Biology:Neosteneosaurus

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Neosteneosaurus is a genus of machimosaurid, known from the Middle Jurassic Oxford Clay of the UK, and Marnes de Dives, France.

History

Skeleton at Peterborough Museum

The type species, N. edwardsi, was originally named as a species of Steneosaurus in 1868 as "Steneosaurus" edwardsi in 1868.[1] The genus Steneosaurus was used as a longtime wastebasket taxon for various teleosauroid specimens and had more than a dozen species. The type species, Steneosaurus rostromajor is undiagnostic, making the genus and species dubious and invalid. Additionally, many species of so called "Steneosaurus" were found to be quite different and unrelated to one another, thus needing new generic names.[2].

In 2020, a study by Michela Johnson and colleagues in 2020 reclassified "Steneosaurus" edwardsi into the new genus Neosteneosaurus. The species "Steneosaurus" durobrivensis and "Steneosaurus" hulkei were also found to be junior synonyms of N. edwardsi.[2]

Classification

Neosteneosaurus is a member of the machimosaurid subfamily Machimosaurinae. It is closely related to other large-bodied members such as Lemmysuchus and Machimosaurus.

The cladogram below is from an analysis by Johnson and colleagues in 2020.[2]

Machimosauridae

Macrospondylus bollensis

Clovesuurdameredeor stephani

Charitomenosuchus leedsi

Seldsienean megistorhynchus

Deslongchampsina larteti

Machimosaurinae

Proexochokefalos hebertii

cf. Proexochokefalos bouchardi

Neosteneosaurus edwardsi

Andrianavoay baroni

Machimosaurini

Lemmysuchus obtusidens

Yvridiosuchus boutilieri

Machimosaurus buffetauti

Machimosaurus mosae

Machimosaurus rex

Machimosaurus hugii

Ecology

In life, Neosteneosaurus was a large and robust predator of large-bodied marine prey. It had well suited for tackling large prey, with large teeth and some of the largest muscle attachment sites among machimosaurids.[3] However, this animal was still not at the level of more derived and more durophagous machimosaurids such as Machimosaurus. Neosteneosaurus was part of a shift of machimosaurids adapting to larger prey, functioning as a generalist predator of large fish and marine reptiles.[3]

In 2015, it was estimated at more than 7 m (23 ft) in length.[4] In 2016, this estimate was revised down to 6.6 m (22 ft), but even with such measurement, this animal remains to be the largest known Middle Jurassic crocodylomorph.[5] This would make Neosteneosaurus amongst the top of the food chain in its ecosystem.

References

  1. Eudes-Deslongchamps E. 1867-1869. Notes Paléontologiques. Caen and Paris: 320-392.
  2. 2.0 2.1 2.2 Johnson, Michela M.; Young, Mark T.; Brusatte, Stephen L. (2020). "The phylogenetics of Teleosauroidea (Crocodylomorpha, Thalattosuchia) and implications for their ecology and evolution". PeerJ 8. doi:10.7717/peerj.9808. PMID 33083104. 
  3. 3.0 3.1 Johnson, M. M.; Foffa, D.; Young, M. T.; Brusatte, S. L. (2022). "The ecological diversification and evolution of Teleosauroidea (Crocodylomorpha, Thalattosuchia), with insights into their mandibular biomechanics". Ecology and Evolution 12 (11). doi:10.1002/ece3.9484. PMID 36415878. Bibcode2022EcoEv..12E9484J. 
  4. Michela M Johnson, Mark Thomas Young, Lorna Steel, Yves Lepage (July 2015). "Steneosaurus edwardsi (Thalattosuchia: Teleosauridae), the largest known crocodylomorph of the Middle Jurassic". Biological Journal of the Linnean Society 115 (4): 911–918. doi:10.1111/bij.12525. https://www.researchgate.net/publication/280098383. 
  5. Young, MT; Rabi, M.; Bell, MA; Foffa, D.; Steel, L.; Sachs, S.; Peyer, K. (2016). "Big-headed marine crocodyliforms and why we must be cautious when using extant species as body length proxies for long-extinct relatives". Palaeontologia Electronica 19 (3): 1–14. doi:10.26879/648. http://palaeo-electronica.org/content/2016/1554-teleosaurid-size-estimation. 

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