Biology:Capulidae

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The Capulidae, the cap snails or cap shells, are a taxonomic family of limpet-like sea snails, marine gastropod molluscs.[1]

Capulidae is the only family in the superfamily Capuloidea. Capulidae is assigned to the paraphyletic order Littorinimorpha,[1] but is more closely related to neogastropods than some other littorinimorphs as it is a member of the clade Latrogastropoda.[2]

The name Trichotropidae was previously used for this family, but this name is invalid, as it is a junior synonym.[3]

Characteristics

(Original description; described as Capulusidae) The foot is complicated on its anterior margin. The shell adheres to the animal by a circular muscle, leaving an opening in front for the issue of the head and the entrance to the branchial cavity. The gills form a single ridge across the roof. The aperture is in the form of an extended proboscis, with a deep groove above. The tentacula, which are two in number, have the eyes at their external base. The anus is on the right side of the branchial cavity. [4]

Cup-shaped or bonnet-like, the shells of this family start with flat, tightly coiled initial whorls. From this beginning, they expand rapidly in a planospiral way, flaring out dramatically like a funnel.

The interior of the shell is porcelaneous. Some forms develop internal partitions or septa, a feature that makes them superficially resemble the Calyptraeidae (slipper limpets). Externally, the shell is covered by a thick periostracum that feels either felt-like or hairy. There is no operculum.

Species in this family are parasites, praying mostly on other molluscs. They steal the food from these animals with a long proboscis.

Genera

Genera within the family Capulidae include:[5]

Subfamily Capulinae J. Fleming, 1822
  • Ariadnaria Habe, 1961
  • Armenostoma Hacobjan, 1976
  • Blackdownia Kollmann, 1976 [6]
  • Capulus Montfort, 1810
  • Cerithioderma Conrad, 1860 (?)
  • Ciliatotropis Golikov, 1986
  • Cryocapulus Schiaparelli, Bouchet, Fassio & Oliverio, 2020
  • Discotrichoconcha Powell, 1951
  • Echinospira Girotti, 1970 [citation needed]
  • Hyalorisia Dall, 1889
  • Icuncula Iredale, 1924
  • Krebsia Montfort, 1810 : synonym of Capulus Montfort, 1810
  • Latticosta S.-I Huang & M.-H. Lin, 2021
  • Lippistes Montfort, 1810 (?)
  • Neoiphinoe Habe, 1978
  • Malluvium Melvill, 1906 : belongs to the family Hipponicidae.
  • Profusinus Bandel, 2000
  • Rufodardanula Ponder, 1965; synonym of Skenella Pfeffer, 1886
  • Separatista Gray, 1847
  • Sirius Hedley, 1900
  • Tintorium Sohl, 1961
  • Torellia Lovén in Jeffreys, 1867
  • Trichamathina Habe, 1962
  • Trichosirius Finlay, 1926 (?)
  • Trichotropis Broderip and G. B. Sowerby I, 1829
  • Turbinopsis Conrad, 1860
  • Turritropis Habe, 1961
  • Vermeijia Amano, 2019
  • Verticosta S.-I Huang & M.-H. Lin, 2020
  • Xuwenospira H.-J. Wang, 1982
  • Zelippistes Finlay, 1927
Subfamily † Lysinae Saul & Squires, 2008
  • Garzasia Saul & Squires, 2008
  • Lysis Gabb, 1864
  • Spirogalerus H. J. Finlay & Marwick, 1937
Genera brought into synonymy
  • Actita Fischer von Waldheim, 1823: synonym of Capulus Montfort, 1810
  • Antitrichotropis Powell, 1951: synonym of Torellia Jeffreys, 1867
  • Ariadna P. Fischer, 1864: synonym of Ariadnaria T. Habe, 1961 (junior homonym, junior homonym of Ariadna Audouin, 1826 [Arachnida]; Ariadnaria is a replacement name)
  • Brocchia Bronn, 1828: synonym of Capulus Montfort, 1810
  • Capulonix Iredale, 1929: synonym of Capulus Montfort, 1810
  • Dolichosirius Iredale, 1931: synonym of Sirius Hedley, 1900 (junior subjective synonym)
  • Eosirius P. A. Maxwell, 1966: synonym of † Trichosirius (Eosirius) P. A. Maxwell, 1966 represented as Trichosirius H. J. Finlay, 1926
  • Iphinoe H. Adams & A. Adams, 1856 : synonym of Neoiphinoe Habe, 1978
  • Mesostoma Deshayes, 1861: synonym of † Cerithioderma Conrad, 1860 (junior subjective synonym, invalid: junior homonym of Mesostoma Ehrenberg, 1836)
  • Neoconcha E.A. Smith, 1907: synonym of Torellia (Neoconcha) E.A. Smith, 1907
  • Opposirius Iredale, 1931: synonym of Sirius Hedley, 1900
  • Ovotropis Egorov & Alexeyev, 1998: synonym of Neoiphinoe Habe, 1978
  • Pileopsis Lamarck, 1822: synonym of Capulus Montfort, 1810
  • Opposirius Iredale, 1931: synonym of Sirius Hedley, 1900
  • Trichoconcha E.A. Smith, 1907: synonym of Torellia Jeffreys, 1867
  • Tropidothais L. R. Cox, 1925: synonym of † Lysis Gabb, 1864 (junior subjective synonym)

References

  1. 1.0 1.1 Gofas, S. (2012). Capulidae. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=139 on 2012-07-24
  2. Colgan, Donald J.; Ponder, Winston F. (2025-07-28). "Redefining Latrogastropoda Again and Searching for Its Sister Group in Hypsogastropoda (Gastropoda: Caenogastropoda)". Diversity 17 (8): 524. doi:10.3390/d17080524. ISSN 1424-2818. Bibcode2025Diver..17..524C. 
  3. ITIS Capulidae Fleming, 1822
  4. Fleming, J. (1822). "The philosophy of zoology: or a general view of the structure, functions and classification of animals". Edinburgh 2: 494. https://www.biodiversitylibrary.org/page/28230157. Retrieved 6 November 2025. 
  5. Gofas, S. (2010). Capulidae. In: Bouchet, P.; Gofas, S.; Rosenberg, G. (2010) World Marine Mollusca database. Accessed through: World Register of Marine Species at http://www.marinespecies.eu/aphia.php?p=taxdetails&id=139 on 2010-06-06
  6. Kiel S. (2003) New taxonomic data for the gastropod fauna of the Umzamba Formation (Santonian–Campanian, South Africa); Cretaceous Research 24 (2003) 449–475
  • Powell A. W. B., New Zealand Mollusca, William Collins Publishers Ltd, Auckland, New Zealand 1979 ISBN 0-00-216906-1
  • Vaught, K.C. (1989). A classification of the living Mollusca. American Malacologists: Melbourne, FL (USA). ISBN 0-915826-22-4. XII, 195 pp
  • Discover Life
  • "Capulidae". Integrated Taxonomic Information System. https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=72606. 

Wikidata ☰ Q3141569 entry