010); "Abstract," page 211.</ref>

<ref name="porfiri-calvo-santos-abs-109">Porfiri, Calvo and Santos (2011); "Abstract," page 109.</ref>

<ref name="gong-etal-abs-1">Gong et al. (2012); "Abstract," page 1.</ref>
<ref name="senter-etal-abs-58">Senter et al. (2012); "Abstract," page 58.</ref>

<ref name="evans-larson-currie-abs-1041">Evans, Larson, and Currie (2013); "Abstract," page 1041.</ref>

<ref name="han-etal-abs">Han et al. (2014); "Abstract".</ref>

<ref name="jasinski-abs">Jasinski (2015); "Abstract", page 79.</ref>

<ref name="lu-brusatte-abs">Lü and Brusatte (2015); "Abstract", page 1.</ref>

<ref name="depalma-etal-2015-inpassim">DePalma et al. (2015); ''in passim''.</ref>

<ref name="bell-currie-inpassim">Bell and Currie (2015); ''in passim''.</ref>

}}

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==External links==


[[Category:Paleontology timelines|Dromaeosaurid]]

{{Sourceattribution|Timeline of dromaeosaurid research}}