Biology:List of eurypterid genera

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Eurypterus, the most commonly found eurypterid fossil and the first eurypterid genus to be described

This list of eurypterid genera is a comprehensive listing of all genera that have ever been included in the order Eurypterida, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful (nomen dubium), or were not formally published (nomen nudum), as well as junior synonyms of more established names and genera that are no longer considered eurypterids. The list currently includes 115 names out of which 74 are considered valid eurypterid genera.

There are approximately 250 species of eurypterids recognized as valid.[1]

Naming conventions and terminology

There is no "official" or "canonical" list of eurypterid genera. The closest thing is found contained in the regularly updated Summary list of fossil spiders and their relatives in the World Spider Catalog. The vast majority of the content of the list below, including the valid genera, preoccupied names, junior synonyms, taxonomical classifications and sites of discovery, is based upon the 2018 edition of the Summary list. Content not based on the list, such as genera described after its publication, is noted through inline citations.

Naming conventions and terminology follow the International Code of Zoological Nomenclature. Technical terms used include:

  • Junior synonym: A name which describes the same taxon as a previously published name. If two or more genera are formally designated and the type specimens are later assigned to the same genus, the first to be published (in chronological order) is the senior synonym, and all other instances are junior synonyms. Senior synonyms are generally used, except by special decision of the ICZN, but junior synonyms cannot be used again, even if deprecated. Junior synonymy is often subjective, unless the genera described were both based on the same type specimen.
  • Nomen nudum (Latin for "naked name"): A name that has appeared in print but has not yet been formally published by the standards of the ICZN. Nomina nuda (the plural form) are invalid, and are therefore not italicized as a proper generic name would be. If the name is later formally published, that name is no longer a nomen nudum and will be italicized on this list. Often, the formally published name will differ from any nomina nuda that describe the same specimen.
  • Nomen manuscriptum (Latin for "manuscript name"): A name that appears in manuscript of a formal, but not-peer-reviewed, publication that has no scientific backing. A nomen manuscriptum is equivalent to a nomen nudum for everything except the method of publication, and description.
  • Nomen oblitum (Latin for "forgotten name"): A name that has not been used in the scientific community for more than fifty years after its original proposal.
  • Preoccupied name: A name that is formally published, but which has already been used for another taxon. This second use is invalid (as are all subsequent uses) and the name must be replaced. As preoccupied names are not valid generic names, they will also go unitalicized on this list.
  • Nomen dubium (Latin for "dubious name"): A name describing a fossil with no unique diagnostic features.

Eurypterid genera

  Valid genus
  Dubious taxon, synonym, preoccupied name or informal name
  Missassigned genus
Genus Author(s) Year Status Age Location Notes

Acanthoeurypterus

Kjellesvig-Waering

N/A

Jr. synonym

N/A

N/A

Nomen manuscriptum, also considered as a synonym of Eurypterus.[2]

Acutiramus Ruedemann 1935 Valid Silurian - Devonian
  •  Australia[3]
  •  United States
  •  Czech Republic
  •  Canada
Adelophthalmus Jordan 1854 Valid Devonian - Permian Nearly cosmopolitan, see article.
Alkenopterus Størmer 1974 Valid Devonian
  •  Germany

Alloeurypterus

Kjellesvig-Waering

N/A

Jr. synonym

N/A

N/A

Nomen manuscriptum, also considered as a synonym of Eurypterus.[2]

Angustidontus

Cooper

1936

Misidentification[4]

N/A

N/A

Misidentified, a crustacean.

Anthraconectes

Meek & Worthen

1868

Jr. synonym

N/A

N/A

Synonym of Adelophthalmus.

Baltoeurypterus

Størmer

1973

Jr. synonym

N/A

N/A

Synonym of Eurypterus.

Bassipterus Kjellesvig-Waering & Leutze 1966 Valid Silurian
  •  United States

Belinuropsis

Matthew

1910

Nomen dubium

Carboniferous

  •  Canada

Too poorly preserved to be assigned to any particular arthropod group.[5]

Bembicosoma

Laurie

1899

Misidentification[6]

N/A

N/A

Misidentified, a stem genus on Planaterga.

Borchgrevinkium

Novojilov

1959

Misidentification[1]

N/A

N/A

Misidentified, possibly a stem prosomapod.

Brachyopterella Kjellesvig-Waering 1966 Valid Silurian
  •  Norway
  •  Scotland
Brachyopterus Størmer 1951 Valid Ordovician
  •  Wales

Bunodella

Matthew

1889

Nomen dubium

Silurian

  •  Canada

Uncertain affinities, probably synonym of Acutiramus.[7]

Buffalopterus Kjellesvig-Waering & Heubusch 1962 Valid Silurian
  •  United States
Campylocephalus Eichwald 1860 Valid Carboniferous - Permian
  •  Russia
  •  Czech Republic

Campylognathus

Diener

1924

Preoccupied[8]

N/A

N/A

Preoccupied by a genus of plant bugs, changed to Campylocephalus.

Carcinosoma Claypole 1890 Valid Silurian
  •  England
  •  Canada
  •  United States
  •  Scotland
Ciurcopterus Tetlie & Briggs 2009 Valid Silurian
  •  United States
Clarkeipterus Kjellesvig-Waering 1966 Valid Silurian
  •  United States

Clintonipterus

Kjellesvig-Waering

N/A

Nomen nudum

N/A

N/A

Informal name that only appears in notes in museum drawers.[9]

Ctenopterus Clarke & Ruedemann 1912 Valid Silurian
  •  United States

Curviramus

Ruedemann

1935

Jr. synonym

N/A

N/A

Synonym of Pterygotus.

Cyrtoctenus

Størmer & Waterston

1968

Jr. synonym

N/A

N/A

Synonym of Hibbertopterus.[10]

Dolichocephala

Claypole

1883

Preoccupied

N/A

N/A

Preoccupied by a genus of flies, synonym of Hallipterus.

Dolichopterus Hall 1859 Valid Silurian
  •  United States
  •  Sweden
Dorfopterus Kjellesvig-Waering 1955 Valid Devonian
  •  United States
Drepanopterus Laurie 1892 Valid Silurian - Devonian
  •  Scotland
  •  England
  •  Canada

Dunsopterus

Waterston

1968

Jr. synonym

N/A

N/A

Synonym of Hibbertopterus.[10]

Echinognathus Walcott 1882 Valid Ordovician
  •  United States
Eocarcinosoma Caster & Kjellesvig-Waering 1964 Valid Ordovician
  •  United States
Erettopterus Salter 1859 Valid Silurian - Devonian
  •  Canada
  •  England
  •  Estonia
  •  Norway
  •  Scotland
  •  Sweden
  •  United States
Erieopterus Kjellesvig-Waering 1958 Valid Silurian - Devonian
  •  United States
  •  Canada
  •  Norway
  •  Germany

Eidothea

Scouler

1831

Preoccupied[11]

N/A

N/A

Preoccupied name, changed to Campylocephalus.

Eurypterella

Matthew

1888

Nomen dubium

Carboniferous

  •  Canada

Too poorly preserved to be assigned to any particular arthropod group.[12]

Eurypterus de Kay 1825 Valid Silurian
  •  Norway
  •  England
  •  United States
  •  Sweden
  •  Estonia

Eurysoma

Claypole

1890

Preoccupied

N/A

N/A

Preoccupied name, changed to Carcinosoma.

Eusarcana Strand 1942 Valid Silurian - Devonian
  •  Scotland
  •  Czech Republic
  •  United States

Eusarcus

Grote & Pitt

1875

Preoccupied

N/A

N/A

Preoccupied name, changed to Eusarcana.

Eysyslopterus Tetlie & Poschmann 2008 Valid Silurian
  •  Estonia

Glaucodes

Pruvost

1923

Jr. synonym[13]

N/A

N/A

Synonym of Mycterops.

Glyptoscorpius

Peach

1882

Jr. synonym

N/A

N/A

Synonym of Adelophthalmus.

Grossopterus Størmer 1934 Valid Devonian
  •  Germany
  •  Israel
Hallipterus Kjellesvig-Waering 1963 Valid Devonian
  •  United States
Hardieopterus Waterston 1979 Valid Silurian
  •  Scotland
  •  England
  •  United States
Hastimima White 1908 Valid Permian
  •  Brazil
  •  South Africa
  •  United States
Herefordopterus Tetlie 2006 Valid Silurian
  •  England
Hibbertopterus Kjellesvig-Waering 1959 Valid Carboniferous
  •  Belgium
  •  Czech Republic
  •  Ireland
  •  Scotland
  •  South Africa

Himantopterus

Salter

1856

Preoccupied[14]

N/A

N/A

Unavailable junior homonym, replaced by name Erettopterus

Holmipterus Kjellesvig-Waering 1979 Valid Silurian
  •  Sweden
Hughmilleria Sarle 1903 Valid Silurian
  •  United States
  •  China
Jaekelopterus Waterston 1964 Valid Devonian
  •  United States
  •  Germany
Kiaeropterus Waterston 1979 Valid Silurian
  •  Norway
  •  Scotland

Kockurus

Chlupáč

1995

Misidentification

N/A

N/A

Misidentified, likely represents an aglaspidid.

Kodymirus

Chlupáč

1995

Misidentification

N/A

N/A

Misidentified, likely represents an aglaspidid.

Kokomopterus Kjellesvig-Waering 1966 Valid Silurian
  •  United States
Lanarkopterus Ritchie 1968 Valid Silurian
  •  Scotland
Lamontopterus Waterston 1979 Valid Silurian
  •  Scotland
Laurieipterus Kjellesvig-Waering 1966 Valid Silurian
  •  Scotland
Leiopterella Lamsdell, Braddy, Loeffler & Dineley 2010 Valid Devonian
  •  Canada

Lepidoderma

Reuss

1855

Jr. synonym

N/A

N/A

Synonym of Adelophthalmus.

Marsupipterus Caster & Kjellesvig-Waering 1955 Valid Silurian
  •  England

Mazonipterus

Kjellesvig-Waering

1963

Misidentification

N/A

N/A

Misidentified, a plant.

Megalograptus Miller 1874 Valid Ordovician
  •  United States
Megarachne Hünicken 1980 Valid Carboniferous - Permian
  •  Argentina

Melbournopterus

Caster & Kjellesvig-Waering

1953

Misidentification

N/A

N/A

Misidentified, a brachipod.

Mixopterus Ruedemann 1921 Valid Silurian
  •  Norway
  •  Estonia
  •  United States
Moselopterus Størmer 1974 Valid Devonian
  •  Germany
  •  Latvia
Mycterops Cope 1886 Valid Carboniferous
  •  Scotland
  •  Belgium
  •  United States
Nanahughmilleria Kjellesvig-Waering 1961 Valid Silurian - Devonian
  •  England
  •  Norway
  •  Russia
  •  Scotland
  •  United States
Necrogammarus Woodward 1870 Valid Silurian
  •  England
Onychopterella Størmer 1951 Valid Ordovician - Silurian
  •  South Africa
  •  United States

Onychopterus

Miller & Gurley

1896

Preoccupied[13]

N/A

N/A

Preoccupied name, changed to Onychopterella.

Orcanopterus Stott, Tetlie, Braddy, Nowlan, Glasser & Devereux 2005 Valid Ordovician
  •  Canada
Pagea Waterston 1962 Valid Devonian
  •  Canada

Paracarcinosoma

Caster & Kjellesvig-Waering

1964

Jr. synonym

N/A

N/A

Synonym of Eusarcana.

Paraeurypterus Lamsdell, Hoşgör & Selden 2013 Valid Ordovician
  •  Turkey
Parahughmilleria Kjellesvig-Waering 1961 Valid Silurian - Devonian
  •  Canada
  •  England
  •  Germany
  •  Luxembourg
  •  Russia
  •  Scotland
  •  United States
Parastylonurus Kjellesvig-Waering 1966 Valid Silurian
  •  Scotland
  •  England

Parka

Fleming

1831

Misidentification[15]

N/A

N/A

Misidentified, a plant.

Pentecopterus[16] Lamsdell et al. 2015 Valid Ordovician
  •  United States
Pentlandopterus Lamsdell, Hoşgör & Selden 2013 Valid Ordovician
  •  Scotland
Pittsfordipterus Kjellesvig-Waering & Leutze 1966 Valid Silurian
  •  United States

Polystomurum[17]

Novojilov

1958

Misidentification

N/A

N/A

Misidentified, an unplaced euchelicerate.[1]

Polyzosternites

Goldenberg

1873

Jr. synonym

N/A

N/A

Synonym of Adelophthalmus.

Pterygotus Agassiz 1839 Valid Silurian - Devonian Nearly cosmopolitan, see article.
Rhenopterus Størmer 1936 Valid Devonian
  •  Germany
Rhinocarcinosoma Novojilov 1962 Valid Silurian
  •  United States
  •  Vietnam
Ruedemannipterus Kjellesvig-Waering 1966 Valid Silurian
  •  United States
Salteropterus Kjellesvig-Waering 1951 Valid Silurian
  •  England

Sidneyia

Walcott

1911

Misidentification[18]

N/A

N/A

Misidentified, an arachnomorph.[19]

Slimonia Page 1856 Valid Silurian
  •  England
  •  Scotland
  •  Bolivia
Soligorskopterus[20] Plax, Lamsdell, Vrazo & Barbikov 2018 Valid Devonian
  •  Belarus
Stoermeropterus Lamsdell 2011 Valid Devonian
  •  Norway
  •  Scotland
  •  United States

Strabops

Beecher

1901

Misidentification[21]

N/A

N/A

Misidentified, a strabopid.

Strobilopterus Ruedemann 1935 Valid Silurian - Devonian
  •  Estonia
  •  United States
Stylonurella Kjellesvig-Waering 1966 Valid Silurian - Devonian
  •  Scotland
  •  United States
Stylonuroides Kjellesvig-Waering 1966 Valid Silurian - Devonian
  •  Norway
  •  Russia
Stylonurus Page 1856 Valid Devonian
  •  Scotland
  •  United States

Syntomopterella

Tellie

2007

Jr. synonym

N/A

N/A

Synonym of Strobilopterus.

Syntomopterus

Kjellesvig-Waering

1961

Preoccupied

N/A

N/A

Preoccupied name, changed to Strobilopterus.

Tarsopterella Størmer 1951 Valid Devonian
  •  Scotland

Tarsopterus

Clarke & Ruedemann

1912

Preoccupied[13]

N/A

N/A

Preoccupied name, changed to Tarsopterella.

Terropterus[22] Wang, Dunlop, Gai, Lei, Jarzembowski & Wang 2021 Valid Silurian
  •  China

Truncatiramus

Kjellesvig-Waering

1961

Jr. synonym

N/A

N/A

Synonym of Erettopterus.

Tylopterella Størmer 1951 Valid Silurian
  •  Canada

Tylopterus

Clarke & Ruedemann

1912

Preoccupied[13]

N/A

N/A

Preoccupied name, changed to Tylopterella.

Unionopterus Chernyshev 1948 Valid Carboniferous
  •  Kazakhstan
Vinetopterus Poschmann & Tellie 2004 Valid Devonian
  •  Germany
Vernonopterus Waterston 1968 Valid Carboniferous
  •  Scotland
Waeringopterus Leutze 1961 Valid Silurian
  •  United States
Wiedopterus[23] Poschmann 2015 Valid Devonian
  •  Germany

Willwerathia

Størmer

1936

Misidentification[24]

N/A

N/A

Misidentified, a xiphosuran.

Woodwardopterus Kjellesvig-Waering 1959 Valid Carboniferous
  •  Scotland
Acutiramus.
Adelophthalmus.
Brachyopterus.
Campylocephalus.
Carcinosoma.
Ciurcopterus.
Dolichopterus.
Drepanopterus.
Erettopterus.
Eurypterus.
Eusarcana.
Hallipterus.
Herefordopterus.
Hibbertopterus.
Hughmilleria.
Jaekelopterus.
Kokomopterus.
Laurieipterus.
Megalograptus.
Megarachne.
Mixopterus.
Nanahughmilleria.
Onychopterella.
Orcanopterus.
Pentecopterus.
Pterygotus.
Slimonia.
Stoermeropterus.
Strobilopterus.
Stylonurus.

Eurypterid ichnogenera

Genus Author(s) Year Status Age Location Notes
Arcuites[25] Vrazo & Ciurca 2017 Valid Silurian
  •  Canada
  •  United States
Merostomichnites[26] Packard 1900 Valid Ordovician - Silurian
  •  Norway
  •  Portugal
Merostomichnites tracks in Norway were likely made by the genus Mixopterus.
Palmichnium[27] Richter 1954 Valid Devonian - Carboniferous
  •  Australia
  •  Canada
  •  United States
  •  Wales

See also

References

General

Citations

  1. 1.0 1.1 1.2 Dunlop, J. A., Penney, D. & Jekel, D. 2018. A summary list of fossil spiders and their relatives. In World Spider Catalog. Natural History Museum Bern
  2. 2.0 2.1 O. Erik Tetlie (2006). "Two new Silurian species of Eurypterus (Chelicerata: Eurypterida) from Norway and Canada and the phylogeny of the genus". Journal of Systematic Palaeontology 4 (4): 397–412. doi:10.1017/S1477201906001921. ISSN 1478-0941. http://www.museunacional.ufrj.br/mndi/Aracnologia/Eurypterida/Tetlie%202006%20Silurian%20Eurypterus.pdf. Retrieved May 20, 2011. 
  3. Bicknell, Russell D. C.; Smith, Patrick M.; Poschmann, Markus (2020). "Re-evaluating evidence of Australian eurypterids". Gondwana Research 86: 164–181. doi:10.1016/j.gr.2020.06.002. Bibcode2020GondR..86..164B. https://www.sciencedirect.com/science/article/abs/pii/S1342937X20301799. 
  4. Rolfe, William; Dzik, Jerzy (2006-12-01). "Angustidontus, a Late Devonian pelagic predatory crustacean". Transactions: Earth Sciences 97: 75–96. doi:10.1017/S0263593300001413. https://www.researchgate.net/publication/233549008. 
  5. Miller, Randall F. (1995-08-01). "The status of Belinuropsis wigudensis Matthew, 1910, a Pennsylvanian merostome from New Brunswick, Canada". Atlantic Geology 31 (2). doi:10.4138/2104. ISSN 1718-7885. https://journals.lib.unb.ca/index.php/ag/article/view/2104. 
  6. Anderson, Lyall I.; Moore, Rachel A. (2003). "Bembicosoma re-examined: a xiphosuran from the Silurian of the North Esk Inlier, Pentland Hills, Scotland". Transactions of the Royal Society of Edinburgh: Earth Sciences 94 (3): 199–206. doi:10.1017/S0263593300000614. https://www.cambridge.org/core/journals/earth-and-environmental-science-transactions-of-royal-society-of-edinburgh/article/bembicosoma-reexamined-a-xiphosuran-from-the-silurian-of-the-north-esk-inlier-pentland-hills-scotland/DFB7E5DD24F184786D1769B5DBBD45F2. 
  7. Miller, Randall F.; Tetlie, O. Erik (2007). "The presumed Synziphosuran Bunodella horrida Matthew, 1889 (Silurian; Cunningham Creek Formation, New Brunswick, Canada) is a eurypterid". Journal of Paleontology 81 (3): 588–590. doi:10.1666/05127.1. 
  8. Waterston, Charles D. (1958). "XII.—The Scottish Carboniferous Eurypterida". Transactions of the Royal Society of Edinburgh 63 (2): 265–288. doi:10.1017/S0080456800009492. https://www.cambridge.org/core/journals/earth-and-environmental-science-transactions-of-royal-society-of-edinburgh/article/xiithe-scottish-carboniferous-eurypterida/27C454542B1D42B89B354EC440ED7126. 
  9. Tetlie, Odd Erik (2004). Eurypterid phylogeny with remarks on the origin of arachnids (PhD). University of Bristol. pp. 1–344.
  10. 10.0 10.1 Hughes, Emily Samantha (2019). Discerning the diets of sweep-feeding eurypterids through analyses of mesh-modified appendage armature (MS thesis). West Virginia University. pp. 1–61. doi:10.33915/etd.3890.
  11. "†Eurypterida" (in en). http://www.helsinki.fi/~mhaaramo/metazoa/protostoma/arthropoda/eurypterida/eurypterida_1.html. 
  12. Dunlop, Jason A.; Miller, Randall F. (2007-10-16). "The fossil arachnid genus Eurymartus MATTHEW, 1895 and the eurypterid genus Eurypterella MATTHEW, 1889 from the Late Carboniferous" (in en). Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 245 (3): 295–300. doi:10.1127/0077-7749/2007/0245-0295. https://www.schweizerbart.de/papers/njgpa/detail/245/59169/The_fossil_arachnid_genus_Eurymartus_MATTHEW_1895_and_the_eurypterid_genus_Eurypterella_MATTHEW_1889_from_the_Late_Carboniferous_Fern_Ledges_of_Saint_John_New_Brunswick_Canada?l=FR. 
  13. 13.0 13.1 13.2 13.3 Størmer, Leif. "A new eurypterid from the Ordovician of Montgomeryshire, Wales." Geological Magazine; November 1951; v. 88; no. 6; p. 409-422.
  14. D. Waterston, Charles (1964-01-01). "II.—Observations on Pterygotid Eurypterids". Transactions of the Royal Society of Edinburgh 66 (2): 9–33. doi:10.1017/S0080456800023309. https://www.researchgate.net/publication/259412608. 
  15. Niklas, Karl J. (1976). "Morphological and Ontogenetic Reconstructions of Parka decipiens Fleming and Pachytheca Hooker from the Lower Old Red Sandstone, Scotland". Transactions of the Royal Society of Edinburgh 69 (21): 483–499. doi:10.1017/S0080456800015507. https://www.cambridge.org/core/journals/earth-and-environmental-science-transactions-of-royal-society-of-edinburgh/article/21morphological-and-ontogenetic-reconstructions-of-parka-decipiens-fleming-and-pachytheca-hooker-from-the-lower-old-red-sandstone-scotland/15260D8681E6CA5739D5C8F1643F85BE. 
  16. Lamsdell, James C.; Briggs, Derek E. G.; Liu, Huaibao P.; Witzke, Brian J.; McKay, Robert M. (2015-09-01). "The oldest described eurypterid: a giant Middle Ordovician (Darriwilian) megalograptid from the Winneshiek Lagerstätte of Iowa". BMC Evolutionary Biology 15: 169. doi:10.1186/s12862-015-0443-9. ISSN 1471-2148. PMID 26324341. 
  17. Novojilov, N. (1958). Aleksandr Grigorevich Sharov. ed. Polystomurum strømeri new genus and species of Eurypterida from the Devonian of the region of Voronell. Mat. Princip. Paleont. (Moscow). 2. pp. 47–50. https://books.google.com/books?id=4XCABwAAQBAJ. 
  18. Walcott, Charles Doolittle (1911). "Cambrian Geology and Paleontology". Smithsonian Miscellaneous Collections 57 (2): 17–40. https://en.wikisource.org/wiki/Cambrian_Geology_and_Paleontology/Volume_2/Middle_Cambrian_Merostomata. 
  19. Zhang, Xingliang; Han, Jian; Shu, Degan (2002). "New occurrence of the Burgess Shale arthropod Sidneyia in the Early Cambrian Chengjiang Lagerstätte (South China), and revision of the arthropod Urokodia". Alcheringa: An Australasian Journal of Palaeontology 26 (1): 1–8. doi:10.1080/03115510208619239. 
  20. Plax, Dmitry P.; Lamsdell, James C.; Vrazo, Matthew B.; Barbikov, Dmitry V. (2018). "A new genus of eurypterid (Chelicerata, Eurypterida) from the Upper Devonian salt deposits of Belarus". Journal of Paleontology 92 (5): 838–849. doi:10.1017/jpa.2018.11. https://www.researchgate.net/publication/326233486. 
  21. Hou, X.-G; Bergström, J (1997-12-22). "Arthropods of the Lower Cambrian Chengjiang fauna, southwest China". Fossils and Strata 45: 1–116. https://www.researchgate.net/publication/279900184. 
  22. Wang, Han; Dunlop, Jason A.; Gai, Zhikun; Lei, Xiaojie; Jarzembowski, Edmund A.; Wang, Bo (2021). "First mixopterid eurypterids (Arthropoda: Chelicerata) from the Lower Silurian of South China". Science Bulletin 66 (22): 2277–2280. doi:10.1016/j.scib.2021.07.019. PMID 36654455. Bibcode2021SciBu..66.2277W. https://www.researchgate.net/publication/353240547. 
  23. Poschmann, Markus (2015-12-01). "Sea scorpions (Chelicerata, Eurypterida) from the Lower Devonian (Siegenian) of the Lahrbach Valley/Westerwald area (SW Germany, Rhineland-Palatinate)" (in en). Paläontologische Zeitschrift 89 (4): 783–793. doi:10.1007/s12542-015-0261-9. ISSN 0031-0220. 
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