The Pinjor Formation is a Cenozoic geologic formation in India. The fossils of mammals are abundant from the formation, with the addition of reptiles, fishes and invertebrates.[2]
Paleobiota
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Color key
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Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
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Reptiles
Pseudosuchians
It is possible that Rhamphosuchus crassidens and probably Hypselornis sivalensis may have been present in Pinjor Formation.[3]
Turtle
| Turtles from the Pinjor Formation
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| Genus |
Species |
Material |
Location |
Notes |
Images
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| Megalochelys[4]
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M. atlas
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Numerous specimens, including ones referred to M. sivalensis.
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Known from Plio-Pleistocene deposits of entire Punjab.
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A large sized testudinid.
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| cf. Geoclemys[4]
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cf. G. hamiltonii
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WIF/A 454, partially preserved carapace and antero-posteriorly broken plastron.
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Ambala
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A geoemydid.
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| Omegachelys[4]
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O. sahnii
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WIF/A 451, terminally broken large carapace.
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Ambala
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A geoemydid.
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Squamates
Birds
It is possible that Leptoptilos falconeri and Struthio asiaticus might have been present in Pinjor Formation.[6][7]
| Birds from the Pinjor Formation
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| Genus |
Species |
Material |
Location |
Notes |
Images
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| Aves[8]
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indet.
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Potwar Plateau
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Indeterminate aves.
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| Struthio
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S. camelus
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eggshells fragments
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Chandigarh region
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A struthionid.
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Mammals
Eulipotyphla
| Eulipotyphlas from the Pinjor Formation
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| Genus |
Species |
Material |
Location |
Notes |
Images
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| Chandisorex[9]
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C. punchkulaensis
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Chandigarh region
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A soricid.
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Rodent
| Rodents from the Pinjor Formation
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| Genus |
Species |
Material |
Location |
Notes |
Images
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| Hystrix[9]
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H. leucurus
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Chandigarh region
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A hystricid.
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| Rattus[9]
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R. sp.
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Chandigarh region
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A murid.
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| Mus[9]
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M. linnaeusi
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Chandigarh region
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A murid.
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| cf. M. flynni
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Chandigarh region
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A murid.
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| Hadromys[9]
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H. loujacobsi
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Chandigarh region
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A murid
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| H. sp.
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Chandigarh region
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A murid
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| Nesokia[9]
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N. punchkulaensis
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Chandigarh region
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A murid
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| Cremnomys[9]
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C. blanfordi
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Chandigarh region
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A murid
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| Dilatomys[9]
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D. sp.
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Chandigarh region
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A murid
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| Golunda[9]
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G. sp.
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Chandigarh region
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A murid
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| Anepsirhizomys[9]
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A. pinjoricus
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Chandigarh region
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A murid
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| Tatera[9]
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T. pinjoricus
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Chandigarh region
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A murid
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| Bandicota[9]
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B. sp
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Chandigarh region
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A murid
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Lagomorph
| Lagomorphs from the Pinjor Formation
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| Genus |
Species |
Material |
Location |
Notes |
Images
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| Pliosiwalagus[9][10]
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P. sivalensis
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Chandigarh region
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A leporid
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Primates
It is possible that Hominins (most likely Archaic humans) were present in Pinjor Formation.[1]
| Primates from the Pinjor Formation
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| Genus |
Species |
Material |
Location |
Notes |
Images
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| Theropithecus[9]
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T. oswaldi[11]
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Chandigarh region
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A cercopithecid.
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| Procynocephalus[9]
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P. subhimalayanus
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Chandigarh region
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A cercopithecid.
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Felids
Canids
Ursids
| Ursid from the Pinjor Formation
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| Genus |
Species |
Material |
Location |
Notes |
Images
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| Agriotherium[15]
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A. sivalensis
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Kasamuri Rao, Saharanpur of North-Western Provinces and Siwalik Ganges valley neighbourhood.[17][18]
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A ailuropodine ursid.
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| Melursus[15]
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M. theobaldi
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Kangra district[17]
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A ursine ursid.
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Viverrids
| Viverrids from the Pinjor Formation
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| Genus |
Species |
Material |
Location |
Notes |
Images
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| Vishnuictis[15]
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V. durandi
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Upper Siwalik
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A viverrine viverrid.
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| V. sp.[19]
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Chandigarh region
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A viverrine viverrid.
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| Viverra[15]
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V. bakeri
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Upper Siwalik
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A viverrine viverrid.
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Mustelids
| Mustelids from the Pinjor Formation
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| Genus |
Species |
Material |
Location |
Notes |
Images
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| Enhydriodon[15]
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E. sivalensis
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Chandigarh region, Potwar Plateau
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A lutrine mustelid.
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| Sinictis[15]
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S. lydekkeri
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Upper Siwaliks
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A mustelid
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| Amblonyx[15]
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A. indicus
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Upper Siwaliks
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A lutrine mustelid.
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| A. sp.
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Upper Siwaliks
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A lutrine mustelid.
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| Mellivora
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M. sivalensis[9]
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Chandigarh region
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A mellivorine mustelid.
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| Lutra[15]
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L. palaeindica
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Upper Siwaliks
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A lutrine mustelid.
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Hyaenids
Proboscideans
| Proboscideans from the Pinjor Formation
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| Genus |
Species |
Material |
Location |
Notes |
Images
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| Stegodon[9]
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S. pinjorensis
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Chandigarh region
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A stegodontid proboscidean.
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| S. insignis
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Chandigarh region
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A stegodontid proboscidean.
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| S. sp.
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Chandigarh region
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A stegodontid proboscidean.
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| S. bombifrons
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Potwar Plateau
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A stegodontid proboscidean, also known from the older Tatrot Formation.
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| Elephas[9]
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E. hysudricus
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Chandigarh region
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An elephantid proboscidean.
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| E. planifrons
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Chandigarh region
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An elephantid proboscidean, also known from the older Tatrot Formation.
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| E. platycephalus
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Chandigarh region
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An elephantid proboscidean.
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| Anancus[9]
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A. sivalensis
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Chandigarh region
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A gomphothere proboscidean, also known from the older Tatrot Formation.
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| Stegolophodon[9]
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S. stegodontoides
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Chandigarh region
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A stegodontid proboscidean, also known from the older Tatrot Formation.
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Rhinocerotids
| Rhinocerotids from the Pinjor Formation
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| Genus |
Species |
Material |
Location |
Notes |
Images
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| Rhinoceros[9]
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R. sivalensis
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Chandigarh region
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A rhinocerotine rhinocerotid.
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| R. platyrhinus
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Chandigarh region
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A large rhinocerotine rhinocerotid, includes Punjabitherium.
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| R. palaeindicus
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Chandigarh region
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A rhinocerotine rhinocerotid.
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| Subchilotherium[9]
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S. intermedium
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Chandigarh region
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A aceratheriine rhinocerotid. Also known from the older Tatrot Formation.
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Bovids
Cervids
| Cervids from the Pinjor Formation
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| Genus |
Species |
Material |
Location |
Notes |
Images
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| Cervus
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C. sivalensis
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A large-size cervid.
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| C. triplidens
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A cervid.
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| C. sp.[1]
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A cervid.
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| Metacervocerus
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M. punjabiensis
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A cervid.
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| Eucladoceros
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E. sp.
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A large cervid.
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| Praesinomegaceros
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P. sp.
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A large cervid.
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| Rucervus[1]
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R. sp.
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A cervid.
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Giraffids
Suids
Other Ungulates
References
- ↑ 1.00 1.01 1.02 1.03 1.04 1.05 1.06 1.07 1.08 1.09 1.10 1.11 1.12 1.13 1.14 1.15 1.16 1.17 1.18 1.19 Kaur, A. P.; Saarinen, J.; Patnaik, R.; Chauhan, P. R.; Dennell, R. (2026). "Contextualizing Early to Middle Pleistocene hominin ecology in the Indian Subcontinent". Journal of Quaternary Science. doi:10.1002/jqs.70088.
- ↑ V. Kumaravel, S. J. Sangode; Rohtash Kumar, N. Siva Siddaiah (2005). "Magnetic polarity stratigraphy of Plio–Pleistocene Pinjor Formation (type locality), Siwalik Group, NW Himalaya, India". Zoological Journal of the Linnean Society. doi:10.1093/zoolinnean/zlad184. https://www.jstor.org/stable/24110713.
- ↑ Martin, J. E. (2019). "The taxonomic content of the genus Gavialis from the Siwalik Hills of India and Pakistan" (in en). Papers in Palaeontology 5 (3): 483–497. doi:10.1002/spp2.1247. Bibcode: 2019PPal....5..483M. https://hal.archives-ouvertes.fr/hal-02123647/file/2019Gavialis.pdf.
- ↑ 4.0 4.1 4.2 4.3 4.4 Srivastava, R., and H. H. Schleich. (2018). "Fossil Turtles and Crocodiles from the Caenozoic of S-Asia". ARCO-Nepal, Muhich, 1-324
- ↑ Chabrol, Nils; Jukar, Advait M.; Patnaik, Rajeev; Mannion, Philip D. (21 March 2024). "Osteology of Crocodylus palaeindicus from the late Miocene-Pleistocene of South Asia and the phylogenetic relationships of crocodyloids". Journal of Systematic Palaeontology 24 (1). doi:10.1080/14772019.2024.2313133.
- ↑ Buffetaut, E. (2022). "The First-Named Fossil Ostrich: A Revision of Struthio asiaticus, from the Siwaliks of India". Diversity 14 (10): 860. doi:10.3390/d14100860.
- ↑ Louchart, A., Vignaud, P., Likius, A., Brunet, M., & White, T. D. (2005). A large extinct marabou stork in African Pliocene hominid sites, and a review of the fossil species of Leptoptilos. Acta Palaeontologica Polonica, 50(3).
- ↑ 8.0 8.1 8.2 8.3 8.4 Dennell, R. W.; Coard, R.; Turner, A. (2008). "Predators and scavengers in Early Pleistocene southern Asia". Quaternary International. 192:78-88.
- ↑ 9.00 9.01 9.02 9.03 9.04 9.05 9.06 9.07 9.08 9.09 9.10 9.11 9.12 9.13 9.14 9.15 9.16 9.17 9.18 9.19 9.20 9.21 9.22 9.23 9.24 9.25 9.26 9.27 9.28 "Pinjor Formation". https://indplex.geolex.org/formations/Pinjor%20Fm.
- ↑ Patnaik, R. (2002). "Pliocene Leporidae (Lagomorpha, Mammalia) from the Upper Siwaliks of India: Implications for phylogenetic relationships". Journal of Vertebrate Paleontology 22(2):443–452
- ↑ Getahun, Dagmawit Abebe; Delson, Eric; Seyoum, Chalachew Mesfin (July 2023). "A review of Theropithecus oswaldi with the proposal of a new subspecies" (in en). Journal of Human Evolution 180. doi:10.1016/j.jhevol.2023.103373. https://www.sciencedirect.com/science/article/pii/S0047248423000520. Retrieved 5 March 2026.
- ↑ 12.0 12.1 Stimpson, Christopher M. (May 2024). "Siwalik sabrecats: review and revised diagnosis of Megantereon fossils from the foothills of the Himalaya" (in en). Royal Society Open Science 11 (5). doi:10.1098/rsos.231788. ISSN 2054-5703. PMID 38720790. Bibcode: 2024RSOS...1131788S.
- ↑ Werdelin, Lars; Lewis, Margaret E. (2001), "A revision of the genus Dinofelis (Mammalia, Felidae)", Zoological Journal of the Linnean Society 132 (2): 147–258, doi:10.1006/zjls.2000.0260
- ↑ Jiangzuo, Q.; Liu, J. (2020). "First record of the Eurasian jaguar in southern Asia and a review of dental differences between pantherine cats". Journal of Quaternary Science 35 (6): 817–830. doi:10.1002/jqs.3222. Bibcode: 2020JQS....35..817J.
- ↑ 15.00 15.01 15.02 15.03 15.04 15.05 15.06 15.07 15.08 15.09 15.10 Jasinski, Steven E.; Abbas, Sayyed Ghyour; Mahmood, Khalid; Babar, Muhammad Adeeb; Khan, Muhammad Akbar (2023-11-02). "New Carnivoran (Mammalia: Carnivora) specimens from the Siwaliks of Pakistan and India and their faunal and evolutionary implications". Historical Biology. 35 (11): 2217–2252.
- ↑ Garrido, Guiomar; Arribas, Alfonso (2008). "Canis accitanus nov. sp., a new small dog (Canidae, Carnivora, Mammalia) from the Fonelas P-1 Plio-Pleistocene site (Guadix basin, Granada, Spain)". Geobios 41 (8): 753. doi:10.1016/j.geobios.2008.05.002. Bibcode: 2008Geobi..41..751G.
- ↑ 17.0 17.1 17.2 Lydekker, R. (1884). "Siwalik and Narbada Carnivora". Paleontologia Indica, 10:178-135.
- ↑ Lydekker, R. (1888). "Notes on Indian fossil Vertebrata; the ulna of Hyaenarctos". Records of Geological Survey of India, 21:145-146.
- ↑ Gaur, R.; Chopra, S. (22 March 1984). "Taphonomy, fauna, environment and ecology of Upper Siwaliks (Plio-Pleistocene) near Chandigarh, India" (in en). Nature 308: 353–355. doi:10.1038/308353a0.
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