Biology:Arganaceras
Arganaceras ("Argana horn") is a medium-sized pareiasaur from the Late Permian Ikakern Formation of Morocco. It was about 2 metres (6.6 ft) in length and had a horn-like structure on its snout.[1]
Description
The holotype and only described specimen of Arganaceras (ARG 518) is a reasonably complete, disarticulated skull. Several features of the skull were interpreted as indicative of the holotype being immature when it died, such as a lack of fusion between the quadrate and elements of the cranial vault, as well as some unossified elements. The authors postulate that the cranial ornamentation seen in the holotype would likely be more prominent in an adult individual. Arganaceras is described as a large pareiasaur, distinguished from its relatives by eleven autapomorphic features. It possessed horn-like protuberances on the squamosal and quadratojugal, which are compared to similar ornamentation in Elginia. A noteworthy feature of Arganaceras' skull is the conical bony protuberance present on the nasal midline; other pareiasaurs, including the aforementioned Elginia as well as Pareiasuchus, Scutosaurus, and Bunostegos also possess nasal protuberances. The nasal protuberance of Arganaceras is most similar to that of Elginia and Pareiasuchus, being located in a medial position along the nasal midline, whereas it is located laterally in Scutosaurus and Bunostegos. The premaxillary teeth of Arganaceras are procumbent, and the dentition is described as pleurothecodont, which the authors note is a widespread tooth morphology among herbivorous land animals.[1]
Palaeoecology
The Ikakern Formation is part of the Argana Group, which also features fossiliferous formations that are dated to the Triassic period. Very few tetrapod remains have been found in the Ikakern Formation. Aside from Arganaceras, only a diplocaulid similar to Diploceraspis[2] and the captorhinid Acrodenta have been described. Despite this, ichnofossils are moderately diverse, ranging from tetrapod footprints, to arthropod burrows, to arthropod feeding traces.[3][4]
Phylogeny
In 2025, Jian Yi and Jun Liu described a new pareiasaur, Yinshanosaurus. The paper featured a phylogenetic analysis focused on the inter-relationships within pareiasauria; the results are displayed in the cladogram below. They found Arganaceras to be most closely related to Elginia and Obirkovia. Together, the three genera are found to represent a novel clade within pareiasauria, dubbed elginiidae.[5]
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References
- ↑ 1.0 1.1 Jalil, Nour-Eddine; Janvier, Phillipe (2005). "The Late Permian pareiasaurs (Amniota, Parareptilia) from the Argana Basin, Morocco". Geodiversitas 27 (1): 35–123. https://www.researchgate.net/publication/256238883.
- ↑ Germain, Damien (2010). "The Moroccan diplocaulid: the last lepospondyl, the single one on Gondwana". Historical Biology 22 (1-3): 4–39. doi:10.1080/08912961003779678. ISSN 0891-2963. https://doi.org/10.1080/08912961003779678.
- ↑ Hminna, Abdelkbir; Voigt, Sebastian; Saber, Hafid; Schneider, Jörg W.; Hmich, Driss (2012). "On a moderately diverse continental ichnofauna from the Permian Ikakern Formation (Argana Basin, Western High Atlas, Morocco)". Journal of African Earth Sciences 68: 15–23. doi:10.1016/j.jafrearsci.2012.03.011. ISSN 1464-343X. https://doi.org/10.1016/j.jafrearsci.2012.03.011.
- ↑ Voigt, Sebastian; Hminna, Abdelkbir; Saber, Hafid; Schneider, Jörg W.; Klein, Hendrik (2010). "Tetrapod footprints from the uppermost level of the Permian Ikakern Formation (Argana Basin, Western High Atlas, Morocco)". Journal of African Earth Sciences 57 (5): 470–478. doi:10.1016/j.jafrearsci.2009.12.003. ISSN 1464-343X. https://doi.org/10.1016/j.jafrearsci.2009.12.003.
- ↑ Yi, Jian; Liu, Jun (2025). "The tetrapod fauna of the upper Permian Naobaogou Formation of China: a new mid‐sized pareiasaur Yinshanosaurus angustus and its implications for the phylogenetic relationships of pareiasaurs" (in en). Papers in Palaeontology 11 (3). doi:10.1002/spp2.70020. ISSN 2056-2799. https://onlinelibrary.wiley.com/doi/10.1002/spp2.70020.
Template:Parareptilia Wikidata ☰ Q1064715 entry
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