Biology:Cascolus

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Cascolus is an extinct genus of stem-mandibulate known from the Coalbrookdale Formation.

Description

File:Cascolus Diagram.png
Diagrammatic reconstruction of Cascolus ravitis

Cascolus is a long, somewhat vermiform arthropod, roughly 9 millimetres long. It has a head segment containing a head shield, stalked eyes and five pairs of limbs. The first appendages bore 3 long flagella similar to the first appendages of many megacheirans, and the other four were biramous with gnathobases. The thorax had at least 9 segments, each bearing appendages with a long, spinose, gently tapered endopod and a large flap shaped exopod and two smaller, similarly shaped epipods. The 4th and 5th head appendages were similar to the trunk appendages, but smaller and the 4th only had 1 epipod. Two paired rows of tubercles ran along dorsal side of the trunk and there may have been two possibly limbless segments near the posterior. The trunk remains similar in size through tergites 1-4, and then decreases in width onwards into the limbless segments. The posterior end of the body is not known.[1]

A life reconstruction of Cascolus ravitis

Ecology

Cascolus appears to have been a nektobenthic animal, possibly a scavenger.

Etymology

Cascolus was named in honour of Sir David Attenborough. The genus name derives from "castrum" ("stronghold") and "colus" ("dwelling in"), alluding to the Middle or Old English source for the name "Attenborough". The specific name ravitis derives from "Ratae" (the Roman name for Leicester), "vita" ("life") and "commeatis" ("messenger")

Distribution

Cascolus is known from a single specimen from the Silurian Coalbrookdale Formation in England, a diverse Silurian Lagerstatte.

Classification

Cascolus was originally considered as a stem-group phyllocarid.[1] According to phylogenic analysis in Pulsipher et al. (2022), it is considered as stem-mandibulate instead.[2]

Arthropoda
 

†Parioscorpio venator

 
 
 

†Cascolus ravitis

 
 
 

†Tanazios dokeron

 
 
 
 

†Captopodus poschmanni

 
 
 

†Acheronauta stimulapis

 
 

†Thylacocephala

 
 
 
 
 

†Occacaris oviformis

 
 
 
 

Myriapoda

 
 
 

†Ercaicunia multinodosa

 
 
 
 

†Clypecaris pteroidea

 
 
 
 
 

†Waptia fieldensis

 
 

†Perspicaris dictynna

 
 
 
 

†Canadaspis perfecta

 
 
 

†Tokummia katalepsis

 
 

†Branchiocaris pretiosa

 
 
 
 
 
 
 

†Nereocaris exilis

 
 

†Odaraia alata

 
 
 
 

†Euthycarcinoidea

 
†Fuxianhuiida
 

†Fuxianhuia

 
 
 

†Chengjiangocaris

 
 

†Shankouia

 
 
 
 
 
 
 
 
 
 
Pancrustacea
 
 

Argulus

 
 

Ostracoda

 
 
 
 
 

†Lepidocaris

 
 
 

Triops

 
 

Artemia

 
 
 
 
 
 
 

Remipedia

 
 

Cephalocarida

 
 
 

Hexapoda

 
 
 

Multicrustacea

 
 
 
 
 
 
 
 
 
 

See also

  • List of things named after David Attenborough and his works

References

  1. ↑ 1.0 1.1 Siveter, David J. (22 March 2017). "A new crustacean from the Herefordshire (Silurian) Lagerstätte, UK, and its significance in malacostracan evolution". Proceedings of the Royal Society B: Biological Sciences 284 (1851). doi:10.1098/rspb.2017.0279. PMID 28330926. 
  2. ↑ Pulsipher, M. A.; Anderson, E. P.; Wright, L. S.; Kluessendorf, J.; Mikulic, D. G.; Schiffbauer, J. D. (2022). "Description of Acheronauta gen. nov., a possible mandibulate from the Silurian Waukesha Lagerstätte, Wisconsin, USA". Journal of Systematic Palaeontology 20 (1): 2109216. doi:10.1080/14772019.2022.2109216. 

Wikidata ☰ Q107259350 entry