Biology:List of Scarabaeidae subfamilies and tribes

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The beetle family Scarabaeidae is made up of about 28 subfamilies containing more than 100 tribes. There are more than 2,500 genera and 35,000 described species in Scarabaeidae.[1]

The following subfamilies and tribes are in accordance with those in Catalog of Life (2023) and Dietz, Ahrens, et al. (2023). The tribe Hopliini is placed as a subfamily in Catalogue of Life, but is listed as a tribe of subfamily Melolonthinae here. The subfamilies Sericinae and Sericoidinae were suggested in Dietz, Ahrens, et al. (2023) and are included in the Catalogue of Life.[1][2] The subfamily Lichniinae is sometimes considered a tribe, Lichniini.[3]

Family Scarabaeidae Latreille, 1802
Subfamily Aclopinae Blanchard, 1850
Aclopini Blanchard, 1850
Holcorobeini Nikolajev, 1992
Phaenognathini Iablokoff-Khnozorian, 1977
Subfamily Aegialiinae Castelnau, 1840
Aegialiini Castelnau, 1840
Saprinianini Nikolajev, 2011
Subfamily Allidiostomatinae Arrow, 1940
Subfamily Aphodiinae Leach, 1815
Aphodiini Leach, 1815
Corythoderini Schmidt, 1910
Eupariini Schmidt, 1910
Hornietiini Minkina, 2020
Odochilini Rakovič, 1987
Odontolochini Stebnicka & Howden, 1996
Psammodiini Mulsant, 1842
Rhyparini Schmidt, 1910
Stereomerini Howden & Storey, 1992
Termitoderini Tangelder & Krikken, 1982
Thinorycterini Semenov & Reichardt, 1925
†Psammaegialiini Nikolajev, Wang & Zhang, 2014
Subfamily Aulonocneminae Janssens, 1946
Subfamily Cetoniinae Leach, 1815
Cetoniini Leach, 1815
Cremastocheilini Burmeister & Schaum, 1841
Diplognathini Burmeister, 1842
Goliathini Latreille, 1829
Gymnetini Kirby, 1827
Phaedimini Schoch, 1894
Schizorhinini Burmeister, 1842
Stenotarsiini Kraatz, 1880
Taenioderini Mikšić, 1976
Xiphoscelidini Krikken, 1984
Subfamily Chironinae Blanchard, 1845
Subfamily Dynamopodinae Arrow, 1911
Subfamily Dynastinae MacLeay, 1819
Agaocephalini Burmeister, 1847
Cyclocephalini Castelnau, 1840
Dynastini Macleay, 1819
Hexodontini Lacordaire, 1855
Oryctini Mulsant, 1842
Oryctoderini Endrödi, 1966
Pentodontini Mulsant, 1842
Phileurini Burmeister, 1847
Subfamily Eremazinae Stebnicka, 1977
Eremazini Stebnicka, 1977
†Yixianscarabaeini Nikolajev, 2015
Subfamily Lichniinae Burmeister, 1844
Subfamily Melolonthinae MacLeay, 1819
Acomini Evans & Smith, 2020
Ardellini Paulsen, 2021
Athliini Smith & Evans, 2018
Chasmatopterini Lacordaire, 1855
Chnaunanthini Evans & Smith, 2020
Colymbomorphini Blanchard, 1850
Comophini Britton, 1978
Dichelonychini Kirby, 1837
Diphycerini Medvedev, 1952
Diplotaxini Kirby, 1837
Euchirini Hope, 1840
Hopliini Latreille, 1829
Langbianellini Prokofiev, 2015
Macrodactylini Kirby, 1837
Melolonthini Leach, 1819
Pachypodini Erichson, 1840
Pachytrichini Burmeister, 1855
Phobetusini Evans & Smith, 2020
Phyllotocidiini Britton, 1957
Rhizotrogini Burmeister, 1855
Systellopini Dalla Torre, 1912
Warwickiini Evans & Smith, 2020
†Cretomelolonthini Nikolajev, 1998
Subfamily Oncerinae LeConte, 1861
Subfamily Orphninae Erichson, 1847
Aegidiini Paulian, 1984
Orphnini Erichson, 1847
Subfamily Pachydeminae Reitter, 1902
Aegosthetini Lacroix, 2007
Pachydemini Burmeister, 1855
Subfamily Phaenomeridinae Erichson, 1847
Subfamily Podolasiinae Howden, 1997
Subfamily Rutelinae MacLeay, 1819
Adoretini Burmeister, 1844
Alvarengiini Frey, 1975
Anatistini Lacordaire, 1855
Anomalini Streubel, 1839
Anoplognathini MacLeay, 1819
Geniatini Burmeister, 1844
Rutelini MacLeay, 1819
Subfamily Scarabaeinae Latreille, 1802
Ateuchini Perty, 1830
Byrrhidiini Davis, Deschodt & Scholtz, 2019
Coprini Leach, 1815
Deltochilini Lacordaire, 1855
Dichotomiini Pereira, 1954
Endroedyolini Davis, Deschodt & Scholtz, 2019
Epactoidini Rossini et al, 2022
Epilissini Lansberge, 1874
Epirinini Lansberge, 1874
Eucraniini Burmeister, 1873
Gymnopleurini Streubel, 1846
Odontolomini Davis, Deschodt & Scholtz, 2019
Oniticellini Kolbe, 1905
Onitini Castelnau, 1840
Onthophagini Streubel, 1846
Parachoriini Tarasov, 2017
Phanaeini Hope, 1838
Scarabaeini Latreille, 1802
Sisyphini Mulsant, 1842
Subfamily Sericinae Kirby, 1837
Ablaberini Blanchard, 1850
Diphucephalini Castelnau, 1840
Sericini Kirby, 1837
Subfamily Sericoidinae Erichson, 1847
Automoliini Britton, 1978
Heteronychini Lacordaire, 1855
Liparetrini Burmeister, 1855
Maechidiini Burmeister, 1855
Phyllotocini Burmeister, 1855
Scitalini Britton, 1957
Sericoidini Erichson, 1847
Subfamily Termitotroginae Wasmann, 1918
Subfamily Trichiinae Kolbe, 1897
Trichiini Fleming, 1821
Subfamily Valginae Schenkling, 1922
Valgini Mulsant, 1842
Subfamily †Cretoscarabaeinae Nikolajev, 1995
Subfamily †Lithoscarabaeinae Nikolajev, 1992
Subfamily †Prototroginae Nikolajev, 2000


References

  1. 1.0 1.1 "Scarabaeidae Latreille, 1802". https://www.catalogueoflife.org/data/taxon/6278C. 
  2. Dietz, Lars; Seidel, Matthias; Eberle, Jonas; Misof, Bernhard et al. (2023). "A transcriptome-based phylogeny of Scarabaeoidea confirms the sister group relationship of dung beetles and phytophagous pleurostict scarabs (Coleoptera)". Systematic Entomology 48 (4). doi:10.1111/syen.12602. 
  3. Mondaca, José (2023). "A checklist of the Scarabaeoidea (Coleoptera) of Chile with exemplar live-photographed". Zootaxa 5285 (2). doi:10.11646/ZOOTAXA.5285.2.1. https://www.mapress.com/zt/article/view/zootaxa.5285.2.1.