Biology:Procolophonidae

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Procolophonidae is an extinct family of small, lizard-like parareptiles known from the Late Permian to Late Triassic that were distributed across Pangaea, having been reported from Europe, North America, China, South Africa, South America, Antarctica and Australia. The most primitive procolophonids were likely insectivorous or omnivorous, more derived members of the clade developed bicusped molars, and were likely herbivorous feeding on high fiber vegetation or durophagous omnivores.[1] Many members of the group are noted for spines projecting from the quadratojugal bone of the skull, which likely served a defensive purpose as well as possibly also for display.[2] At least some taxa were likely fossorial burrowers.[3] While diverse during the Early and Middle Triassic, they had very low diversity during the Late Triassic, and were extinct by the beginning of the Jurassic.[4]

Phylogeny

The family is defined as all taxa more closely related to Procolophon trigoniceps than to Owenetta rubidgei.[5] Below is a cladogram from Ruta et al. (2011):[6]

Procolophonidae

Coletta seca

Pintosaurus magnidentis

Sauropareion anoplus

Kitchingnathus untabeni

Phaanthosaurus ignatjevi

Phaanthosaurus simus

Theledectinae

Eumetabolodon dongshengensis

Theledectes perforatus

Tichvinskia vjatkensis

Leptopleuroninae

Pentaedrusaurus ordosianus

Neoprocolophon asiaticus

Sclerosaurus armatus

Scoloparia glyphanodon

Leptopleuron lacertinum

Soturnia caliodon

Hypsognathus fenneri

Procolophoninae

Eumetabolodon bathycephalus

Procolophon trigoniceps

Teratophon spinigenis

Thelerpeton oppressus

Timanophon raridentatus

Thelephon contritus

Anomoiodon liliensterni

Kapes amaenus

Kapes bentoni

Kapes komiensis

Kapes majmesculae

Below are three cladograms that follow phylogenetic analyses by Butler et al. (2023). Analysis 1: Strict consensus of 760 most parsimonious trees (MPTs):[7]

Procolophonidae

Eomurruna

Procolina

Anomoiodon

Lasasaurus

Kitchingnathus

Phonodus

Thelephon

Kapes bentoni

Kapes majmesculae

Timanophon

Tichvinskia

Theledectes

Eumetabolodon dongshengensis

Phaantosaurus (sic)

Sauropareion

Pintosaurus

Coletta

Eumetabolodon bathycephalus

Procolophoninae

Teratophon

Thelerpeton

Procolophon

Leptopleuroninae

Neoprocolophon

Pentaedrusaurus

Mandaphon

Scoloparia

Sclerosaurus

Hwiccewyrm

Hypsognathus

Soturnia

Leptopleuron

Analysis 2: Single MPT:[7]

Procolophonidae

Coletta

Lasasaurus

Sauropareion

Kitchingnathus

Pintosaurus

Phaantosaurus (sic)

Eomurruna

Eumetabolodon dongshengensis

Timanophon

Tichvinskia

Procolophoninae

Eumetabolodon bathycephalus

Thelerpeton

Teratophon

Procolophon

Leptopleuroninae

Kapes bentoni

Thelephon

Kapes majmesculae

Pentaedrusaurus

Anomoiodon

Theledectes

Procolina

Phonodus

Mandaphon

Neoprocolophon

Scoloparia

Sclerosaurus

Hwiccewyrm

Leptopleuron

Hypsognathus

Soturnia

Analysis 3: Strict consensus of 18 MPTs:[7]

Procolophonidae

Coletta

Sauropareion

Kitchingnathus

Phaantosaurus (sic)

Eumetabolodon dongshengensis

Eomurruna

Theledectes

Procolina

Phonodus

Lasasaurus

Anomoiodon

Thelephon

Kapes bentoni

Kapes majmesculae

Timanophon

Tichvinskia

Eumetabolodon bathycephalus

Procolophoninae

Teratophon

Thelerpeton

Procolophon

Leptopleuroninae

Neoprocolophon

Pentaedrusaurus

Mandaphon

Scoloparia

Sclerosaurus

Hwiccewyrm

Hypsognathus

Soturnia

Leptopleuron

References

  1. Pinheiro, Felipe L.; Silva-Neves, Eduardo; Da-Rosa, Átila A. S. (August 2021). Ruta, Marcello. ed. "An early-diverging procolophonid from the lowermost Triassic of South America and the origins of herbivory in Procolophonoidea" (in en). Papers in Palaeontology 7 (3): 1601–1612. doi:10.1002/spp2.1355. ISSN 2056-2799. Bibcode2021PPal....7.1601P. https://onlinelibrary.wiley.com/doi/10.1002/spp2.1355. Retrieved 2022-06-20. 
  2. Zaher, Marta; Coram, Robert A.; Benton, Michael J. (February 2019). Angielczyk, Kenneth. ed. "The Middle Triassic procolophonid Kapes bentoni: computed tomography of the skull and skeleton" (in en). Papers in Palaeontology 5 (1): 111–138. doi:10.1002/spp2.1232. Bibcode2019PPal....5..111Z. 
  3. Botha-Brink, Jennifer; Smith, Roger Malcolm Harris (September 2012). "Palaeobiology of Triassic procolophonids, inferred from bone microstructure" (in en). Comptes Rendus Palevol 11 (6): 419–433. doi:10.1016/j.crpv.2012.03.002. Bibcode2012CRPal..11..419B. https://linkinghub.elsevier.com/retrieve/pii/S1631068312000735. 
  4. MacDougall, Mark J.; Brocklehurst, Neil; Fröbisch, Jörg (2019-03-20). "Species richness and disparity of parareptiles across the end-Permian mass extinction". Proceedings of the Royal Society B: Biological Sciences 286 (1899). doi:10.1098/rspb.2018.2572. ISSN 0962-8452. PMID 30890099. 
  5. Cite error: Invalid <ref> tag; no text was provided for refs named CJC08
  6. Ruta, M.; Cisneros, J. C.; Liebrecht, T.; Tsuji, L. A.; Müller, J. (2011). "Amniotes through major biological crises: Faunal turnover among Parareptiles and the end-Permian mass extinction". Palaeontology 54 (5): 1117–1137. doi:10.1111/j.1475-4983.2011.01051.x. Bibcode2011Palgy..54.1117R. 
  7. 7.0 7.1 7.2 Butler, R. J.; Meade, L. E.; Cleary, T. J.; McWhirter, K. T.; Brown, E. E.; Kemp, T. S.; Benito, J.; Fraser, N. C. (2023). "Hwiccewyrm trispiculum gen. et sp. nov., a new leptopleuronine procolophonid from the Late Triassic of southwest England". The Anatomical Record. doi:10.1002/ar.25316. PMID 37735997. 

Sources

Template:Parareptilia Wikidata ☰ Q133347 entry