Biology:Howesia

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Howesia is an extinct genus of basal rhynchosaur from early Middle Triassic (early Anisian stage) deposits of Eastern Cape, South Africa. It is known from the holotype. SAM-PK-5884 (a partial skeleton with palate and partial lower jaws), two paratypes (SAM-PK-5885 and SAM-PK-5886).[1] The known fossils were found in the Burgersdorp Formation of the middle deposits of the Beaufort Group (Karoo Basin) and referred to Subzone B of the Cynognathus Assemblage Zone.[2] The genus contains a single species, Howesia browni, named by Robert Broom in 1905, after Alfred Brown.[3] The contemporary Eohyosaurus wolvaardti (holotype SAM-PK-K10159) has been suggested to be a junior synonym of H. browni based on shared anatomical traits not recognized when the former was named in 2015.[4]

Classification

Life restoration

The placement of Howesia as a basal member of the Rhynchosauria is shown in the cladogram below, following the phylogenetic results of Spiekman et al. (2026).[4]

 
 

Mesosuchus browni

 
 

Noteosuchus colletti

 
 
 
 

Howesia browni

 
Rhynchosauridae
 

Rhynchosaurus articeps

 
 
 

Eifelosaurus triadicus

 
 
Stenaulorhynchinae

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Bentonyx sidensis

 
 
 

Langeronyx brodiei

 
 
 

Fodonyx spenceri

 
Hyperodapedontinae

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References

  1. ↑ David M. Dilkes (1995). "The rhynchosaur Howesia browni from the Lower Triassic of South Africa". Palaeontology 38: 665–685. 
  2. ↑ Richard J. Butler; Martín D. Ezcurra; Felipe C. Montefeltro; Adun Samathi; Gabriela Sobral (2015). "A new species of basal rhynchosaur (Diapsida: Archosauromorpha) from the early Middle Triassic of South Africa, and the early evolution of Rhynchosauria". Zoological Journal of the Linnean Society 174 (3): 571–588. doi:10.1111/zoj.12246. 
  3. ↑ Robert Broom (1905). "Preliminary notice of some new fossil Reptiles collected by Mr. Alfred Brown at Aliwal North, S. Africa". Records of the Albany Museum 1 (4): 269–275. 
  4. ↑ 4.0 4.1 Spiekman, Stephan N. F.; Browning, Claire; Tshibalanganda, Muofhe; Mukherjee, Debarati; Smith, Roger M. H.; Du Plessis, Anton; Wolvaardt, Frederik (2026-07-09). "Micro‐CT informed cranial anatomy of the early rhynchosaur Howesia browni, with implications for rhynchosaur skull evolution" (in en). Papers in Palaeontology 12 (4). doi:10.1002/spp2.70105. ISSN 2056-2799. 

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