Biology:Lichida

From HandWiki

Lichida is an order of typically spiny trilobite that lived from the Furongian to the Devonian period.[1] These trilobites usually have 8–13 thoracic segments.[2] Their exoskeletons often have a grainy texture or have wart or spine-like tubercles. Some species are extraordinarily spiny, having spiny thoracic segments that are as long or longer than the entire body, from cephalon (head) to pygidium (tail). The sections of the pygidia are leaf-like in shape and also typically end in spines.

The order is divided into two families, Lichidae, and Lichakephalidae.[2] Some experts group the families of the closely related order Odontopleurida within Lichida, too, whereupon the family is then divided into three superfamilies, Dameselloidea, containing the family Damesellidae, Lichoidea, containing the families Lichidae and Lichakephalidae, and Odontopleuroidea, containing the family Odontopleuridae.

Lichids are some of the largest trilobites, with the second and third largest trilobites (Uralichas hispanicus, and Teratapsis grandis) being in the order.[3]

Taxa traditionally placed within Lichida

Parvilichas marochii

As mentioned earlier, the order Lichida is divided into two families.

Lichidae

  • Acanthopyge
  • Akantharges
  • Allolichas
  • Amphilichas
  • Apatolichas
  • Arctinurus
  • Autoloxolichas
  • Borealarges
  • Ceratarges
  • Conolichas
  • Craspedarges
  • Dicranogmus
  • Dicranopeltis
  • Echinolichas
  • Eifliarges
  • Gaspelichas
  • Hemiarges
  • Homolichas
  • Hoplolichas
  • Hoplolichoides
  • Jasperia
  • Leiolichas
  • Lichas
  • Lobopyge
  • Lyralichas
  • Mephiarges
  • Metaleiolichas
  • Metalichas
  • Metopolichas
  • Neolichas
  • Nipponarges
  • Ohleum
  • Oinochoe
  • Otarozoum
  • Paraleiolichas
  • Perunaspis
  • Platylichas
  • Probolichas
  • Pseudotupolichas
  • Radiolichas
  • Richterarges
  • Rontrippia
  • Terataspis
  • Terranovia
  • Trimerolichas
  • Trochurus
  • Uralichas
  • Uripes

Lichakephalidae

  • Acidaspidella
  • Acidaspides
  • Acidaspidina
  • Archikainella
  • Belovia
  • Bestjubella
  • Brutonia
  • Colossaspis
  • Eoacidaspis
  • Lichakephalus
  • Lichakephalina
  • Metaacidaspis
  • Paraacidaspis
  • Usoviana

Cladogram

Pollit et. al. (2005) described the internal relationships below family level for Lichida.[4]

Lichidae
Lichinae
 
 
 
Echinolichini
 

Echinolichas

 
 

Terataspis

 
 

Conolichas

 
 

Otarozoum

 
 
Tetralichini
 

Apatolichas

 
 

Lyralichas

 
 

Amphilichas

 
 

Pseudotupolichas

 
 
 
Platylichini
 

Platylichas

 
 

Autoloxolichas

 
 

Allolichas

 
 

Arctinurus

 
 
 
Lichini
 

Lichas

 
 
 
Dicranopeltini
 

Dicranopeltis

 
 

Uralichas

 
 

Dicranogmus

 
 

Oinochoe

 
 

Metopolichas

 
 
 
Trochurinae
 

Acanthopyge

 
 

Trochurus

 
 

Ceratarges

 
 

Radiolichas

 
 

Borealarges

 
 

Richterarges

 
 

Hemiarges

 
 

Akantharges

 
 

Uripes

 
 

Hoplolichas

 
 

Hoplolichoides

 
 
 
Lichakephalidae
 

Lichakephalus

 
 

Lichakephalina

 
 

Eoacidaspis

 
 

Acidaspidina

 
 

Acidaspidella

 
 

References

Further reading

  • Margaret J. Campbell & Brian D. E. Chatterton (2009). "Silurian lichid trilobites from northwestern Canada: ontogeny and phylogeny of lichids". Journal of Paleontology 83 (2): 263–279. doi:10.1666/07-140R1.1. Bibcode: 2009JPal...83..263C. 
  • Data related to Lichida at Wikispecies

Wikidata ☰ Q3108659 entry