Biology:Bagaraatan

From HandWiki

Bagaraatan (/'ba-ɣa-raa-tan/ meaning 'small' baɣa + 'carnivorous animal, beast of prey' araatan in Mongolian) is a genus of theropod dinosaur from the Late Cretaceous period of what is now the Nemegt Formation of Mongolia. It may have been a small predatory dinosaur measuring around 3 to 4 metres (9.8 to 13.1 ft) in length.

History

1996 skeletal diagram showing holotype and assigned elements that are now known to belong to a different species
Partial mandible

The type species, B. ostromi, was described by Halszka Osmólska in 1996. Initially, the post-cranial (ZPAL MgD-I/108) skeleton had been described as "bird-like", while the skull was noted to exhibit features of several different theropod groups.

The material that warranted this conclusion was later found to be a chimaera of two non-avian dinosaurs, with some of the post-crania (hand bones, left femur, tibiotarsus, and rib) being referred to an indeterminate caenagnathid, possibly Elmisaurus. The material that is considered the holotype, which includes the mandible, axial skeleton, pelvis, and one pedal phalanx, likely indicates that Bagaraatan is an indeterminate tyrannosaurid. Similarities between the material and young specimens of Tyrannosaurus and Tarbosaurus indicate that the holotype represents a juvenile tyrannosaurid, one of the smallest currently known.[1]

Classification

Size compared to a human

Holtz classified Bagaraatan as a basal tyrannosauroid, Coria identified it as a troodontid, and Rauhut placed it in Maniraptora.[2] Mark Loewen et al. placed it in basal Tyrannosauroidea, agreeing with the placement by Holtz.[3]

Below is the cladogram by Loewen et al. in 2013.[3]

Tyrannosauroidea
Proceratosauridae
 
 

Proceratosaurus bradleyi

 
 

Kileskus aristotocus

 
 

Guanlong wucaii

 
 
 
 

Sinotyrannus kazuoensis

 
 
 

Juratyrant langhami

 
 

Stokesosaurus clevelandi

 
 
 
 
 
 

Dilong paradoxus

 
 
 

Eotyrannus lengi

 
 
 

Bagaraatan ostromi

 
 
 

Raptorex kriegsteini

 
 
 

Dryptosaurus aquilunguis

 
 
 
 

Alectrosaurus olseni

 
 

Xiongguanlong baimoensis

 
 
 
 

Appalachiosaurus montgomeriensis

 
 
 
 

Alioramus altai

 
 

Alioramus remotus

 
 
 

Tyrannosauridae

 
 
 
 
 
 
 
 
 
 

In their 2024 reassessment of Bagaraatan, Słowiak, Brusatte & Szczygielski determined that the initial material referred to Bagaraatan is chimaeric. They suggested that the material recognized as the holotype can more confidently be identified as a member of the Tyrannosauridae or a closely related tyrannosauroid. When tested in a phylogenetic analysis, Bagaraatan was recovered in a polytomy with derived tyrannosauroids within the Tyrannosauridae. However, they caution that its exact phylogenetic position remains uncertain since juvenile specimens tend to be recovered in more basal positions than adults of the same species. Their results are shown in the cladogram below:[1]

 
 

Sinotyrannus 85px

 
 

Yutyrannus 85px

 
 

Proceratosaurus

 
  Guanlong
80px
 
  Kileskus
85px
 
 
 

Suskityrannus

 
 
  Timurlengia
80px
 
 
  Xiongguanlong
80px
 
 
 

Dryptosaurus 85px

 
 
 

Raptorex

 
Tyrannosauridae
 

Bagaraatan

 
 

Bistahieversor

 
  Qianzhousaurus
85px
 
  Alioramus altai
75px
 
  Alioramus remotus
85px
 
  Appalachiosaurus
85px
 
  Albertosaurinae
85px
 
 

Tyrannosaurinae 85px

 
 
 
 
 
 
 
 

References

  1. ↑ 1.0 1.1 Słowiak, Justyna; Brusatte, Stephen L; Szczygielski, Tomasz (2024-02-16). "Reassessment of the enigmatic Late Cretaceous theropod dinosaur, Bagaraatan ostromi" (in en). Zoological Journal of the Linnean Society 202 (3). doi:10.1093/zoolinnean/zlad169. ISSN 0024-4082. 
  2. ↑ O. W. M. Rauhut (2003). The interrelationships and evolution of basal theropod dinosaurs. Special Papers in Palaeontology 69: 1-213.
  3. ↑ 3.0 3.1 Loewen, M.A.; Irmis, R.B.; Sertich, J.J.W.; Currie, P. J.; Sampson, S. D. (2013). Evans, David C. ed. "Tyrant Dinosaur Evolution Tracks the Rise and Fall of Late Cretaceous Oceans". PLoS ONE 8 (11). doi:10.1371/journal.pone.0079420. PMID 24223179. Bibcode: 2013PLoSO...879420L. 

Sources

  • Osmolska, H. (1996). "An unusual theropod dinosaur from the Late Cretaceous Nemegt Formation of Mongolia". Acta Palaeontologica Polonica 41; 1-38 [1]

Data related to Bagaraatan at Wikispecies

Wikidata ☰ Q134285 entry