Biology:Cochemiea

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Cochemiea is a genus of cactus.[1] The group was previously synonymized with Mammillaria, until molecular phylogenetic studies determined that—when broadly circumscribedMammillaria is not monophyletic; thus, Cochemiea has been accepted as a distinct genus.[2]

Description

Species within the genus Cochemiea are characterized by their elongated, cylindrical stems covered in spirally arranged tubercles. These stems are typically green or bluish-green, measuring 3 to 7 cm in diameter and reaching heights of 7 to 50 cm, with some individuals exceptionally growing up to 200 cm. The areoles bear 7 to 25 radial spines, each 1 to 2 cm long and varying in color from white to yellow or reddish-brown. Additionally, they possess 1 to 6 central spines (occasionally up to 11), measuring 1 to 5 cm in length. These central spines can be white with black tips, reddish-brown, or black. Flowers are zygomorphic, arranged in two rows, and emerge from the axils at the stem apex. They measure 3 to 5 cm in length, open during the day, and are typically red or purple, spherical, and indehiscent with a smooth surface and a scar at the tip. The seeds are black and measure 0.5 to 1 mm in diameter.[3][4]

Distribution

The native range of Cochemiea extends from the southwestern and south-central United States into Mexico. Specific locations include Arizona, California, the Mexican Pacific Islands, the Gulf of Mexico region, Nevada, New Mexico, Texas, and Utah. These plants thrive in diverse habitats, from full sun to the shade of shrubs, and can be found on rocky cliffs and canyons at altitudes up to 1,800 meters above sea level.

Taxonomy

The genus Cochemiea was initially described by American botanist Mary Katharine Brandegee as a subgenus within Mammillaria in 1897. In 1899, British botanist Frederick Arthur Walton elevated it to its current genus status in the publication Cactus Journal 2: 50. The genus name honors the extinct Cochimí indigenous tribe, who historically inhabited Baja California, a region within the natural distribution of these plants.[5]

A 2021 molecular phylogenetic study of the "mammilloid clade", which included the genera Cochemiea, Coryphantha, Cumarinia, Escobaria, Mammillaria, Neolloydia and Ortegocactus, showed that it consisted of four monophyletic groups, which the authors re-circumscribed into four genera: Cumarinia; Mammillaria, with a reduced number of species; Coryphantha, expanded to include species previously placed in Mammillaria and Escobaria; and Cochemiea, expanded to include a large number of species previously placed in Mammillaria, as well as Neolloydia conoidea.[2]

Mammilloid clade

Cumarinia

Mammillaria s.s.

Coryphantha s.l.

Cochemiea s.l.

Species

In 2021, Breslin, Wojciechowski and Majure placed the following species in the genus, some already placed there and some moved from Mammillaria, Neolloydia, Neomammillaria and Ortegocactus.[2] As of January 2026, Plants of the World Online accepted in the genus the species moved by Breslin et al.[1]

Image Scientific name Distribution
120px Cochemiea albicans (Britton & Rose) P.B.Breslin & Majure Mexico (Baja California)
120px Cochemiea angelensis (R.T.Craig) P.B.Breslin & Majure Mexico
120px Cochemiea armillata (K.Brandegee) P.B.Breslin & Majure Mexico (Baja California Sur)
120px Cochemiea barbata (Engelm.) Doweld Mexico (Chihuahua, Durango)
120px Cochemiea blossfeldiana (Boed.) P.B.Breslin & Majure Mexico (Guadalupe, Baja California)
120px Cochemiea boolii (G.E.Linds.) P.B.Breslin & Majure, Mexico (Sonora)
Cochemiea bullardiana (H.E.Gates) P.B.Breslin & Majure, Mexico (Baja California)
120px Cochemiea capensis (H.E.Gates) Doweld Mexico (Baja California Sur, Sonora)
Cochemiea cerralboa (Britton & Rose) P.B.Breslin & Majure Mexico (Baja California Sur).
120px Cochemiea dioica (K.Brandegee) Doweld S. California to NW. Mexico
120px Cochemiea estebanensis (G.E.Linds.) P.B.Breslin & Majure, Mexico (Island San Esteban)
120px Cochemiea fraileana (Britton & Rose) P.B.Breslin & Majure Mexico (Baja California Sur).
120px Cochemiea goodridgei (Scheer ex Salm-Dyck) P.B.Breslin & Majure Mexico (Baja California).
120px Cochemiea grahamii (Engelm.) Doweld SE. California to W. Texas and Mexico (Baja California Norte to Chihuahua)
120px Cochemiea guelzowiana (Werderm.) P.B.Breslin & Majure Mexico (Durango, Coahuila, Nuevo León)
120px Cochemiea halei (K.Brandegee) Walton Mexico (Baja California)
120px Cochemiea hutchisoniana (H.E.Gates) P.B.Breslin & Majure Mexico (Baja California)
120px Cochemiea insularis (H.E.Gates) P.B.Breslin & Majure Mexico (Baja California)
120px Cochemiea mainiae (K.Brandegee) P.B.Breslin & Majure S. Arizona to NW. Mexico
120px Cochemiea maritima H.E.Gates ex. Shurly Mexico (W. Central Baja California)
120px Cochemiea mazatlanensis (K.Schum.) D.Aquino & Dan.Sánchez Mexico (Colima, Jalisco, Michoacan, Nayarit, Sinaloa, Sonora)
120px Cochemiea multidigitata (Radley & G.E.Linds.) P.B.Breslin Mexico (Sonora).
120px Cochemiea palmeri (J.M.Coult.) P.B.Breslin & Majure Mexico (Baja California)
120px Cochemiea phitauiana (E.M.Baxter) Doweld Mexico (Baja California).
120px Cochemiea pondii (Greene) Walton Mexico (Island Cedros)
120px Cochemiea poselgeri (Hildm.) Britton & Rose Mexico (Baja California Sur)
120px Cochemiea saboae (Glass) Doweld Mexico (E. Sonora, W. Chihuahua)
120px Cochemiea schumannii (Hildm.) P.B.Breslin & Majure Mexico (Baja California Sur).
120px Cochemiea setispina Walton Mexico (C. Baja California).
120px Cochemiea sheldonii (Britton & Rose) Doweld Arizona to Mexico (Sonora, Sinaloa, W. Chihuahua)
120px Cochemiea tetrancistra (Engelm.) P.B.Breslin & Majure SW. U.S.A. to NW. Mexico
120px Cochemiea theresae (Cutak) Doweld Mexico (Durango, Zacatecas).
120px Cochemiea thomasii García-Mor. & al. Mexico (Sinaloa)
120px Cochemiea thornberi (Orcutt) P.B.Breslin & Majure S. Arizona to Mexico (Sonora).
Cochemiea viridiflora (Britton & Rose) P.B.Breslin & Majure Arizona to SW. New Mexico.
120px Cochemiea wrightii (Engelm.) Doweld E. Arizona to W. Texas and N. Mexico.

References

  1. 1.0 1.1 "Cochemiea (K.Brandegee) Walton", Plants of the World Online (Royal Botanic Gardens, Kew), https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:1154805-2, retrieved 2022-03-05 
  2. 2.0 2.1 2.2 Breslin, Peter B.; Wojciechowski, Martin F.; Majure, Lucas C. (2021), "Molecular phylogeny of the Mammilloid clade (Cactaceae) resolves the monophyly of Mammillaria", Taxon 70 (2): 308–323, doi:10.1002/tax.12451, Bibcode2021Taxon..70..308B 
  3. "Floraculture". https://www.booksite.ru/blumen/1997/1997_4.pdf. 
  4. "On-line Guide to the positive identification of Members of the Cactus Family". https://cactiguide.com/cactus/?genus=Cochemiea. 
  5. "Cochemiea". 2021-04-29. https://www.cactusnames.org/cochemiea/. 
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