Biology:Caypullisaurus

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Caypullisaurus is an extinct genus of platypterygiine ophthalmosaurid ichthyosaur from the Late Jurassic to the Early Cretaceous (Tithonian and Berriasian stages) of Argentina. Its holotype was collected from the Vaca Muerta Formation of Cerro Lotena, Neuquen, dating to the early Tithonian stage of the Late Jurassic, about 150 million years ago. Caypullisaurus was first named by Marta Fernández in 1997 and the type species is Caypullisaurus bonapartei. It was a large ichthyosaur, measuring about 7 m (23 ft) long.[1][2] The forelimbs of Caypullisaurus contained 10 digits each.[3]

Classification

It is a member of the family Ophthalmosauridae, and closely related to Platypterygius and Brachypterygius.[4] In 2012, Caypullisaurus was found to be most closely related to Athabascasaurus and "Platypterygius" australis, and to nest within the subfamily Platypterygiinae.[5]

Phylogeny

The following cladogram shows a possible phylogenetic position of Caypullisaurus in Ophthalmosauridae according to the analysis performed by Zverkov and Jacobs (2020).[6]

Ophthalmosauria
 
Ophthalmosaurinae
 
 
 
 

Acamptonectes densus

 
 
 

Mollesaurus periallus

 
 
 

Ophthalmosaurus natans

 
 

Ophthalmosaurus icenicus

 
 
 
 
 
 
 
 
 

Gengasaurus nicosiai

 
 
 
 

Nannopterygius yasykovi

 
 
 

Nannopterygius enthekiodon

 
 

Nannopterygius saveljeviensis

 
 

Nannopterygius borealis

 
 
 
 
 
 

Arthropterygius volgensis

 
 
 

Arthropterygius lundi

 
 

Arthropterygius thalassonotus

 
 
 

Arthropterygius hoybergeti

 
 

Arthropterygius chrisorum

 
 
 
 
 
 
 
 
 
Platypterygiinae
 
 

Brachypterygius extremus

 
 

Aegirosaurus leptospondylus

 
 
 

Muiscasaurus catheti

 
 
 

Leninia stellans

 
 
 

Sveltonectes insolitus

 
 
 

Athabascasaurus bitumineus

 
 
 

Platypterygius americanus

 
 
 
 

Acuetzpalin carranzai

 
 

Platypterygius sachicarum

 
 

Caypullisaurus bonapartei

 
 
 
 
 
 

Grendelius mordax

 
 

Grendelius alekseevi

 
 

Grendelius pseudoscythicus

 
 

Grendelius zhuravlevi

 
 
 
 
 
 

Undorosaurus kielanae

 
 
 

Undorosaurus nessovi

 
 

Undorosaurus gorodischensis

 
 
 
 
 
 
 
 

Platypterygius australis

 
 
 

Plutoniosaurus bedengensis

 
 

Simbirskiasaurus birjukovi

 
 

Platypterygius hercynicus

 
 

Sisteronia seeleyi

 
 

Platypterygius platydactylus

 
 

Maiaspondylus lindoei

 
 
 
 
 
 
 
 
 
 
 
 
 
 


See also

References

  1. ↑ Fernández, M.S. (1997). "A new Ichthyosaur from the Tithonian (Late Jurassic) of the Neuquen Basin, Northwestern Patagonia, Argentina". Journal of Paleontology 71 (3): 479–484. doi:10.1017/S0022336000039494. Bibcode: 1997JPal...71..479F. 
  2. ↑ Gasparini, Z.; Fernández, M. (2005). "Jurassic marine reptiles of the Neuquén Basin: records, faunas and their palaeobiogeographic significance". Geological Society, London, Special Publications 252 (1): 279–294. doi:10.1144/GSL.SP.2005.252.01.14. Bibcode: 2005GSLSP.252..279G. 
  3. ↑ Motani, Ryosuke (2005-05-31). "EVOLUTION OF FISH-SHAPED REPTILES (REPTILIA: ICHTHYOPTERYGIA) IN THEIR PHYSICAL ENVIRONMENTS AND CONSTRAINTS" (in en). Annual Review of Earth and Planetary Sciences 33 (1): 395–420. doi:10.1146/annurev.earth.33.092203.122707. ISSN 0084-6597. https://www.annualreviews.org/doi/10.1146/annurev.earth.33.092203.122707. 
  4. ↑ Fernández M. 2007. Redescription and phylogenetic position of Caypullisaurus (Ichthyosauria: Ophthalmosauridae). Journal of Paleontology 81 (2): 368-375.
  5. ↑ Valentin Fischer; Michael W. Maisch; Darren Naish; Ralf Kosma; Jeff Liston; Ulrich Joger; Fritz J. Krüger; Judith Pardo Pérez et al. (2012). "New Ophthalmosaurid Ichthyosaurs from the European Lower Cretaceous Demonstrate Extensive Ichthyosaur Survival across the Jurassic–Cretaceous Boundary". PLOS ONE 7 (1). doi:10.1371/journal.pone.0029234. PMID 22235274. Bibcode: 2012PLoSO...729234F. 
  6. ↑ Nikolay G. Zverkov; Megan L. Jacobs (2021). "Revision of Nannopterygius (Ichthyosauria: Ophthalmosauridae): reappraisal of the 'inaccessible' holotype resolves a taxonomic tangle and reveals an obscure ophthalmosaurid lineage with a wide distribution". Zoological Journal of the Linnean Society 191 (1): 228–275. doi:10.1093/zoolinnean/zlaa028. 

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