Biology:List of Cholevinae genera

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These 267 genera belong to the subfamily Cholevinae, small carrion beetles. There are at least 1,100 described species in Cholevinae.[1][2][3]


  • Acrotrychiopsis Normand, 1946
  • Adelopidius Apfelbeck, 1907
  • Adelopsella Jeannel, 1908
  • Adelopsis Portevin, 1907
  • Adelopspeleon Salgado, 2012
  • Afrocatops Jeannel, 1964
  • Albaniola Jeannel, 1924
  • Albanodirus Giachino & Vailati, 1998
  • Amplexella Gnaspini, 1996
  • Anemadiola Szymczakowski, 1963
  • Anemadus Reitter, 1884
  • Anillocharis Reitter, 1903
  • Anillochlamys Jeannel, 1909
  • Anisoscapha Muller, 1917
  • Anthroherpon Reitter, 1889
  • Antrocharis Abeille de Perrin, 1878
  • Antrodulus Knirsch, 1927
  • Antrosedes Reitter, 1912
  • Aphaobiella Gueorguiev, 1976
  • Aphaobius Abeille de Perrin, 1878
  • Aphaotus Breit, 1914
  • Apholeuonus Reitter, 1889
  • Apocatops Zwick, 1968
  • Apterocatops Miyama, 1985
  • Araucaniopsis Salgado Costas, 2005
  • Archeoboldoria Ghidini, 1937
  • Astagobius Reitter, 1886
  • Attaephilus Motschoulsky, 1870
  • Atticiella Coiffait, 1955
  • Attumbra Gozis, 1886
  • Augustia Zariquiey, 1927
  • Austrocholeva Zwick, 1979
  • Austronargus Zwick, 1979
  • Austronemadus Zwick, 1979
  • Babuniella Karaman, 1954
  • Balcanobius Gueorguiev, 1965
  • Banatiola Decu, 1967
  • Baronniesia Fresneda, Bourdeau & Faille, 2009
  • Baryodirus Perreau, 2000
  • Bathyscia Schiödte, 1848
  • Bathyscidius Jeannel, 1910
  • Bathysciella Jeannel, 1906
  • Bathyscimorphus Jeannel, 1910
  • Bathysciola Jeannel, 1910
  • Bathysciopsis Muller, 1941
  • Bathysciotes Jeannel, 1910
  • Bellesia Fresneda & Hernando, 1994
  • Beronia Gueorguiev, 1960
  • Beroniella Giachino & Gueorguiev, 1991
  • Beskovia Gueorguiev, 1960
  • Besuchetiola Rampini & Zoia, 1991
  • Bithyniella Jeannel, 1955
  • Blattochaeta Reitter, 1910
  • Blattodromus Reitter, 1904
  • Boldoria Jeannel, 1924
  • Breuilia Jeannel, 1909
  • Breuilites Salgado, 1980
  • Bulgariella Karaman, 1958
  • Bureschiana Gueorguiev, 1963
  • Canavesiella Giachino, 1993
  • Cansiliella Paoletti, 1972
  • Cantabrogeus Salgado, 2000
  • Capraiola Zoia & Rampini, 1994
  • Catopidius Jeannel, 1922
  • Catopodes Portevin, 1914
  • Catoposchema Jeannel, 1936
  • Catops Paykull, 1798
  • Catopsimorphus Aubé, 1850
  • Catoptrichus Murray, 1856
  • Cavazzutiella Casale & Giachino, 1985
  • Ceretophyes Comas & Escola, 1989
  • Ceuthmonocharis Jeannel, 1910
  • Ceuthophyes Jeannel, 1924
  • Chappuisiotes Jeannel, 1957
  • Charonites Apfelbeck, 1907
  • Chionocatops Ganglbauer, 1899
  • Choleva Latreille, 1796
  • Cholevinus Reitter, 1901
  • Cholevodes Portevin, 1928
  • Closania Jeannel, 1928
  • Coiffaitiola Jeannel, 1955
  • Coreobathyscia Szymczakowski, 1975
  • Croatodirus Casale, Giachino & Jalzic, 2000
  • Cryptobathyscia Vailati, 1980
  • Cytodromus Abeille de Perrin, 1876
  • Dalmatiola Jeannel, 1924
  • Deelemaniella Perreau, 2002
  • Dellabeffaella Capra, 1924
  • Diaprysius Abeille de Perrin, 1878
  • Dissochaetus Reitter, 1884
  • Dreposcia Jeannel, 1922
  • Drimeotus Miller, 1856
  • Dzungarites Jeannel, 1936
  • Elladoherpon Casale, 1983
  • Eocatops Peyerimhoff, 1924
  • Espanoliella Gueorguiev, 1976
  • Eucatops Portevin, 1903
  • Eunemadus Portevin, 1914
  • Euryspeonomus Jeannel, 1919
  • Excelsiorella Salgado, 2008
  • Falkocholeva Hatch, 1928
  • Falkonemadus Szymczakowski, 1961
  • Fissocatops Zwick, 1968
  • Fusi Perkovsky, 1989
  • Genestiellina Giachino, 1996
  • Gesciella Giachino & Guéorguiev, 1989
  • Graciliella Njunjić et al., 2016
  • Gueorguievella Giachino & Gueorguiev, 2006
  • Hadesia Muller, 1911
  • Halbherria Conci & Tamanini, 1951
  • Haplotropidius Muller, 1903
  • Henrotiella Perreau, 1999
  • Hexaurus Reitter, 1884
  • Himalops Perreau, 1986
  • Hoffmannella Muller, 1912
  • Huetheriella Jeannel, 1934
  • Hussonella Jeannel, 1934
  • Icharonia Reitter, 1912
  • Insubriella Vailati, 1990
  • Iranobathyscia Zoia & Rampini, 1994
  • Isereus Reitter, 1886
  • Josettekia Belles & Deliot, 1983
  • Karadeniziella Casale & Giachino, 1989
  • Katobatizon Knirsch, 1928
  • Kircheria Giachino & Vailati, 2006
  • Lagariella Fresneda, 2000
  • Laneyriella Gueorguiev, 1976
  • Leonesiella Salgado, 1996
  • Leonhardella Reitter, 1903
  • Leonhardia Reitter, 1901
  • Leptodirus Schmidt, 1832
  • Leptomeson Jeannel, 1924
  • Leptostagus Karaman, 1954
  • Lessiniella Pavan, 1941
  • Lotharia Mandl, 1944
  • Magdelainella Jeannel, 1924
  • Maroniella Casale & Giachino, 1985
  • Mehadiella Csiki, 1899
  • Mesocatops Szymczakowski, 1961
  • Mesocolon Broun, 1880
  • Micronemadus Jeannel, 1936
  • Monguzziella Vailati, 1993
  • Muelleriella Jeannel, 1924
  • Nafarroa Fresneda & Dupré, 2010
  • Nargiotes Jeannel, 1936
  • Nargomorphus Jeannel, 1936
  • Nargus Thomson, 1867
  • Naspunius Fresneda, Hernando & Lagar, 1994
  • Nauticiella Moravec & Mlejnek, 2002
  • Nemadiolus Jeannel, 1936
  • Nemadiopsis Jeannel, 1936
  • Nemadotropis Szymczakowski, 1971
  • Nemadus Thomson, 1867
  • Neobathyscia Muller, 1917
  • Neoeocatops Peck & Cook, 2007
  • Netolitzkya Muller, 1913
  • Newtoniopsis Salgado Costas, 2005
  • Nipponemadus Perreau, 2004
  • Nonveilleriella Perreau & Pavicevic, 2008
  • Notidocharis Jeannel, 1956
  • Ochridiola Sbordoni, 1971
  • Oresigenus Jeannel, 1948
  • Oritocatops Jeannel, 1921
  • Orostygia Muller, 1912
  • Oryotus Miller, 1856
  • Ovobathysciola Jeannel, 1924
  • Pallaresiella Fresneda, 1998
  • Pandania Szymczakowski, 1964
  • Pangaeoniola G. & M.Etonti, 1985
  • Parabathyscia Jeannel, 1908
  • Paracatops Portevin, 1907
  • Paranemadus Zwick, 1979
  • Paranillochlamys Zariquiey, 1940
  • Parantrophilon Noesske, 1914
  • Parapaulipalpina Gnaspini, 1996
  • Parapropus Ganglbauer, 1899
  • Paraspeonomus Coiffait, 1952
  • Paratroglophyes Fourès, 1954
  • Parvospeonomus Bellés & Escolà, 1977
  • Patriziella Jeannel, 1956
  • Paulipalpina Gnaspini & Peck, 1996
  • Pavicevicia Perreau, 2008
  • Peckardia Salgado Costas, 2005
  • Peckena Gnaspini, 1996
  • Perriniella Jeannel, 1910
  • Phacomorphus Jeannel, 1908
  • Phaneropella Jeannel, 1910
  • Philomessor Jeannel, 1936
  • Pholeuodromus Breit, 1913
  • Pholeuon Hampe, 1856
  • Pholeuonella Jeannel, 1910
  • Pholeuonidius Jeannel, 1911
  • Pholeuonopsis Apfelbeck, 1901
  • Pisidiella Jeannel, 1930
  • Platycholeus Horn, 1880
  • Pretneria Muller, 1931
  • Prionochaeta Horn, 1880
  • Proleonhardella Jeannel, 1910
  • Proleptodirina Perkovsky, 1997
  • Proptomaphaginus Szymczakowski, 1969
  • Prospelaeobates Giachino & Etonti, 1996
  • Protobracharthron Reitter, 1889
  • Protopholeuon Jeannel, 1923
  • Pseudobathyscidius Karaman, 1964
  • Pseudoboldoria Ghidini, 1937
  • Pseudonargiotes Salgado Costas, 2005
  • Pseudonemadus Portevin, 1914
  • Pseudospeonomus Comas, Fresneda & Salgado, 2007
  • Ptomaphaginus Portevin, 1914
  • Ptomaphagus Hellwig, 1795
  • Ptomaphaminus Perreau, 2000
  • Purkynella Knirsch, 1926
  • Quaestus Schaufuss, 1861
  • Radevia Knirsch, 1925
  • Radziella Casale & Jalzic, 1988
  • Rangiola Jeannel, 1936
  • Ravasinia Muller, 1922
  • Redensekia Karaman, 1953
  • Reinholdina Giachino & Moravec, 2009
  • Remyella Jeannel, 1931
  • Rhaetiella Giachino & Vailati, 2005
  • Rhodopiola Gueorguiev, 1960
  • Roubaliella Jeannel, 1925
  • Royerella Jeannel, 1910
  • Rozajella Curcic, Brajkovic & Curcic, 2007
  • Rybinskiella Reitter, 1906
  • Salgadoia Fresneda, 1998
  • Sbordoniola Zoia & Rampini, 1994
  • Sciaphyes Jeannel, 1924
  • Sciodrepoides Hatch, 1933
  • Sengletiola Zoia & Rampini, 1994
  • Serbopholeuonopsis Ćurčić & Boškova, 2002
  • Setnikia Breit, 1913
  • Sinobathyscia Perreau, 1999
  • Sinuicollia Piva, 2008
  • Sophrochaeta Reitter, 1884
  • Spelaeobates Muller, 1901
  • Spelaeochlamys Dieck, 1871
  • Spelaeodromus Reitter, 1884
  • Spelaites Apfelbeck, 1907
  • Speocharidius Jeannel, 1919
  • Speocharinus Espaol & Escola, 1977
  • Speodiaetus Jeannel, 1908
  • Speonemadus Jeannel, 1922
  • Speonesiotes Jeannel, 1910
  • Speonomidius Jeannel, 1924
  • Speonomites Jeannel, 1910
  • Speonomus Jeannel, 1908
  • Speophyes Jeannel, 1910
  • Speoplanes Muller, 1911
  • Sphaerobathyscia Muller, 1917
  • Stygiophyes Fresneda, 1998
  • Takobiella Ruzicka, 1992
  • Tartariella Nonveiller & Pavicevic, 1999
  • Tismanella Jeannel, 1928
  • Trapezodirus Jeannel, 1924
  • Trocharanis Reitter, 1884
  • Troglocharinus Reitter, 1908
  • Troglodromus Sainte-Claire Deville, 1901
  • Troglophyes Abeille de Perrin, 1894
  • Velebitodromus Casale, Giachino & Jalžic, 2004
  • Viallia Pavan, 1950
  • Weiratheria Zariquiey, 1927
  • Zariquieyella Jeannel, 1928
  • Aranzadiella Espaol, 1972
  • Petkovskiella Gueorguiev, 1976


References