Biology:Tylototriton

From HandWiki

Tylototriton is a genus of newts commonly known as crocodile newts or knobby newts. About 38 known species are in this genus, with many species having been described recently.[1][2][3] They range from northeastern India and Nepal through Burma to northern Thailand, Laos, Vietnam, and southern China.[2]

Species

This genus currently contains around 40 described species as of April 2026:[4][5]

  • Tylototriton anguliceps Le, Nguyen, Nishikawa, Nguyen, Pham, Matsui, Bernardes, and Nguyen, 2015
  • Tylototriton anhuiensis Qian, Sun, Li, Guo, Pan, Kang, Wang, Jiang, Wu, and Zhang, 2017
  • Tylototriton asperrimus Unterstein, 1930
  • Tylototriton broadoridgus Shen, Jiang, and Mo, 2012
  • Tylototriton dabienicus Chen, Wang, and Tao, 2010
  • Tylototriton daloushanensis Zhou, Xiao, and Luo, 2022
  • Tylototriton gaowangjienensis ({{{2}}}) Huang, Xiang, Wu, Zhang, Zhang, Wang, Lan, Huang, Jiang, and Jiang, 2024
  • Tylototriton guilinensis Xiao et al., 2026[6]
  • Tylototriton hainanensis Fei, Ye, and Yang, 1984
  • Tylototriton himalayanus Khatiwada, Wang, Ghimire, Vasudevan, Paudel, and Jiang, 2015
  • Tylototriton kachinorum Zaw, Lay, Pawangkhanant, Gorin, and Poyarkov, 2019
  • Tylototriton kweichowensis Fang and Chang, 1932
  • Tylototriton liuyangensis Yang, Jiang, Shen, and Fei, 2014
  • Tylototriton lizhengchangi Hou, Zhang, Jiang, Li and Lu, 2012
  • Tylototriton maolanensis Li, Wei, Cheng, Zhang, and Wang, 2020
  • Tylototriton ngarsuensis Grismer, Wood, Quah, Thura, Espinoza, Grismer, Murdoch, and Lin, 2018
  • Tylototriton ngoclinhensis Phung, Pham, Nguyen, Ninh, Nguyen, Bernardes, Le, Ziegler & Nguyen, 2023
  • Tylototriton notialis Stuart, Phimmachak, Sivongxay, and Robichaud, 2010
  • Tylototriton panhai Nishikawa, Khonsue, Pomchote, and Matsui, 2013
  • Tylototriton panwaensis Grismer, Wood, Quah, Thura, Espinoza, and Murdoch, 2019
  • Tylototriton pasmansi Bernardes, Le, Nguyen, Pham, Pham, Nguyen, and Ziegler, 2020
  • Tylototriton phukhaensis Pomchote, Khonsue, Thammachoti, Hernandez, Peerachidacho, Suwannapoom, Onishi, and Nishikawa, 2020
  • Tylototriton podichthys Phimmachak, Aowphol, and Stuart, 2015
  • Tylototriton pseudoverrucosus Hou, Gu, Zhang, Zeng, and Lu, 2012
  • Tylototriton pulcherrimus Hou, Zhang, Li, and Lu, 2012
  • Tylototriton shanjing Nussbaum, Brodie, and Yang, 1995
  • Tylototriton shanorum Nishikawa, Matsui, and Rao, 2014
  • Tylototriton sini Lyu, Wang, Zeng, Zhou, Qi, Wan, and Wang, 2021
  • Tylototriton sparreboomi Bernardes, Le, Nguyen, Pham, Pham, Nguyen, and Ziegler, 2020
  • Tylototriton taliangensis Liu, 1950
  • Tylototriton thaiorum Poyarkov, Nguyen, and Arkhipov, 2021
  • Tylototriton tongziensis Li, Liu, Shi, Wei, and Wang, 2022
  • Tylototriton umphangensis Pomchote, Peerachidacho, Hernandez, Sapewisut, Khonsue, Thammachoti, and Nishikawa, 2021
  • Tylototriton uyenoi Nishikawa, Khonsue, Pomchote, and Matsui, 2013
  • Tylototriton verrucosus Anderson, 1871
  • Tylototriton vietnamensis Böhme, Schöttler, Nguyen, and Köhler, 2005
  • Tylototriton vietnamirabilis Ong AV, Phan TQ, Hoang CV, Nguyen MHT, Nguyen TT, Ziegler T, Nguyen TQ, Pham CT, 2026
  • Tylototriton wenxianensis Fei, Ye, and Yang, 1984
  • Tylototriton wufengensis Li et al., 2026[7]
  • Tylototriton yangi Hou, Zhang, Zhou, Li, and Lu, 2012
  • Tylototriton zaimeng Decemson, Lalremsanga, Elangbam, Vabeiryureilai, Shinde, Purkayastha, Arkhipov, Bragin & Poyarkov, 2023
  • Tylototriton ziegleri Nishikawa, Matsui, and Nguyen, 2013

Phylogeny

The following phylogeny of Tylototriton is from Nishikawa, Matsui & Rao (2014).[8]

Tylototriton
Subgenus Yaotriton
 

Tylototriton panhai

 
 

Tylototriton asperrimus

 
 
Subgenus Tylototriton
 

Tylototriton taliangensis

 
 
 

Tylototriton kweichowensis

 
 
 
 

Tylototriton verrucosus verrucosus (Nepal)

 
 

Tylototriton verrucosus verrucosus (Myanmar)

 
 
 
 

Tylototriton yangi

 
 

Tylototriton uyenoi

 
T. shanjing complex
 

Tylototriton verrucosus pulcherrima

 
 
 

Tylototriton shanjing

 
 

Tylototriton verrucosus verrucosus

 
 
 
 
 
 
 
 

References

  1. ↑ Max Sparreboom (2013). "Tylototriton Anderson, 1871". Salamanders of the Old World. http://science.naturalis.nl/hosted-sites/salamanders/salamanders-of-the-old-world/species-list/tylototriton. 
  2. ↑ 2.0 2.1 Frost, Darrel R. (2021). "Tylototriton Anderson, 1871". Amphibian Species of the World 6.1, an Online Reference. American Museum of Natural History. http://research.amnh.org/vz/herpetology/amphibia/Amphibia/Caudata/Salamandridae/Pleurodelinae/Tylototriton. 
  3. ↑ Nishikawa, Kanto; Masafumi Matsui; Tao Thien Nguyen (2013). "A new species of Tylototriton from northern Vietnam (Amphibia: Urodela: Salamandridae)". Current Herpetology 32: 34–49. doi:10.5358/hsj.32.34. http://www.bioone.org/doi/abs/10.5358/hsj.32.34. 
  4. ↑ Phung, Trung My; Pham, Cuong The; Nguyen, Truong Quang; Ninh, Hoa Thi; Nguyen, Huy Quoc; Bernardes, Marta; Le, Son Thanh; Ziegler, Thomas et al. (2023-07-03). "Southbound – the southernmost record of Tylototriton (Amphibia, Caudata, Salamandridae) from the Central Highlands of Vietnam represents a new species". ZooKeys (1168): 193–218. doi:10.3897/zookeys.1168.96091. ISSN 1313-2970. PMID 38328626. PMC 10848839. Bibcode: 2023ZooK.1168..193P. https://zookeys.pensoft.net/article/96091/. 
  5. ↑ Ong, An Vinh; Phan, Tien Quang; Hoang, Chung Van; Nguyen, Mai Hong Thi; Nguyen, Tao Thien; Ziegler, Thomas; Nguyen, Truong Quang; Pham, Cuong The (2026-04-08). "A new species of Tylototriton (Amphibia, Salamandridae) from Phu Xai Lai Leng Mountain, Nghe An Province, Vietnam" (in en). ZooKeys (1276): 285–305. doi:10.3897/zookeys.1276.173848. ISSN 1313-2970. PMID 42004729. Bibcode: 2026ZooK.1276..285O. 
  6. ↑ Xiao; Chen; Wu; Pan; Chen; Yu; Huang (2026). "A new species of genus Tylototriton (Caudata, Salamandridae) from northeastern Guangxi, China". Zoosystematics and Evolution 102 (2): 671–686. doi:10.3897/zse.102.184792. Bibcode: 2026ZooEv.102..671X. 
  7. ↑ Li, Shize; Shi, Shengchao; Liu, Jing; Luo, Zhonghao; Wang, Jiajun; Liao, Linhong; Wang, Yeqing; Gong, Renhu et al. (2026). "Description of a new species of the Asian newt genus Tylototriton Anderson, 1871 (Urodela, Salamandridae) from central China" (in en). Zoosystematics and Evolution 102 (1): 181–197. doi:10.3897/zse.102.173283. 
  8. ↑ Nishikawa, Matsui & Rao, 2014 : A new species of Tylototriton (Amphibia: Urodela: Salamandridae) from central Myanmar. Natural History Bulletin of the Siam Society, vol. 60, no. 1, p. 9–22.

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