Biology:Iris lactea

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Short description: Species of plant

Iris lactea
Iris lactea 03.JPG
Scientific classification edit
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Monocots
Order: Asparagales
Family: Iridaceae
Genus: Iris
Subgenus: Iris subg. Limniris
Section: Iris sect. Limniris
Series: Iris ser. Ensatae
Species:
I. lactea
Binomial name
Iris lactea
Pall.
Synonyms[1]
  • Eremiris lactea (Pall.) Rodion.
  • Lindl. (Illegitimate) (Unknown)
  • (Fisch. ex Link) Turcz. Iris sibirica subsp. triflora
  • Iris longispatha (Balb.) Nyman
  • Iris haematophylla Fisch. ex Sims
  • Balb. Joniris fragrans
  • Iris moorcroftiana Klatt
  • Iris fragrans Fisch. ex Link
  • Wall. ex D.Don (Fisch. ex Sims) Klatt
  • Joniris triflora Iris oxypetala
  • (Balb.) Klatt Xiphion triflorum
  • Iris lactea var. lactea C.A.Mey.(Illegitimate)
  • (Balb.) Alef. Joniris longispatha
  • Iris triflora Iris sibirica var. haematophylla

Iris lactea is a species in the genus Iris. It is also in the subgenus Limniris and is the only species in the series Ensatae. The Japanese water iris, Iris ensata, is actually in series Laevigatae. It is a rhizomatous perennial, from central Asia, with pale blue or violet flowers. It is cultivated as an ornamental plant in temperate regions.

Description

Iris lactea has a thick creeping rhizome,[2][3] that is covered in reddish purple fibres.[4][5][6]

It grows to a height of between, 3–50 cm (1–18 in),[4][5][7] with a 10–30 cm (4–12 in) flowering stem.[8][9]

It has 2–4 flowers per stem, blooming between April and June,[2][4][5] or May and August in the UK.[9] The violet scented flowers,[10] can last for 2–3 weeks,[3] and measure about 5–7.5 cm in diameter.[2][3][5]

It has lanceolate (lance-shaped), green spathes, measuring 4.5–10 x 0.8–1.6 cm.[2][4][5]

The flowers come in a range of shades from pale blue to violet,[3][9] white[7][11] or yellow.[2][4][5] It has dark standards, delicate white falls, which are striated with blue, red-purple[8] or violet.[9]

It has flower stalks (pedicel) measure about 4–7 cm long, with a very short perianth tube (3 mm), 2.5–3.2 cm stamens and yellow anthers.[4][5][6]

The leaves are linear, mostly ribbed, greyish green, rising from the base of the plant. They are between 14–70 cm long and between 3–7 mm wide.[2][4][5]

It fruits (makes seeds) between June and September (after flowering), the seed capsule is narrow and cylindrical in shape, with 6 ribs running along the side of the capsule, which ends in a beak-like point. The capsule measures 6.5–7.5 × 1–1.4 cm. The fruiting stems are unequal, ranging from 4–10 cm. Inside the capsule, are maroon-brown seeds which are pyriform (pear shaped).[2][4][5]

Biochemistry

In 2008, a study was carried out on the anatomical structure of the leaf and drought resistance of 4 different species of Iris (Iris songarica, Iris potaninii, Iris loczyi and Iris lactea) from Qinghai, China. It showed that all the species were strongly adaptable to drought conditions.[12]

Genetics

In 2009, a karyotype analysis was carried out on 10 irises found in China, it found the chromosome counts, including Iris lactea of 2n=40.[13]

As most irises are diploid, having two sets of chromosomes, this can be used to identify hybrids and classification of groupings.[9] It has a chromosome count of 2n=40, 44, 50.[7][10]

Taxonomy

It is commonly known in the UK as the 'white flowered iris'[14] or in the US as 'milky iris'.[4][15] It is known as 'nomajiris', in Sweden.[16]

In Ladakhi of Tibet, it is known as 'Tesmamentok',[4] meaning pale purple flower.[17]

The Latin specific epithet Lactea means milky color,[10] hence the common name 'milky iris'.

In China, it is written as 白花马蔺 白花马蔺 in Chinese script and known as 'bai hua ma lin' in Pidgin in China.[5][16] It also known as Ma Lin or Malan flower.[citation needed]

It was first published and described by Pallas in 'Reise Russ. Reich' (Reise durch Verschiedene Provinzen des Russischen Reichs – translated as 'Travel through various provinces of the Russian Empire') in 1776.[16][18]

The taxonomy of this species has been very confused. It was originally named Iris ensata lactea (Thunberg) in 'Transactions of the Linnean Society of London' (page 328) on 1 May 1794 but later Iris ensata was re-classified as Iris kaempferi, which is now a synonym of Iris ensata (within Iris series Laevigatae). Even Dykes in his books Iris Genus (1913) and Handbook of Iris (19 ) got the name incorrect and recorded it as 'Iris ensata'. Later, writers have tried to rectify the mistake. Fritz Kohlein in his book Iris (1987) called it 'Iris ensata auct. non Thunberg.'[10]

Many new names were considered, but determining which name was first used and should have precedence has been difficult, so it has generally been left as Iris lactea (Pallas).[8][10]

Iris lactea is an accepted name by the RHS,[14] and it was verified by United States Department of Agriculture Agricultural Research Service on 2 October 2014.[16]

Distribution and habitat

Iris lactea comes from wide range of areas. It is native to temperate and tropical regions of Asia.[16]

Range

Iris lactea in Mongolia

It is found in Afghanistan, Kashmir (including Guraiz and Himachal Pradesh),[4][6] Kazakhstan, Central Asia, India (including Ladakh),[4] Pakistan ,[4] Russia (in Siberia[3] and Primorye[2]), Tibet, China , Mongolia and Korea.[7][8][11] In China, it is found within the Chinese provinces of Anhui, Gansu, Hebei, Heilongjiang, Henan, Hubei, Jiangsu, Jilin, Liaoning, Nei Mongol, Ningxia, Qinghai, Shaanxi, Shandong, Shanxi, Sichuan,[13] Xinjiang and Xi-zang (Chinese Tibet).[5]

It is listed with Iris bloudowii, Iris humilis, Iris ruthenica, Iris sibirica, Iris tenuifolia and Iris tigridia as being found in the Altai-Sayan region (where Russia, China, Mongolia and Kazakhstan come together).[19]

It is regarded as a wildflower in India and China.[4][5]

Habitat

Plant label showing wide range of distribution

I. lactea grows in a wide range of habitats. Including steppe meadows, turf slopes,[3] heavily grazed river shore within desert steppe,[20] grasslands, roadsides,[5] grassy slopes and hillsides.[2][5]

It grows at altitudes of between 600–3800 m above sea level.[2][5][6]

It flowers as late as late summer in the wild.[21]

Cultivation

Since Iris lactea, is one of the most common wild irises across all of China, living it a diverse range of habitats. It is very tolerant of most garden conditions.[10]

It is hardy to USDA Zones 3–9.[7]

It is very tolerant plant, growing in a variety of soils (including those that dry out in summer),[9][21] salty areas and can be used as a soil improver.[9] It prefers sunny places, but can grow successfully in shady places.[3]

It has been planted in mass on high leaded soils to reduce the lead contamination.[22]

It is suitable for cultivation in rock gardens and group planting.[3]

Propagation

It can be propagated by seed and by division.[3]

Hybrids and cultivars

Many of the synonyms may represent distinct varieties of lactea. The following is a list of named variations botanical and horticultural:

  • 'Biglumis' (Robert Sweet, 1835, Siberia) – now called Iris lactea Pallas var. biglumis Koidz.)[10]
  • 'Chinensis' (which grows in Korea, Russia and India)[9][10]
  • 'Chrysantha' (yellow flowers)[5][9]
  • 'Ensata Chinensis'
  • 'Ensata Grandiflora' (Dykes 1913)
  • 'Ensata Grandiflora Alba' (white form of 'Ensata Grandiflora', found at Ohio State University Bot. Garden 1933)
  • 'Hyacinthiana' (Collected by Reginald Farrer, 1914–1915, Tibet or W. China-Kansu)
  • 'Iliensis' (Poljakov, From near the Ili River, Kazakhstan)
  • 'Illini Fountain' (1993, Budapest, Hungary)
  • 'Mani' (1935, Tibet)
  • 'Moorcroftiana' (Wallich 1828)
  • 'Pabularia' (1888, Kashmir)
  • 'Redundant' (2002, Denver Botanic Garden)[10]
  • var. lactea (white flowers,[9] with purple veins and pale violet inner segments[5])

Iris lactea has been used in hybridization for the selection of dwarf bearded irises.[3]

Known hybrids:

  • 'Calsata Hybrids' (Tomas Tamberg, 1979, cross of Iris douglasiana × Iris ensata)
  • 'Chrysata Charme' (Tamberg, 2001, cross of Sino-sibirica × I. lactea)
  • 'Hamadryad' (1931, Iris 'Watsoniana' × Iris ensata)[10]

Uses

It is widely grown throughout China, where it is often used as an ingredient in a herbal contraceptive.[9] It has also been used in the anti-cancer drug 'Irisquinone', which comes from a herbal remedy.[23] The rhizomes are also used in traditional oriental medicine,[2][4] including Tibet.[24]

The leaves are used as fodder for animals, and for thatching, matting and basket work,[2][4] and its leaf fibres are also used in paper making and for brushes.[9] The flowers contain the pigment – anthocyanin.[4]

It has also been used in veterinary medicine. The consumption of the flowers and seeds is thought to increase a cow's milk yield.[25]

Culture

In Afghanistan, it has been banned by the Taliban.[26] This maybe due to its use in herbal medicines.

References

  1. ↑ "Iris lactea Pall". theplantlist.org. http://www.theplantlist.org/tpl1.1/record/kew-322026?ref=tpl1. 
  2. ↑ 2.00 2.01 2.02 2.03 2.04 2.05 2.06 2.07 2.08 2.09 2.10 2.11 Hoskovec, Ladislav. "IRIS LACTEA Pall. – White-flower Chinese Iris". botany.cz. http://botany.cz/en/iris-lactea/. 
  3. ↑ 3.0 3.1 3.2 3.3 3.4 3.5 3.6 3.7 3.8 3.9 "Iris lactea Pall.". efloras.org. http://www.efloras.org/florataxon.aspx?flora_id=120&taxon_id=200028182. 
  4. ↑ 4.00 4.01 4.02 4.03 4.04 4.05 4.06 4.07 4.08 4.09 4.10 4.11 4.12 4.13 4.14 4.15 4.16 "Milky Iris". flowersofindia.net. http://www.flowersofindia.net/catalog/slides/Milky%20Iris.html. 
  5. ↑ 5.00 5.01 5.02 5.03 5.04 5.05 5.06 5.07 5.08 5.09 5.10 5.11 5.12 5.13 5.14 5.15 5.16 "Flora of China 24: 297–312. 2000, 1. IRIS Linnaeus, Sp. Pl. 1: 38. 1753.". flora.huh.harvard.edu. 2000. pp. 7–8. http://flora.huh.harvard.edu/china/PDF/PDF24/iris.pdf. 
  6. ↑ 6.0 6.1 6.2 6.3 Akhter, Chesfeeda; Khuroo, Anzar A.; Malik, Akhtar H.; Dar, G. H. (8 November 2012). "A TAXONOMIC APPRAISAL OF GENUS IRIS L. (IRIDACEAE) IN KASHMIR HIMALAYA, INDIA". academia.edu. https://www.academia.edu/7635873. 
  7. ↑ 7.0 7.1 7.2 7.3 7.4 Kramb, D (25 September 2004). "Iris lactea". signa.org. http://www.signa.org/index.pl?Iris-lactea. 
  8. ↑ 8.0 8.1 8.2 8.3 Cassidy, George E.; Linnegar, Sidney (1982). Growing Irises. Bromley: Christopher Helm. p. 138. ISBN 0-88192-089-4. 
  9. ↑ 9.00 9.01 9.02 9.03 9.04 9.05 9.06 9.07 9.08 9.09 9.10 9.11 Austin, Claire (2005). Irises: A Gardener's Encyclopedia. Timber Press, Incorporated. ISBN 978-0881927306. 
  10. ↑ 10.0 10.1 10.2 10.3 10.4 10.5 10.6 10.7 10.8 10.9 "(SPEC) Iris lactea Pallas". wiki.irises.org. http://wiki.irises.org/bin/view/Spec/SpecLactea. 
  11. ↑ 11.0 11.1 "Beardless Irises Two". pacificbulbsociety.org. 2 March 2013. http://www.pacificbulbsociety.org/pbswiki/index.php/BeardlessIrisesTwo. 
  12. ↑ Wang, Jun; Gao, Yi-ke (May 2008). "Study on the Relationship between Leaf Anatomical Structure and Drought Resistance of Four Species of Iris". Heilongjiang Agricultural Sciences (Landscape Architecture College of Beijing Forestry University, Beijing). http://en.cnki.com.cn/Article_en/CJFDTOTAL-HLJN200805040.htm. Retrieved 22 January 2015. 
  13. ↑ 13.0 13.1 Yu, Xiao-Fang; Zhang, Hai-Qing; Yuan, Ming; Zhou, Yong-Hong (2009). "Karyotype studies on ten Iris species (Iridaceae) from Sichuan, China". Caryologia 62 (3): 253–260. doi:10.1080/00087114.2004.10589690. http://www.caryologia.unifi.it/past_volumes/62_3/01-1574-(253-260)-VF.pdf. Retrieved 23 March 2015. 
  14. ↑ 14.0 14.1 "Iris lactea". rhs.org.uk. https://www.rhs.org.uk/Plants/9293/White-flowered-iris/Details. 
  15. ↑ Scott Ogden and Lauren Springer OgdenPlant Driven Design: Creating Gardens That Honor Plants, Place and Spirit, p. 114, at Google Books
  16. ↑ 16.0 16.1 16.2 16.3 16.4 {{citation | mode = cs1 | title = Iris lactea | work = Germplasm Resources Information Network (GRIN) | url = | publisher = [[Organization:Agricultural Research ServAgricultural Research Service (ARS), United States Department of Agriculture (USDA) | access-date = 31 March 2015 }}
  17. ↑ Sanjeev KumarAmazing Land Ladakh: Places, People, and Culture, p. 44, at Google Books
  18. ↑ "Iridaceae Iris lactea Pall.". ipni.org (International Plant Names Index). http://www.ipni.org/ipni/idPlantNameSearch.do?id=438774-1. 
  19. ↑ "Biodiversity of Altai-Sayan Ecoregion". bioaltai-sayan.ru. http://www.bioaltai-sayan.ru/regnum/eng/species_all.php?right=box-spec-p/iris.php. 
  20. ↑ "Iris lactea Pall.". greif.uni-greifswald.de. http://greif.uni-greifswald.de/floragreif/?flora_search=Image&record_id=12968. 
  21. ↑ 21.0 21.1 Stebbings, Geoff (1997). The Gardener's Guide to Growing Irises. Newton Abbot: David and Charles. p. 17. ISBN 0715305395. https://archive.org/details/gardenersguideto00steb. 
  22. ↑ Various editors Amidohydrolases – Advances in Research and Application 2013 Edition, p. 276, at Google Books
  23. ↑ Michael A. Lerner Choices in Healing: Integrating the Best of Conventional and Complementary, p. 381, at Google Books
  24. ↑ Dr Barry Clark (translator) The Quintessence Tantras of Tibetan Medicine at Google Books
  25. ↑ Umberto Quattrocchi CRC World Dictionary of Medicinal and Poisonous Plants: Common Names, p. 2105, at Google Books
  26. ↑ "Iris lactea". plantlust.com. http://www.plantlust.com/plants/iris-lactea/. 

Sources

  • Aldén, B., S. Ryman & M. Hjertson. 2009. Våra kulturväxters namn – ursprung och användning. Formas, Stockholm (Handbook on Swedish cultivated and utility plants, their names and origin).
  • Chinese Academy of Sciences. 1959–. Flora reipublicae popularis sinicae.
  • Czerepanov, S. K. 1995. Vascular plants of Russia and adjacent states (the former USSR).
  • Huxley, A., ed. 1992. The new Royal Horticultural Society dictionary of gardening.
  • Khassanov, F. O. & N. Rakhimova. 2012. Taxonomic revision of the genus Iris L. (Iridaceae Juss.) for the flora of Central Asia. Stapfia 97:175.
  • Kitagawa, M. 1979. Neo-lineamenta florae Manshuricae.
  • Komarov, V. L. et al., eds. 1934–1964. Flora SSSR. [= I. ensata Thunb.].
  • Mathew, B. 1981. The Iris. 125–127.
  • Nasir, E. & S. I. Ali, eds. 1970–. Flora of [West] Pakistan.
  • Waddick, J. W. & Zhao Yu-tang. 1992. Iris of China.
  • Walters, S. M. et al., eds. 1986–. European garden flora.

External links


Wikidata ☰ Q6747387 entry



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