Biology:Dryopteris hybrid complex

The fern genus Dryopteris is known to be one of the most freely-hybridizing of fern genera.

Parent species

• Dryopteris expansa
• Dryopteris goldieana
• Dryopteris intermedia
• Dryopteris ludoviciana
• Dryopteris marginalis

Hybrid species

• Dryopteris carthusiana (D. intermedia × unknown parent; allotetraploid)
• Dryopteris campyloptera (D. intermedia × D. expansa; allotetraploid)
• Dryopteris celsa (D. goldieana × D. ludoviciana; allotetraploid)
• Dryopteris clintoniana (D. cristata × D. goldieana; allohexaploid)
• Dryopteris cristata (D. goldieana × unknown parent; allotetraploid)

Other hybrids

• Dryopteris × bootii (D. cristata × D. intermedia; triploid)
• Dryopteris × critica (D. borreri × D. filix-mas)
• Dryopteris × complexa aggregate (D. filix-mas and D. affinis; tetraploid)
• Dryopteris × convoluta (D. cambrensis × D. filix-mas)
• Dryopteris × deweveri (D. dilatata × D. carthusiana)
• Dryopteris × neo-wherryi (D. goldieana × D. marginalis; diploid)
• Dryopteris × triploidea (D. carthusiana × D. intermedia; triploid)