Biology:Homoeosaurus

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Homoeosaurus is an extinct genus of rhynchocephalian reptile, known from the Late Jurassic-earliest Cretaceous of Europe, with specimens being reported from France (Canjuers Lagerstatte), England (Purbeck Group) and Germany (Solnhofen Limestone). Several species have been described within the genus, based on varying proportions of the limb bones to the body length based on the presacral vertebrae. Specimen C.M.6438 of H. maximiliani from Germany has a total length of around 17 centimetres (6.7 in), with a skull length of about 1.7 centimetres (0.67 in). In comparison to other rhynchocephalians, the limbs are proportionally long.[1] Recent studies have classified Homoeosaurus as a member of Neosphenodontia, with some studies including it as part of the clade Leptorhynchia, also including sapheosaurs, pleurosaurs, Kallimodon and Vadasaurus.[2] Despite being found in aquatic deposits, it is suggested to have been terrestrial.[3] Analysis of its limb bone morphology suggests that it was probably capable of climbing and may have climbed trees and/or rocky surfaces at least some of the time.[4] It is thought to have been a carnivore/insectivore.[5] One specimen was found as stomach contents of the fish Belonostomus.[6]

Cladogram after Beccari et al. 2025:[4]

Rhynchocephalia

Gephyrosaurus

Sphenodontia

Diphydontosaurus

Planocephalosaurus

Eusphenodontia

Clevosaurus (paraphyletic)

Neosphenodontia
Sphenodontidae
†Eilenodontinae

Priosphenodon

Sphenotitan

Sphenodontinae

Opisthiamimus

Navajosphenodon

Sphenofontis

Sphenodon (tuatara)

Homoeosaurus solnhofensis

Homoeosaurus maximiliani

Homoeosaurus parvipes

Sphenodraco

Kallimodon

Pleurosauridae

Derasmosaurus

Ankylosphenodon

Palaeopleurosaurus

Arcosaurus

Pleurosaurus

Vadasaurus

Sapheosauridae

Sapheosaurus

Piocormus

Oenosaurus

References

  1. Wu X-C. The comparative anatomy and systematics of Mesozoic sphenodontians. Ph.D. Thesis, McGill University. 1991. p. 84-113.
  2. DeMar, David G.; Jones, Marc E. H.; Carrano, Matthew T. (2022-12-31). "A nearly complete skeleton of a new eusphenodontian from the Upper Jurassic Morrison Formation, Wyoming, USA, provides insight into the evolution and diversity of Rhynchocephalia (Reptilia: Lepidosauria)" (in en). Journal of Systematic Palaeontology 20 (1): 1–64. doi:10.1080/14772019.2022.2093139. ISSN 1477-2019. https://www.tandfonline.com/doi/full/10.1080/14772019.2022.2093139. 
  3. Reynoso, Víctor-Hugo (2005-09-30). "Possible evidence of a venom apparatus in a Middle Jurassic Sphenodontian from the Huizachal red beds of Tamaulipas, México" (in en). Journal of Vertebrate Paleontology 25 (3): 646–654. doi:10.1671/0272-4634(2005)025[0646:PEOAVA2.0.CO;2]. ISSN 0272-4634. http://www.tandfonline.com/doi/abs/10.1671/0272-4634%282005%29025%5B0646%3APEOAVA%5D2.0.CO%3B2. 
  4. 4.0 4.1 Beccari, Victor; Guillaume, Alexandre R. D.; Jones, Marc E. H.; Villa, Andrea; Cooper, Natalie; Regnault, Sophie; Rauhut, Oliver W. M. (2025-07-01). "An arboreal rhynchocephalian from the Late Jurassic of Germany, and the importance of the appendicular skeleton for ecomorphology in lepidosaurs". Zoological Journal of the Linnean Society 204 (3). doi:10.1093/zoolinnean/zlaf073. ISSN 0024-4082. 
  5. Rauhut, Oliver W. M.; Heyng, Alexander M.; López-Arbarello, Adriana; Hecker, Andreas (2012-10-31). Farke, Andrew A.. ed. "A New Rhynchocephalian from the Late Jurassic of Germany with a Dentition That Is Unique amongst Tetrapods" (in en). PLOS ONE 7 (10). doi:10.1371/journal.pone.0046839. ISSN 1932-6203. PMID 23118861. Bibcode2012PLoSO...746839R. 
  6. Frey, Eberhard; Tischlinger, Helmut (2012-03-07). Fenton, Brock. ed. "The Late Jurassic Pterosaur Rhamphorhynchus, a Frequent Victim of the Ganoid Fish Aspidorhynchus?" (in en). PLOS ONE 7 (3). doi:10.1371/journal.pone.0031945. ISSN 1932-6203. PMID 22412850. Bibcode2012PLoSO...731945F. 

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