Biology:Charitomenosuchus

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Charitomenosuchus (meaning "graceful crocodile") is an extinct genus of machimosaurid teleosauroid from the Callovian Oxford Clay Formation of England.[1]

Illustration of the holotype

History

The type species was originally named "Steneosaurus" leedsi by Andrews in 1909 on the basis of three specimens.[2] The genus Steneosaurus was used as a longtime wastebasket taxon for various teleosauroid specimens and had more than a dozen species. The type species, Steneosaurus rostromajor is undiagnostic, making the genus and species dubious and invalid. Additionally, many species of so called "Steneosaurus" were found to be quite different and unrelated to one another, thus needing new generic names.[1].

In her unpublished 2019 thesis, Michela Johnson coined the nomen ex dissertationae Charitomenosuchus for S. leedsi.[3] The new genus and combination, Charitomenosuchus leedsi was formally published by Johnson and colleagues in 2020.[1]

Classification

Charitomenosuchus is classified as one of the most basal machimosaurids, being outside the subfamily Machimosaurinae. It's more gracile and lightly-built morphology is typical of basal machimosaurids while more derived members were large and much more robust.

The cladogram below is from an analysis by Johnson and colleagues in 2020.[1]

Machimosauridae

Macrospondylus bollensis

Clovesuurdameredeor stephani

Charitomenosuchus leedsi

Seldsienean megistorhynchus

Deslongchampsina larteti

Machimosaurinae

Proexochokefalos hebertii

cf. Proexochokefalos bouchardi

Neosteneosaurus edwardsi

Andrianavoay baroni

Machimosaurini

Lemmysuchus obtusidens

Yvridiosuchus boutilieri

Machimosaurus buffetauti

Machimosaurus mosae

Machimosaurus rex

Machimosaurus hugii

Ecology

In life, Charitomenosuchus was a marine hunter of small prey. It had proportionally one of the most gracile jaws of all machimosaurids, with a quick bite but relatively weak bite force. Additionally, it had a relatively weak front bite, meaning it could grab prey quickly but likely took longer to overpower and process. This suggests that Charitomenosuchus was adapted for catching quick prey such as small fish and cephalopods that it coexisted with.[4]

References

  1. 1.0 1.1 1.2 1.3 Johnson, Michela M.; Young, Mark T.; Brusatte, Stephen L. (2020). "The phylogenetics of Teleosauroidea (Crocodylomorpha, Thalattosuchia) and implications for their ecology and evolution". PeerJ 8. doi:10.7717/peerj.9808. PMID 33083104. 
  2. C. W. Andrews. 1909. On some new steneosaurs from the Oxford Clay of Peterborough. Annals and Magazine of Natural History 3:299-308
  3. Johnson, Michela Maria Angeline (2019) (in English). The taxonomy, systematics and ecomorphological diversity of Teleosauroidea (Crocodylomorpha, Thalattosuchia), and the evaluation of the genus 'Steneosaurus'. https://era.ed.ac.uk/handle/1842/36656. 
  4. Johnson, M. M.; Foffa, D.; Young, M. T.; Brusatte, S. L. (2022). "The ecological diversification and evolution of Teleosauroidea (Crocodylomorpha, Thalattosuchia), with insights into their mandibular biomechanics". Ecology and Evolution 12 (11). doi:10.1002/ece3.9484. PMID 36415878. Bibcode2022EcoEv..12E9484J. 

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