Biology:Kirengellida

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Short description: Extinct family of shelled animals

Kirengellida
Temporal range: Late Cambrian–Mid Ordovician
Scientific classification e
Kingdom: Animalia
Phylum: Brachiopoda (?)
Order: Kirengellida
Rozov, 1975
Superfamilies[1]
  • Hypseloconoidea
  • Kirengelloidea
Synonyms[1]

Romaniellida

The Kirengellids are a group of problematic Cambrian fossil shells of marine organisms. The shells bear a number of paired muscle scars on the inner surface of the valve.

These fossils have conventionally been regarded as monoplacophoran molluscs, and possibly ancestral to gastropods or cephalopods.[2] They were presumed to be exogastric on the presumption that their larger muscle scars were anterior,[3][4] but it may be dangerous to compare these scars with molluscan musculature.[5] In any case, they coiled in the opposite direction to Romaniella.[5] However, their calcitic shells, the position of the muscle scars, and putative association with secondary shell elements, make a brachiopod affinity possible, by analogy with the mobergellans: a group of phosphatic shells from the same time period, with a similar set of muscle scars.[5] There is also strong similarity to the contemporary brachiopod group, the Craniopsids. In the case of this diagnosis, a simple lophophore apparatus is postulated to sit between the muscle scars and the edges of the shell.[5] On the other hand, Vendrasco (2012) reaffirmed the interpretation of the kirengellids as molluscs, noting that the Kirengella muscle scar pattern is also similar to what occurs in monoplacophorans.[6][7] Bouchet et al. (2017) classified Kirengellida as an order within the subclass Tergomya.[8]

Included taxa

After [5]: fig. 5 

  • Kirengella Rozov, 1968 (Upper Cambrian)
    • Kirengella alta Whitfield 1889
    • Kirengella ayaktchica - type species
    • Kirengella expansus
    • Kirengella kultavasaensis Doguzhaeva 1972
    • Kirengella oregonensis
    • Kirengella pyramidalis
    • Kirengella rectilateralis Berkey 1898
    • Kirengella stabilis Berkey 1898
    • Kirengella washingtonense
  • Hypseloconus (Upper Cambrian)
  • Lenaella (Tremadoc / Lower Ordovician)
  • Nyuella (Tremadoc / Lower Ordovician)
  • Romaniella (Arenig / late Lower Ordovician)
  • Moyerokania (Arenig / late Lower Ordovician)
  • Angarella (Arenig / late Lower Ordovician)
  • Pygmaeoconus (Llanvirn / early Middle Ordovician)

References

  1. 1.0 1.1 "Kirengellida". MolluscaBase. World Register of Marine Species. 2026. http://www.marinespecies.org/aphia.php?p=taxdetails&id=1601051. 
  2. e.g. Yochelson, E. L.; Flower, R. H.; Webers, G. F. (1973), "The bearing of the new Late Cambrian monoplacophoran genus Knightoconus upon the origin of the Cephalopoda", Lethaia 6 (3): 275–309, doi:10.1111/j.1502-3931.1973.tb01199.x 
  3. Yochelson, E.L. (1978). "An alternative approach to the interpretation of the phylogeny of ancient mollusks". Malacologia 17 (2): 165–191. https://archive.org/details/malacologia171978inst. 
  4. Yochelson, E.L; Webers, G.F. (2006). "A restudy of the Late Cambrian Molluscan fauna of Berkey (1898) from Taylors Falls, Minnesota". Minnesota Geological Survey Report of Investigations 64: 60. http://conservancy.umn.edu/bitstream/58939/1/RI64%5B1%5D.pdf. 
  5. 5.0 5.1 5.2 5.3 5.4 Dzik, Jerzy (2010). "BRACHIOPOD IDENTITY OF THE ALLEGED MONOPLACOPHORAN ANCESTORS OF CEPHALOPODS". Malacologia 52 (1): 97–113. http://www.paleo.pan.pl/people/Dzik/Publications/Angarella.pdf. 
  6. Vendrasco, M. J. (2012). "Early evolution of molluscs". Mollusks: Morphology, Behavior and Ecology. Nova Science Publishers. pp. 1–43. ISBN 9781621009870. https://www.researchgate.net/publication/246548102. 
  7. Cope, J. C. W.; Ebbestad, J. O. R. (2023). "Tergomyan molluscs from the Early Ordovician of the Llangynog Inlier, South Wales, UK". PalZ. doi:10.1007/s12542-023-00667-5. 
  8. Bouchet, P.; Rocroi, J.-P.; Hausdorf, B.; Kaim, A.; Kano, Y.; Nützel, A.; Parkhaev, P.; Schrödl, M. et al. (2017). "Revised classification, nomenclator and typification of gastropod and monoplacophoran families". Malacologia 61 (1-2): 1-526. doi:10.4002/040.061.0201. 
  • S. N. Rozov. 1975. A new order of the Monoplacophora. Paleontological Journal 15(1):39-43
  • G.P. Wahlman. 1992. Middle and Upper Ordovician symmetrical univalved mollusks (Monoplacophora and Bellerophontina) of the Cincinnati Arch region. United States Geological Survey Professional Paper 1066(O):1-123

Wikidata ☰ Q138648830 entry