Biology:List of pinnipeds

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Short description: Species in mammal infraorder Pinnipedia
Five pinniped species, clockwise from top left: New Zealand fur seal (Arctocephalus forsteri), southern elephant seal (Mirounga leonina), Steller sea lion (Eumetopias jubatus), walrus (Odobenus rosmarus), and grey seal (Halichoerus grypus)
Combined range of all pinnipeds

Pinnipedia is an infraorder of mammals in the order Carnivora, composed of seals, sea lions, and the walrus. A member of this group is called a pinniped or a seal.[lower-alpha 1] They are widespread throughout the ocean and some larger lakes, primarily in colder waters. Pinnipeds range in size from the 1.1 m (3 ft 7 in) and 50 kg (110 lb) Baikal seal to the 6 m (20 ft) and 3,700 kg (8,200 lb) male southern elephant seal, which is also the largest member of Carnivora.[1] Several species exhibit sexual dimorphism, such as the southern elephant seal, where the males can be more than three times as long and six times as massive as the females, or the Ross seal, which has females typically larger than the males. Four seal species are estimated to have over one million members, while six are classified as endangered with population counts as low as 600, and two, the Caribbean monk seal and the Japanese sea lion, went extinct in the 20th century.

The 34 extant species of Pinnipedia are split into 22 genera within 3 families: Odobenidae, comprising the walrus; Otariidae, the eared seals, split between the sea lions and fur seals; and Phocidae, the earless or true seals. Odobenidae and Otariidae are combined into the superfamily Otarioidea, with Phocidae in Phocoidea. Extinct species have also been placed into the three extant families, as well as the extinct family Desmatophocidae, though most extinct species have not been categorized into a subfamily. Nearly one hundred extinct Pinnipedia species have been discovered, though due to ongoing research and discoveries the exact number and categorization is not fixed.

Conventions

Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the pinniped's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted. All extinct species or subspecies listed alongside extant species went extinct after 1500 CE, and are indicated by a dagger symbol "Extinct".

Classification

The infraorder Pinnipedia consists of 3 families containing 34 extant species belonging to 22 genera and divided into 48 extant subspecies, as well the extinct Caribbean monk seal and Japanese sea lion species, which are the only pinniped species to go extinct since prehistoric times. This does not include hybrid species or extinct prehistoric species.

Pinnipedia  
 Odobenidae  

Odobenus  

 Otariidae  

Arctocephalus  

Neophoca  

Phocarctos  

Otaria  

Zalophus  

Eumetopias  

Callorhinus  

Otarioidea
Phocidae  

Pusa  

Phoca  

Halichoerus  

Histriophoca  

Pagophilus  

Cystophora  

Erignathus  

Leptonychotes  

Hydrurga  

Lobodon  

Ommatophoca  

Mirounga  

Monachus  

Neomonachus  

Phocoidea

Pinnipeds

The following classification is based on the taxonomy described by Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis. This includes splitting the monk seal genus Monachus into Monachus and Neomonachus, the reorganization of grey seal subspecies, and the removal of the Laptev walrus subspecies.[2][3][4]

Family Odobenidae

Genus OdobenusBrisson, 1762 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Walrus

Brown walrus

O. rosmarus
(Linnaeus, 1758)

Arctic Ocean and subarctic seas
Map of range
Size: Male: 270–356 cm (106–140 in) long; 800–1,700 kg (1,764–3,748 lb)
Female: 225–312 cm (89–123 in) long; 400–1,250 kg (882–2,756 lb)[5]

Habitat: Neritic marine, oceanic marine, intertidal marine, coastal marine, and other[6]

Diet: Primarily eats bivalve mollusks, as well as other invertebrates, slow-moving fish, and occasionally birds, seals, and other marine mammals[6]
 VU 


112,500 Template:Population change unknown[6]

Family Otariidae

Genus ArctocephalusGeoffroy, F. Cuvier, 1826 – eight species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Antarctic fur seal

Brown and gray seal

A. gazella
(Peters, 1875)
Subantarctic islands
Map of range
Size: Male: 180 cm (71 in) long; 130–200 kg (287–441 lb)
Female: 120–140 cm (47–55 in) long; 22–50 kg (49–110 lb)[7]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[7]

Diet: Eats krill, cephalopods, fish, and penguins[7]
 LC 


700,000–1,000,000 Population declining[7]

Brown fur seal

Brown and gray seal

A. pusillus
(Schreber, 1775)

Southern African and Australian coasts (dark blue indicates breeding grounds)
Map of range
Size: Male: 201–227 cm (79–89 in) long; 218–360 kg (481–794 lb)
Female: 136–171 cm (54–67 in) long; 41–113 kg (90–249 lb)[8]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[8]

Diet: Eat a wide variety of fish, cephalopods, and crustaceans, and sometimes African penguins and other seabirds[8]
 LC 


1,060,000 Population increasing[8]

Galápagos fur seal

Brown and gray seal

A. galapagoensis
Heller, 1904
Galápagos Islands
Map of range
Size: Male: 150–160 cm (59–63 in) long; 60–68 kg (132–150 lb)
Female: 110–130 cm (43–51 in) long; 27–33 kg (60–73 lb)[9]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[9]

Diet: Eats small squids and a variety of fish[9]
 EN 


10,000 Population declining[9]

Guadalupe fur seal

Gray seal

A. townsendi
Merriam, 1897
Islands off of the southern Pacific coast of North America (dark blue indicates breeding grounds)
Map of range
Size: Male: 180 cm (71 in) long; 160–170 kg (353–375 lb)
Female: 148 cm (58 in) long; 49 kg (108 lb)[10]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[10]

Diet: Primarily eats squid, as well as fish[10]
 LC 


10,000 Population increasing[10]

Juan Fernández fur seal

Gray seal

A. philippii
(Peters, 1866)
Islands off of the Pacific coast of South America
Map of range
Size: Male: 150–200 cm (59–79 in) long; 140 kg (309 lb)
Female: 140 cm (55 in) long; 50 kg (110 lb)[11]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[12]

Diet: Primarily eat lanternfish, as well as a narrow range of fish from the Scomberesocidae, Carangidae, Engraulidae, and Bathylagidae families, and cephalopods[12]
 LC 


16,000 Population increasing[12]

New Zealand fur seal

Brown and gray seal

A. forsteri
(Lesson, 1828)
Southern Australian and New Zealand coasts
Map of range
Size: Male: 150–250 cm (59–98 in) long; 120–180 kg (265–397 lb)
Female: 100–150 cm (39–59 in) long; 30–50 kg (66–110 lb)[13]

Habitat: Forest, shrubland, neritic marine, oceanic marine, intertidal marine, and coastal marine[14]

Diet: Eat a wide variety of cephalopods, fish, and birds[14]
 LC 


100,000 Population increasing[14]

South American fur seal

Brown and gray seal

A. australis
(Zimmermann, 1783)

Southern South American coasts
Map of range
Size: Male: 200 cm (79 in) long; 90–200 kg (198–441 lb)
Female: 140 cm (55 in) long; 60 kg (132 lb)[15]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[15]

Diet: Primarily eat pelagic fish, demersal fish, and cephalopods[15]
 LC 


109,500 Population increasing[15]

Subantarctic fur seal

Brown and gray seal

A. tropicalis
(Gray, 1872)
Southern parts of the Indian, Pacific, and Atlantic Oceans
Map of range
Size: Male: 180 cm (71 in) long; 70–165 kg (154–364 lb)
Female: 119–152 cm (47–60 in) long; 25–67 kg (55–148 lb)[16]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[16]

Diet: Primarily eats lanternfish, cod icefish, and cephalopods, as well as crustaceans and rockhopper penguins[16]
 LC 


200,000 Population steady[16]

Genus CallorhinusGray, 1859 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Northern fur seal

Brown seal

C. ursinus
(Linnaeus, 1758)
Northern Pacific Ocean (dark blue indicates breeding grounds)
Map of range
Size: Male: 213 cm (84 in) long; 180–275 kg (397–606 lb)
Female: 142 cm (56 in) long; 40–50 kg (88–110 lb)[17]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[17]

Diet: Eats a variety of epipelagic and vertically migrating mesopelagic fish and squid[17]
 VU 


650,000 Population declining[17]

Genus EumetopiasGill, 1866 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Steller sea lion

Brown seal

E. jubatus
(Schreber, 1776)
Northern Pacific Ocean (red indicates breeding grounds)
Map of range
Size: Male: 300–340 cm (118–134 in) long; 1,120 kg (2,469 lb)
Female: 230–290 cm (91–114 in) long; 350 kg (772 lb)[18]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[19]

Diet: Eats a variety of fish and cephalopods, as well as northern fur seal, harbor seals, and ringed seals[19]
 NT 


81,300 Population increasing[19]

Genus NeophocaGray, 1866 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Australian sea lion

Brown seal

N. cinerea
(Péron, 1816)
Southwestern Australian coast
Map of range
Size: Male: 180–250 cm (71–98 in) long; 180–250 kg (397–551 lb)
Female: 130–180 cm (51–71 in) long; 61–105 kg (134–231 lb)[20]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[20]

Diet: Eats cephalopods, fish, and crustaceans[20]
 EN 


6,500 Population declining[20]

Genus OtariaPéron, 1816 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
South American sea lion

Brown seal

O. flavescens
(Shaw, 1800)
Southeastern and western South American coast and islands
Map of range
Size: Male: 210–260 cm (83–102 in) long; 300–350 kg (661–772 lb)
Female: 150–200 cm (59–79 in) long; 170 kg (375 lb)[21]

Habitat: Inland wetlands, neritic marine, oceanic marine, intertidal marine, and coastal marine[21]

Diet: Eats a wide variety of benthic fish, pelagic fish, and invertebrates[21]
 LC 


222,500 Population steady[21]

Genus PhocarctosPeters, 1866 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
New Zealand sea lion

Brown seal

P. hookeri
(Gray, 1844)
Southern New Zealand coast and islands
Map of range
Size: Male: 210–270 cm (83–106 in) long; 300–450 kg (661–992 lb)
Female: 180–200 cm (71–79 in) long; 90–165 kg (198–364 lb)[22]

Habitat: Forest, shrubland, neritic marine, oceanic marine, intertidal marine, and coastal marine[22]

Diet: Eats a wide variety of fish, cephalopods, and crustaceans, as well as penguins[22]
 EN 


3,000 Population declining[22]

Genus ZalophusGill, 1866 – three species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
California sea lion

Brown and gray seal

Z. californianus
(Lesson, 1828)
Pacific North American coast (dark blue indicates breeding grounds)
Map of range
Size: 165–220 cm (65–87 in) long; 275–390 kg (606–860 lb)[23]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[24]

Diet: Eats a variety of fish and squid[24]
 LC 


180,000 Population increasing[24]

Galápagos sea lion

Brown seal

Z. wollebaeki
Sivertsen, 1953
Galápagos Islands
Map of range
Size: Male: 200–250 cm (79–98 in) long; 200–400 kg (441–882 lb)
Female: 150–200 cm (59–79 in) long; 50–110 kg (110–243 lb)[25]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[26]

Diet: Eats sardines, as well as lanternfish, deep-sea smelt, and small squid[26]
 EN 


9,200–10,600 Population declining[26]

Japanese sea lionExtinct

Black and white photo of seals

Z. japonicus
(Peters, 1866)
Sea of Japan Size: Male: 230–250 cm (91–98 in) long; 450–560 kg (992–1,235 lb)
Female: 160 cm (63 in) long[27]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[27]

Diet: Unknown; believed to be similar to California sea lions[27]
 EX 


0 Population steady[27]

Family Phocidae

Genus CystophoraAgardh, 1841 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Hooded seal

Yellow and black hooded seal

C. cristata
(Erxleben, 1777)
Central and western North Atlantic ocean (blue indicates breeding grounds)
Map of range
Size: Male: 250–270 cm (98–106 in) long; 200–400 kg (441–882 lb)
Female: 200–220 cm (79–87 in) long; 145–300 kg (320–661 lb)[5]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[28]

Diet: Preys on fish and invertebrates throughout the water column[28]
 VU 


340,000 Template:Population change unknown[28]

Genus ErignathusGill, 1866 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Bearded seal

Gray seal

E. barbatus
(Erxleben, 1777)

Arctic ocean
Map of range
Size: 200–260 cm (79–102 in) long; 200–360 kg (441–794 lb) tail[5]

Habitat: Neritic marine, oceanic marine, and intertidal marine[29]

Diet: Primarily eats crabs, shrimp, clams, snails, benthic and demersal fish, and spoon worms[29]
 LC 


Unknown Template:Population change unknown[29]

Genus HalichoerusNilsson, 1820 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Grey seal

Gray seal

H. grypus
(Fabricius, 1791)

Shores of the North Atlantic Ocean
Map of range
Size: Male: 195–230 cm (77–91 in) long; 170–310 kg (375–683 lb)
Female: 165–195 cm (65–77 in) long; 105–186 kg (231–410 lb)[5]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[30]

Diet: Primarily eats benthic and demersal fish[30]
 LC 


316,000 Population increasing[30]

Genus HistriophocaGill, 1873 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Ribbon seal

Black and gray seal

H. fasciata
(Zimmermann, 1783)
Arctic and subarctic regions of the North Pacific Ocean (blue indicates reduced summer range)
Map of range
Size: 165–175 cm (65–69 in) long; 72–90 kg (159–198 lb)[31]

Habitat: Neritic marine and oceanic marine[32]

Diet: Preys on fish, crustaceans, and other invertebrates[32]
 LC 


183,000 Template:Population change unknown[32]

Genus HydrurgaGistel, 1848 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Leopard seal

Black and gray seal

H. leptonyx
(Blainville, 1820)
Antarctic Ocean
Map of range
Size: Male: 250–320 cm (98–126 in) long; 200–455 kg (441–1,003 lb)
Female: 241–338 cm (95–133 in) long; 225–591 kg (496–1,303 lb)[5]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[33]

Diet: Primarily eats krill, fish, squid, penguins, other seabirds, and juvenile seals[33]
 LC 


18,000 Template:Population change unknown[33]

Genus LeptonychotesGill, 1872 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Weddell seal

Black and gray seal

L. weddellii
(Lesson, 1826)
Coastal Antarctic Ocean
Map of range
Size: 280–330 cm (110–130 in) long; 400–600 kg (882–1,323 lb)[34]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[35]

Diet: Primarily eats cod icefish, as well as Antarctic toothfish, lanternfish, and cephalopods[35]
 LC 


300,000 Template:Population change unknown[35]

Genus LobodonGray, 1844 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Crabeater seal

Gray seal

L. carcinophaga
(Hombron, Jacquinot, 1842)
Antarctic Ocean
Map of range
Size: Male: 203–241 cm (80–95 in) long; 200–300 kg (441–661 lb)
Female: 216–241 cm (85–95 in) long; 200–300 kg (441–661 lb)[5]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[36]

Diet: Primarily eats Antarctic krill, as well as fish and squid[36]
 LC 


4,000,000 Template:Population change unknown[36]

Genus MiroungaGray, 1827 – two species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Northern elephant seal

Gray seal

M. angustirostris
Gill, 1866
Northeastern Pacific Ocean (red dots indicate breeding grounds)
Map of range
Size: Male: 400–500 cm (157–197 in) long; 2,000–2,700 kg (4,409–5,952 lb)
Female: 200–300 cm (79–118 in) long; 600–900 kg (1,323–1,984 lb)[5]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[37]

Diet: Primarily eats squid, lanternfish, and other mesopelagic fish[37]
 LC 


110,000 Population increasing[37]

Southern elephant seal

Brown seal

M. leonina
(Linnaeus, 1758)
Antarctic and subantarctic region
Map of range
Size: Male: 450–600 cm (177–236 in) long; 1,500–3,700 kg (3,307–8,157 lb)
Female: 200–300 cm (79–118 in) long; 400–600 kg (882–1,323 lb)[5]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[38]

Diet: Primarily eats lanternfish, cod icefish, and squid[38]
 LC 


325,000 Population steady[38]

Genus MonachusFleming, 1822 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Mediterranean monk seal

Gray seal

M. monachus
(Hermann, 1779)
Scattered portions of the Mediterranean Sea Size: 230–280 cm (91–110 in) long; 240–300 kg (529–661 lb)[39]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[40]

Diet: Eats benthic fish, pelagic fish, cephalopods, and lobsters[39][40]
 VU 


400–600 Population increasing[40]

Genus NeomonachusSlater, Helgen, 2014 – two species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Caribbean monk sealExtinct

Gray seal

N. tropicalis
(Gray, 1850)
Caribbean Sea Size: 200–240 cm (79–94 in) long; 200 kg (441 lb)[5][39]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[41]

Diet: Believed to have eaten eels, lobsters, octopus, and coral reef fish[41]
 EX 


0 Population steady[41]

Hawaiian monk seal

Gray seal

N. schauinslandi
(Matschie, 1905)
Hawaiian islands
Map of range
Size: 210–250 cm (83–98 in) long; 170–240 kg (375–529 lb)[39]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[42]

Diet: Eats benthic fish, pelagic fish, cephalopods, and lobsters[39][42]
 EN 


600 Population decreasing[42]

Genus OmmatophocaGray, 1844 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Ross seal

Gray seal

O. rossii
Gray, 1844
Coastal Antarctic Ocean
Map of range
Size: Male: 168–208 cm (66–82 in) long; 129–216 kg (284–476 lb)
Female: 190–250 cm (75–98 in) long; 159–204 kg (351–450 lb)[5]

Habitat: Neritic marine, oceanic marine, and coastal marine[43]

Diet: Primarily eats squid, as well as fish and krill[43]
 LC 


40,000 Template:Population change unknown[43]

Genus PagophilusGray, 1844 – one species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Harp seal

Black and gray seal

P. groenlandicus
(Erxleben, 1777)

Northern Atlantic Ocean
Map of range
Size: Male: 171–190 cm (67–75 in) long; 135 kg (298 lb)
Female: 168–183 cm (66–72 in) long; 120 kg (265 lb)[44]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[45]

Diet: Eats a wide variety of fish and invertebrates[45]
 LC 


4,500,000 Population increasing[45]

Genus PhocaLinnaeus, 1758 – two species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Spotted seal

Gray seal

P. largha
Pallas, 1811
Northern Pacific Ocean (dark areas indicate breeding grounds)
Map of range
Size: Male: 161–176 cm (63–69 in) long; 85–110 kg (187–243 lb)
Female: 151–169 cm (59–67 in) long; 65–115 kg (143–254 lb)[39]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[46]

Diet: Eats a variety of fish, cephalopods, and crustaceans[46]
 LC 


320,000 Template:Population change unknown[46]

Harbor seal

Gray seal

P. vitulina
Gray, 1864

Northern Hemisphere coastlines
Map of range
Size: Male: 160–186 cm (63–73 in) long; 87–170 kg (192–375 lb)
Female: 148–169 cm (58–67 in) long; 60–142 kg (132–313 lb)[39]

Habitat: Neritic marine, oceanic marine, intertidal marine, and coastal marine[47]

Diet: Eats a wide variety of fish, cephalopods, and crustaceans[47]
 LC 


190,000 Population increasing[47]

Genus PusaScopoli, 1771 – three species
Common name Scientific name and subspecies Range Size and ecology IUCN status and estimated population
Baikal seal

Gray seal

P. sibirica
Gmelin, 1788
Lake Baikal
Map of range
Size: 110–140 cm (43–55 in) long; 50–130 kg (110–287 lb)[48]

Habitat: Inland wetlands[48]

Diet: Primarily eat sculpin fish, as well as other fish[48]
 LC 


54,000 Population steady[48]

Caspian seal

Gray seal

P. caspica
Gmelin, 1788
Caspian Sea
Map of range
Size: 126–140 cm (50–55 in) long; 50–86 kg (110–190 lb)[49][50]

Habitat: Inland wetlands[49]

Diet: Eats a variety of fish[49]
 EN 


68,000 Template:Population change unknown[49]

Ringed seal

Gray seal

P. hispida
Schreber, 1775

Arctic Ocean
Map of range
Size: 110–175 cm (43–69 in) long; 32–124 kg (71–273 lb)[51]

Habitat: Inland wetlands, neritic marine, and oceanic marine[52]

Diet: Eats a variety of invertebrates and small fish[52]
 LC 


1,500,000 Template:Population change unknown[52]

Notes

  1. "Seal" can refer to the entire pinniped infraorder, but is often also used to mean either pinnipeds with the exception of the walrus, or fur seals and true seals while excluding the walrus and sea lions.

References

  1. "Southern Elephant Seal". Marine Life Encyclopedia. Oceana. http://oceana.org/marine-life/marine-mammals/southern-elephant-seal. Retrieved December 1, 2020. 
  2. Scheel, D.-M.; Slater, G. J.; Kolokotronis, S.-O.; Potter, C. W.; Rotstein, D. S.; Tsangaras, K.; Greenwood, A. D.; Helgen, K. M. (May 2014). "Biogeography and taxonomy of extinct and endangered monk seals illuminated by ancient DNA and skull morphology". ZooKeys (409): 1–33. doi:10.3897/zookeys.409.6244. PMID 24899841. 
  3. Olsen, M. T.; Galatius, A.; Biard, V.; Gregersen, K.; Kinze, C. C. (April 2016). "The forgotten type specimen of the grey seal [Halichoerus grypus (Fabricius, 1791)] from the island of Amager, Denmark". Zoological Journal of the Linnean Society 178 (3): 713–720. doi:10.1111/zoj.12426. 
  4. Higdon, J. W.; Stewart, D. B. (2018). State of Circumpolar Walrus Populations: Odobenus rosmarus. WWF Arctic Programme. p. 6. https://nammco.no/wp-content/uploads/2018/10/higdon-stewart-2018-1.pdf. Retrieved November 9, 2020. 
  5. 5.0 5.1 5.2 5.3 5.4 5.5 5.6 5.7 5.8 5.9 Nowak, Ronald M.; Walker, Ernest Pillsbury (2003). Walker's Marine Mammals of the World. Johns Hopkins University Press. pp. 92–113. ISBN 978-0-8018-7343-0. 
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  18. Keranen, Danielle (2013). "Eumetopias jubatus". University of Michigan. https://animaldiversity.org/accounts/Eumetopias_jubatus/. 
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