Biology:List of ground beetle genera

Short description: none

This page lists the genera in the family Carabidae, according to Catalogue of Life and Carabcat Database as of March 2023, excluding Tiger Beetles (which are considered to be in the family Cicindelidae).

Family Carabidae

Subfamily Anthiinae Bonelli, 1813

Tribe Anthiini Bonelli, 1813

Anthia Weber, 1801

Atractonotus Perroud, 1847

Baeoglossa Chaudoir, 1850

Cycloloba Chaudoir, 1850

Cypholoba Chaudoir, 1850

Eccoptoptera Chaudoir, 1878

Gonogenia Chaudoir, 1844

Netrodera Chaudoir, 1850

Tribe Helluonini Hope, 1838

Subtribe Helluonina Hope, 1838

Aenigma Newman, 1836

Ametroglossus Sloane, 1914

Dicranoglossus Chaudoir, 1872

Epimicodema Sloane, 1914

Gigadema J.Thomson, 1859

Helluapterus Sloane, 1914

Helluarchus Sloane, 1914

Helluo Bonelli, 1813

Helluodema Laporte, 1867

Helluonidius Chaudoir, 1872

Helluopapua Darlington, 1968

Helluosoma Laporte, 1867

Neohelluo Sloane, 1914

Platyhelluo Baehr, 2005

Subtribe Omphrina Jedlicka, 1941

Colfax Andrewes, 1920

Creagris Nietner, 1857

Dailodontus Reiche, 1843

Erephognathus Alluaud, 1932

Helluobrochus Reichardt, 1974

Helluomorpha Laporte, 1834

Helluomorphoides Ball, 1951

Macrocheilus Hope, 1838

Meladroma Motschulsky, 1855

Omphra Dejean, 1825

Pleuracanthus Gray, 1832

Triaenogenius Chaudoir, 1877

Tribe Physocrotaphini Chaudoir, 1863

Anguloderus Anichtchenko & Sciaky, 2017

Foveocrotaphus Anichtchenko, 2014

Helluodes Westwood, 1847

Holoponerus Fairmaire, 1883

Physocrotaphus Parry, 1849

Physoglossus Akhil & Sabu, 2020

Pogonoglossus Chaudoir, 1863

Schuelea Baehr, 2004

Subfamily Apotominae LeConte, 1853

Apotomus Illiger, 1807

Subfamily Brachininae Bonelli, 1810

Tribe Brachinini Bonelli, 1810

Aptinoderus Hubenthal, 1919

Aptinus Bonelli, 1810

Brachinulus Basilewsky, 1958

Brachinus Weber, 1801

Mastax Fischer von Waldheim, 1828

Pheropsophus Solier, 1833

Styphlodromus Basilewsky, 1959

Styphlomerus Chaudoir in Putzeys, 1875

Tribe Crepidogastrini Jeannel, 1949

Brachynillus Reitter, 1904

Crepidogaster Boheman, 1848

Crepidogastrillus Basilewsky, 1959

Crepidogastrinus Basilewsky, 1957

Crepidolomus Basilewsky, 1959

Crepidonellus Basilewsky, 1959

Subfamily Broscinae Hope, 1838

Tribe Broscini Hope, 1838

Subtribe Axonyina Roig-Juñent, 2000

Broscodes Bolivar y Pieltain, 1914

Rawlinsius Davidson & Ball, 1998

Subtribe Baripodina Jeannel, 1941

Baripus Dejean, 1828

Bembidiomorphum Champion, 1918

Subtribe Broscina Hope, 1838

Broscodera Lindroth, 1961

Broscosoma Rosenhauer, 1846

Broscus Panzer, 1813

Chaetobroscus Semenov, 1900

Craspedonotus Schaum, 1863

Eobroscus Kryzhanovskij, 1951

Kashmirobroscus J.Schmidt; Wrase & Sciaky, 2013

Miscodera Eschscholtz, 1830

Zacotus LeConte, 1869

Subtribe Creobiina Jeannel, 1941

Acallistus Sharp, 1886

Adotela Laporte, 1867

Anheterus Putzeys, 1868

Bountya Townsend, 1971

Brithysternum W.J.MacLeay, 1873

Cascellius Curtis, 1838

Cerotalis Laporte, 1867

Creobius Guérin-Méneville, 1838

Gnathoxys Westwood, 1842

Nothocascellius Roig-Juñent, 1995

Promecoderus Dejean, 1829

Subtribe Nothobroscina Roig-Juñent, 2000

Chylnus Sloane, 1920

Diglymma Sharp, 1886

Eurylychnus Bates, 1891

Mecodema Blanchard, 1843

Monteremita Seldon & Holwell, 2019

Nothobroscus Roig-Juñent & Ball, 1995

Oregus Putzeys, 1868

Orthoglymma Liebherr; Marris; Emberson; Syrett & Roig-Juñent, 2011

Percolestus Sloane, 1892

Percosoma Schaum, 1858

Subfamily Carabinae Linnaeus, 1802

Tribe Carabini Linnaeus, 1802

Subtribe Carabina Linnaeus, 1802

Aplothorax G.R.Waterhouse, 1842

Calosoma Weber, 1801

Carabus Linnaeus, 1758

Subtribe Ceroglossina Lapouge, 1927

Ceroglossus Solier, 1848

Tribe Cychrini Perty, 1830

Subtribe Cychrina Perty, 1830

Cychropsis Boileau, 1901

Cychrus Fabricius, 1794

Scaphinotus Dejean, 1826

Sphaeroderus Dejean, 1826

Subtribe Pamborina Hope, 1838

Maoripamborus Brookes, 1944

Pamborus Latreille, 1812

Subfamily Ctenodactylinae Laporte, 1834

Tribe Ctenodactylini Laporte, 1834

Alachnothorax Liebke, 1929

Amblycoleus Chaudoir, 1872

Antipionycha Liebke, 1928

Askalaphium Liebke, 1938

Ctenodactyla Dejean, 1825

Leptotrachelon Liebke, 1928

Leptotrachelus Latreille, 1829

Oilea Liebke, 1931

Parapionycha Liebke, 1929

Pionycha Chaudoir, 1848

Plagiotelum Solier, 1849

Propionycha Liebke, 1928

Pseudometabletus Liebke, 1930

Schidonychus Klug, 1834

Teukrus Liebke, 1931

Wate Liebke, 1928

Tribe Hexagoniini G.Horn, 1881

Dinopelma Bates, 1889

Hexagonia Kirby, 1825

Omphreoides Fairmaire, 1896

Subfamily Dryptinae Bonelli, 1810

Tribe Dryptini Bonelli, 1810

Desera Hope, 1831

Drypta Latreille, 1797

Megadrypta Sciaky & Anichtchenko, 2020

Neodrypta Basilewsky, 1960

Nesiodrypta Jeannel, 1949

Prionodrypta Jeannel, 1949

Tribe Galeritini Kirby, 1825

Ancystroglossus Chaudoir, 1863

Eunostus Laporte, 1835

Galerita Fabricius, 1801

Trichognatha Latreille, 1829

Tribe Zuphiini Bonelli, 1810

Subtribe Dicrodontina Machado, 1992

Dicrodontus Chaudoir, 1872

Subtribe Leleupidiina Basilewsky, 1951

Colasidia Basilewsky, 1954

Gunvorita Landin, 1955

Leleupidia Basilewsky, 1951

Neoleleupidia Basilewsky, 1953

Paraleleupidia Basilewsky, 1951

Subtribe Planetina Jedlicka, 1941

Planetes W.S.MacLeay, 1825

Subtribe Zuphiina Bonelli, 1810

Acrogenys W.J.MacLeay, 1864

Agastus Schmidt-Goebel, 1846

Chaudoirella Mateu, 1982

Coarazuphium Gnaspini; Vanin & Godoy, 1998

Ildobates Español, 1966

Metaxidius Chaudoir, 1852

Metazuphium Mateu, 1992

Mischocephalus Chaudoir, 1863

Parazuphium Jeannel, 1942

Polistichus Bonelli, 1810

Pseudaptinus Laporte, 1834

Speothalpius B.Moore, 1995

Speozuphium B.Moore, 1995

Typhlozuphium Baehr, 2014

Zuphioides Ball & Shpeley, 2013

Zuphium Latreille, 1805

Subfamily Elaphrinae Latreille, 1802

Blethisa Bonelli, 1810

Diacheila Motschulsky, 1844

Elaphrus Fabricius, 1775

†Elaphrotites Haupt, 1856

Subfamily Gineminae Ball & Shpeley, 2002

Ginema Ball & Shpeley, 2002

Subfamily Harpalinae Bonelli, 1810

Tribe Anisodactylini Lacordaire, 1854

Allendia Noonan, 1974

Allocinopus Broun, 1903

Amphasia Newman, 1838

Anisodactylus Dejean, 1829

Anisostichus Emden, 1953

Cenogmus Sloane, 1898

Chydaeus Chaudoir, 1854

Crasodactylus Guérin-Méneville, 1847

Criniventer Emden, 1953

Diachromus Erichson, 1837

Dicheirus Mannerheim, 1843

Gaioxenus Broun, 1910

Geopinus LeConte, 1847

Gnathaphanus W.S.MacLeay, 1825

Gynandromorphus Dejean, 1829

Haplaner Chaudoir, 1878

Harpalomimetes Schauberger in Csiki, 1932

Hypharpax W.S.MacLeay, 1825

Maoriharpalus Larochelle & Larivière, 2005

Nornalupia Kataev, 2002

Notiobia Perty, 1830

Parabaris Broun, 1881

Progonochaetus G.Müller, 1938

Pseudanisotarsus Noonan, 1973

Pseudognathaphanus Schauberger, 1932

Pseudorhysopus Kataev & Wrase, 2001

Rhysopus Andrewes, 1929

Scybalicus Schaum, 1862

Triplosarus Bates, 1874

Tuiharpalus Larochelle & Larivière, 2005

Xestonotus LeConte, 1853

Tribe Harpalini Bonelli, 1810

Subtribe Amblystomina Fauvel, 1889

Amblystomus Erichson, 1837

Anomostomus LaFerté-Sénectère, 1853

Barysomus Dejean, 1829

Oosoma Nietner, 1857

Subtribe Ditomina Bonelli, 1810

Bronislavia Semenov, 1891

Carenochyrus Solsky, 1874

Carterus Dejean, 1830

Chilotomus Chaudoir, 1842

Ditomus Bonelli, 1810

Dixus Billberg, 1820

Eocarterus Stichel, 1923

Eucarterus Reitter, 1900

Graniger Motschulsky, 1864

Indocarterus Kataev & Wrase, 2018

Liochirus Tschitscherine, 1897

Machozetus Chaudoir, 1850

Odotoncarus Solier, 1835

Oedesis Motschulsky, 1850

Pachycarus Solier, 1835

Parapenthes Kataev & Wrase, 2018

Penthus Chaudoir, 1843

Phorticosomus Schaum, 1863

Proditomus Schauberger, 1934

Pseudaristus Reitter, 1900

Tschitscherinellus Csiki, 1906

Subtribe Harpalina Bonelli, 1810

Acinopus Dejean, 1821

Afromizonus Basilewsky, 1947

Allosiopelus N.Ito, 1995

Amblygnathus Dejean, 1829

Anisocnemus Chaudoir, 1843

Athrostictus Bates, 1878

Axinotoma Dejean, 1829

Aztecarpalus Ball, 1970

Bleusea Bedel, 1897

Boeomimetes Péringuey, 1896

Bradybaenus Dejean, 1829

Coleolissus Bates, 1892

Cratacanthus Dejean, 1829

Cratognathus Dejean, 1829

Daptus Fischer von Waldheim, 1823

Dioryche W.S.MacLeay, 1825

Discoderus LeConte, 1853

Dregus Motschulsky, 1864

Ectinothorax Alluaud, 1941

Eriophonus Tschitscherine, 1901

Euryderus LeConte, 1846

Geodromus Dejean, 1829

Harpalinus Jeannel, 1946

Harpaliscus Bates, 1892

Harpalobrachys Tschitscherine, 1899

Harpalodiodes Bousquet, 2002

Harpalomimus Facchini & Giachino, 2020

Harpalomorphus Péringuey, 1896

Harpalus Latreille, 1802

Harpathaumas Basilewsky, 1947

Harponixus Basilewsky, 1950

Hartonymus Casey, 1914

Heteracantha Brullé, 1835

Hyphaereon W.S.MacLeay, 1825

Indiophonus N.Ito, 1996

Kareya Andrewes, 1919

Lampetes Andrewes, 1940

Laparhetes Jeannel, 1946

Liodaptus Bates, 1889

Meroctenus Gemminger & Harold, 1868

Microderes Faldermann, 1836

Neoaulacoryssus Noonan, 1985

Neodiachipteryx Noonan, 1985

Neohyparpalus Clarke, 1981

Neophygas Noonan, 1976

Nesacinopus Tschitscherine, 1900

Nesarpalus Bedel, 1897

Nipponoharpalus Habu, 1973

Nothodaptus Maindron, 1906

Oesyperus Andrewes, 1923

Omostropus Péringuey, 1896

Ooidius Chaudoir, 1847

Ophoniscus Bates, 1892

Ophonus Dejean, 1821

Oxycentrus Chaudoir, 1854

Panagrius Andrewes, 1933

Pangus Dejean, 1821

Parasiopelus Basilewsky, 1946

Paraulacoryssus Shpeley; Hunting & Ball, 2017

Parophonus Ganglbauer, 1891

Paulianoscirtus Basilewsky, 1976

Penthophonus Reitter, 1900

Piosoma LeConte, 1847

Platymetopsis Ball & Maddison, 1987

Platymetopus Dejean, 1829

Prakasha Andrewes, 1919

Pseudodiachipteryx Burgeon, 1936

Pseudohyparpalus Basilewsky, 1946

Pseudoselenophorus Péringuey, 1896

Selenophorus Dejean, 1829

Selenotichnus Kataev, 1999

Siopelophonus Kataev, 2015

Siopelus Murray, 1859

Stenomorphus Dejean, 1831

Trichopselaphus Chaudoir, 1843

Trichotichnus A.Morawitz, 1863

Trichoxycentrus N.Ito, 2000

Typsiharpalus Tschitscherine, 1901

Tribe Pelmatellini Bates, 1882

Hakaharpalus Larochelle & Larivière, 2005

Kupeharpalus Larochelle & Larivière, 2005

Lecanomerus Chaudoir, 1850

Nemaglossa Solier, 1849

Notospeophonus B.Moore, 1962

Pelmatellus Bates, 1882

Syllectus Bates, 1878

Trachysarus Reed, 1874

Tribe Stenolophini Kirby, 1837

Acupalpus Latreille, 1829

Agonoleptus Casey, 1914

Amerinus Casey, 1884

Angionychus Klug, 1853

Anthracus Motschulsky, 1850

Batoscelis Dejean, 1836

Bradycellus Erichson, 1837

Cratosoma Jeannel, 1948

Cyptomicrus Vinson, 1939

Dicheirotrichus Jacquelin du Val, 1855

Egadyla Alluaud, 1916

Euthenarus Bates, 1874

Fuminoria Morita, 2006

Goniocellus Casey, 1914

Gugheorites Basilewsky, 1951

Haplanister B.Moore, 1996

Hemiaulax Bates, 1892

Hippoloetis Laporte, 1835

Idiomelas Tschitscherine, 1900

Kaffovatus Clarke, 1972

Kenyacus Alluaud, 1917

Kiwiharpalus Larochelle & Larivière, 2005

Lioholus Tschitscherine, 1897

Loxoncus Schmidt-Goebel, 1846

Pachytrachelus Chaudoir, 1852

Parabradycellus N.Ito, 2003

Paramecus Dejean, 1829

Philodes LeConte, 1861

Pholeodytes Britton, 1962

Pogonodaptus G.Horn, 1881

Polpochila Solier, 1849

Psychristus Andrewes, 1930

Rhabidius Basilewsky, 1948

Stenolophus Dejean, 1821

Uenanthracus Kasahara, 1994

Subfamily Hiletinae Schiödte, 1847

Eucamaragnathus Jeannel, 1938

Hiletus Schiödte, 1847

Subfamily Lebiinae Bonelli, 1810

Tribe Cyclosomini Laporte, 1834

Subtribe Corsyrina Ganglbauer

Corsyra Dejean, 1825

Discoptera Semenov, 1889

Subtribe Cyclosomina Laporte, 1834

Cyclosomus Latreille, 1829

Sarothrocrepis Chaudoir, 1850

Tetragonoderus Dejean, 1829

Subtribe Graphipterina Latreille, 1802

Graphipterus Latreille, 1802

Piezia Brullé, 1834

Trichopiezia Nègre, 1955

Subtribe Masoreina Chaudoir, 1871

Anaulacus W.S.MacLeay, 1825

Atlantomasoreus Mateu, 1984

Leuropus Andrewes, 1947

Lophidius Dejean, 1831

Masoreus Dejean, 1821

Mnuphorus Chaudoir, 1873

Odontomasoreus Darlington, 1968

Somoplatodes Basilewsky, 1986

Somoplatus Dejean, 1829

Subtribe Nemotarsina Bates, 1883

Nemotarsus LeConte, 1853

Tribe Lachnophorini LeConte, 1853

Subtribe Calophaenina Jeannel, 1948

Calophaena Klug, 1821

Calophaenoidea Liebke, 1930

Subtribe Lachnophorina LeConte, 1853

Amphithasus Bates, 1871

Anchonoderus Reiche, 1843

Aporesthus Bates, 1871

Asklepia Liebke, 1938

Balligratus Moret & Ortuño, 2017

Calybe Laporte, 1834

Ega Laporte, 1834

Eucaerus LeConte, 1853

Euphorticus G.Horn, 1881

Grundmannius Basilewsky, 1965

Guatemalteca Erwin, 2004

Lachnaces Bates, 1872

Lachnophorus Dejean, 1831

Peruphorticus Erwin & Zamorano, 2014

Pseudophorticus Erwin, 2004

Selina Motschulsky, 1858

Tribe Lebiini Bonelli, 1810

Subtribe Agrina Kirby, 1837

Abaditicus Ball & Hilchie, 1983

Abrodiella Bousquet, 2002

Agra Fabricius, 1801

Allardina Basilewsky, 1963

Amelus Chaudoir, 1873

Anomotariella Baehr, 2012

Anomotarus Chaudoir, 1875

Apterodromites Mateu, 1976

Aspasiola Chaudoir, 1877

Calleida Latreille, 1824

Calleidomorpha Motschulsky, 1855

Callidadelpha Steinheil, 1875

Chaudoirina Mateu, 1955

Cryptobatis Eschscholtz, 1829

Cyanotarus Reed, 1874

Cylindronotum Putzeys, 1845

Demetrida White, 1846

Diabaticus Bates, 1878

Do Baehr, 2009

Dromidea Perroud & Montrouzier, 1864

Epikastea Liebke, 1935

Eujalmenus Bousquet, 2002

Euplatia Chaudoir, 1873

Eurycalleida Maindron, 1905

Falsodromius Mateu, 1976

Glycia Chaudoir, 1842

Hyboptera Chaudoir, 1873

Hybopteroides Erwin & Ball, 2012

Infernophilus Larson, 1969

Kteatus Liebke, 1936

Lipostratia Chaudoir, 1873

Merizomena Chaudoir, 1873

Mimodromius Chaudoir, 1873

Mimophilorhizus Mateu, 1993

Oechalius Liebke, 1935

Ogygium Liebke, 1935

Onota Chaudoir, 1873

Otoglossa Chaudoir, 1873

Paraglycia Bedel, 1904

Phacocerus Chaudoir, 1873

Philophuga Motschulsky, 1859

Phloeocarabus W.J.MacLeay, 1871

Plochionus Dejean, 1821

Pontonoa Liebke, 1935

Pseudotoglossa Mateu, 1961

Pylartesius Liebke, 1939

Rugitarus Baehr, 2009

Speotarus B.Moore, 1964

Straneotia Mateu, 1961

Tecnophilus Chaudoir, 1877

Teiresia Liebke, 1935

Thoasia Liebke, 1939

Titaresius Liebke, 1935

Trigonothops W.J.MacLeay, 1864

Valeriaaschero Erwin, 2004

Vianasia Mateu, 1955

Subtribe Apenina Ball

Apenes LeConte, 1851

Cymindoidea Laporte, 1833

Habutarus Ball & Hilchie, 1983

Platytarus Fairmaire, 1850

Trymosternus Chaudoir, 1873

Subtribe Celaenephina Habu, 1982

Celaenephes Schmidt-Goebel, 1846

Subtribe Cymindidina Laporte, 1834

Afrotarus Jeannel, 1949

Assadecma Basilewsky, 1982

Cymindis Latreille, 1805

Hystrichopus Boheman, 1848

Leptosarcus Péringuey, 1896

Metaxymorphus Chaudoir, 1850

Petrimagnia Kryzhanovskij & Mikhailov, 1971

Pinacodera Schaum, 1857

Plagiopyga Boheman, 1848

Pseudomasoreus Desbrochers des Loges, 1904

Taridius Chaudoir, 1875

Subtribe Demetriadina Bates, 1886

Cylindrocranius Chaudoir, 1878

Demetrias Bonelli, 1810

Subtribe Dromiusina Bonelli, 1810

Afrodromius Basilewsky, 1958

Axinopalpus LeConte, 1846

Barrymooreana Baehr, 1997

Brachynopterus Bedel, 1898

Brigalowia Baehr, 2006

Calodromius Reitter, 1905

Carbonellia Mateu, 1968

Disciferella Kataev & Muilwijk, 2020

Dromiops Péringuey, 1898

Dromius Bonelli, 1810

Dromoceryx Schmidt-Goebel, 1846

Geoffreyella Baehr, 2012

Klepteromimus Péringuey, 1898

Mesolestes Schatzmayr, 1943

Metadromius Bedel, 1907

Microdaccus Schaum in Baudi di Selve, 1864

Microlestes Schmidt-Goebel, 1846

Microlestodes Baehr, 1987

Monnea Mateu, 1970

Negrea Mateu, 1968

Omophagus Andrewes, 1937

Oxoides Solier, 1849

Paradromius Fowler, 1887

Philorhizus Hope, 1838

Polyaulacus Chaudoir, 1878

Psammodromius Peyerimhoff, 1927

Pseudomonnea Mateu, 1983

Somalodromius Mateu, 1967

Xenodromius Bates, 1891

Zolotarevskyella Mateu, 1953

Subtribe Gallerucidiina Chaudoir, 1872

Gallerucidia Chaudoir, 1872

Lebidia A.Morawitz, 1862

Subtribe Lebiina Bonelli, 1810

Alkestis Liebke, 1939

Aristolebia Bates, 1892

Daer Semenov & Znojko, 1929

Lachnolebia Maindron, 1905

Lebia Latreille, 1802

Lebidema Motschulsky, 1864

Lebiomorphica Lorenz, 1998

Lebistina Motschulsky, 1864

Lebistinida Péringuey, 1898

Lionedya Chaudoir, 1871

Matabele Péringuey, 1896

Megalebia Mateu, 1972

Orthobasis Chaudoir, 1871

Pachylebia Jeannel, 1949

Pachylebiodes Mateu, 1972

Paulianolebia Mateu, 1972

Pseudopachylebia Mateu, 1972

Rhopalostyla Chaudoir, 1850

Scalidion Schmidt-Goebel, 1846

Setolebia Jedlicka, 1941

Sofota Jedlicka, 1951

Subtribe Lichnasthenina J.Thomson

Astastus Péringuey, 1896

Australovelinda Baehr, 2012

Imasakaia Arai & Morita, 2010

Lichnasthenus J.Thomson, 1858

Mimovelindopsis Mateu, 1963

Velinda Andrewes, 1921

Velindastus Schüle & Lorenz, 2008

Velindomimus Jeannel, 1955

Velindopsis Burgeon, 1937

Subtribe Lionychina Jeannel, 1948

Apristomimus Mateu, 1969

Apristus Chaudoir, 1846

Eremolestes Maindron, 1905

Lionychus Wissmann, 1846

Lorestania Anichtchenko, 2011

Metablus Jedlicka, 1958

Omobrus Andrewes, 1930

Singiliomimus Péringuey, 1896

Syntomus Hope, 1838

Tilius Chaudoir, 1876

Trichidema Basilewsky, 1956

Subtribe Metallicina Basilewsky, 1984

Euproctinus Leng & Mutchler, 1927

Metallica Chaudoir, 1873

Pachycallida Jeannel, 1949

Parena Motschulsky, 1860

Subtribe Peliocypadina Basilewsky, 1984

Demetriola Jeannel, 1949

Peliocypas Schmidt-Goebel, 1846

Subtribe Pericalina Hope, 1838

Agonocheila Chaudoir, 1848

Allophanes Andrewes, 1939

Allophanopsis Louwerens, 1952

Amphimenes Bates, 1873

Amphimenoides Kirschenhofer, 1999

Antimerina Alluaud, 1897

Arsinoe Laporte, 1834

Auchmerus Andrewes, 1929

Bellavalentis Hunting & Yang, 2019

Brachichila Chaudoir, 1870

Brachyctis Chaudoir, 1870

Catascopellus Straneo, 1969

Catascopus Kirby, 1825

Cheilagona Baehr, 2006

Coptodera Dejean, 1825

Coptoglossus Chaudoir, 1870

Coptoptera Chaudoir, 1837

Coptopterella Basilewsky, 1961

Coptopterina Basilewsky, 1956

Crassagena Baehr, 2006

Cylindropectus Lorenz, 1998

Dolichoctis Schmidt-Goebel, 1846

Dontolobus Basilewsky, 1970

Drymatus Motschulsky, 1862

Eucheila Dejean, 1829

Eurycoleus Chaudoir, 1848

Eurydera Laporte, 1831

Formosiella Jedlicka, 1951

Gidda Andrewes, 1920

Glyphodactyla Chaudoir, 1837

Holcoderus Chaudoir, 1870

Horniulus Jedlicka, 1932

Labocephalus Chaudoir, 1848

Lelis Chaudoir, 1870

Lioptera Chaudoir, 1870

Lobodontidius Basilewsky, 1970

Lobodontulus Basilewsky, 1970

Lobodontus Chaudoir, 1842

Madecassina Jeannel, 1949

Mascarenhia Alluaud, 1933

Menarus Andrewes, 1939

Metascopus Basilewsky, 1970

Minuphloeus Darlington, 1968

Minuthodes Andrewes, 1941

Miscelus Klug, 1834

Mochtherus Schmidt-Goebel, 1846

Mormolyce Hagenbach, 1825

Mormolycina Jeannel, 1949

Neocoptodera Jeannel, 1949

Nycteis Laporte, 1835

Oreodicastes Maindron, 1905

Oxyodontus Chaudoir, 1870

Paradolichoctis Baehr, 2006

Pareurydera Jeannel, 1949

Pectinitarsus Fairmaire, 1881

Pericalus W.S.MacLeay, 1825

Philophlaeus Chaudoir, 1844

Phloeoxena Chaudoir, 1870

Pristacrus Chaudoir, 1870

Pseudoplatiella Baehr, 2012

Serrimargo Chaudoir, 1870

Sfitakantha Andrewes, 1919

Sinurus Chaudoir, 1870

Stenognathus Chaudoir, 1843

Stenotelus Bouchard, 1903

Stilboma Andrewes, 1933

Stricklandiana Bousquet, 2002

Tantillus Chaudoir, 1870

Thyreochaetus Basilewsky, 1959

Thyreopterus Dejean, 1831

Thysanotus Chaudoir, 1848

Trichocoptodera Louwerens, 1958

Xanthos Kirschenhofer, 2003

Xenitenopsis Basilewsky, 1956

Xenitenus Péringuey, 1896

Subtribe Physoderina Chaudoir, 1877

Allocota Motschulsky, 1860

Anchista Nietner, 1856

Dasiosoma Britton, 1937

Diamella Shi & Liang, 2013

Endynomena Chaudoir, 1873

Lachnoderma W.J.MacLeay, 1873

Metallanchista Shi & Liang, 2013

Orionella Jedlicka, 1964

Paraphaea Bates, 1873

Physodera Eschscholtz, 1829

Subtribe Pseudotrechina Basilewsky, 1984

Pseudotrechus Rosenhauer, 1856

Subtribe Singilina Jeannel, 1949

Singilis Rambur, 1837

Subtribe Somotrichina Mateu, 1963

Oecornis Britton, 1940

Paulianites Jeannel, 1949

Pephrica Alluaud, 1936

Somotrichus Seidlitz, 1887

Subtribe Sugimotoina Habu, 1975

Sugimotoa Habu, 1975

Subtribe Trichina Basilewsky, 1984

Trichis Klug, 1832

†Cymindoides Motschulsky, 1856

†Lebina Germar, 1813

†Miolebidia Zhang; Liu & Shangguan, 1989

†Protoscalidion L.Schaufuss, 1888

Tribe Odacanthini Laporte, 1834

Subtribe Actenonycina Bates, 1871

Actenonyx White, 1846

Subtribe Homethina Liebherr, 2016

Aeolodermus Andrewes, 1929

Cyphocoleus Chaudoir, 1877

Diplacanthogaster Liebke, 1932

Homethes Newman, 1842

Quammenis Erwin, 2000

Stenocheila Laporte, 1832

Subtribe Odacanthina Laporte, 1834

Anasis Laporte, 1867

Andrewesia Liebke, 1938

Arame Andrewes, 1919

Archicolliuris Liebke, 1931

Asios Liebke, 1933

Aulacolius Sloane, 1923

Basistichus Sloane, 1917

Clarencia Sloane, 1917

Colliuris DeGeer, 1774

Crassacantha Baehr, 1995

Cryptocolliuris Basilewsky, 1955

Deipyrodes Bousquet, 2002

Dicraspeda Chaudoir, 1863

Dobodura Darlington, 1968

Erectocolliuris Liebke, 1931

Eucolliuris Liebke, 1931

Eudalia Laporte, 1867

Gestroania Liebke, 1938

Giachinoana Baehr, 2003

Lachnothorax Motschulsky, 1862

Lasiocera Dejean, 1831

Mimocolliuris Liebke, 1933

Myrmecodemus Sloane, 1923

Neoeudalia Baehr, 2005

Odacantha Paykull, 1798

Ophionea Klug, 1821

Polydamasium Liebke, 1938

Porocara Sloane, 1917

Protocolliuris Liebke, 1931

Renneria Baehr, 1999

Smeringocera Chaudoir, 1863

Stenidia Brullé, 1834

Tricharnhemia Baehr, 2009

Subtribe Pentagonicina Bates, 1873

Parascopodes Darlington, 1968

Pentagonica Schmidt-Goebel, 1846

Scopodes Erichson, 1842

Tribe Perigonini G.Horn, 1881

Diploharpus Chaudoir, 1850

Mizotrechus Bates, 1872

Perigona Laporte, 1835

Ripogenites Basilewsky, 1954

Subfamily Licininae Bonelli, 1810

Tribe Chaetogenyini Emden, 1958

Camptotoma Reiche, 1843

Tribe Chlaeniini Brullé, 1834

Subtribe Callistina Laporte, 1834

Callistomimus Chaudoir, 1872

Callistus Bonelli, 1810

Subtribe Chlaeniina Brullé, 1834

Actodus Alluaud, 1915

Chlaenius Bonelli, 1810

Eccoptomenus Chaudoir, 1850

Ectenognathus Murray, 1858

Globulipalpus Sciaky & Facchini, 2019

Harpaglossus Motschulsky, 1858

Hololeius LaFerté-Sénectère, 1851

Holosoma Semenov, 1889

Mirachlaenius Facchini, 2011

Parachlaenius Kolbe, 1894

Perissostomus Alluaud, 1930

Procletodema Péringuey, 1898

Procletus Péringuey, 1896

Rhopalomelus Boheman, 1848

Sphodroschema Alluaud, 1930

Stenoodes Basilewsky, 1953

Straneomelus Sciaky & Facchini, 2019

Stuhlmannium Kolbe, 1894

Viridagonum Lassalle, 2015

†Rhopalochlaenius Zhang; Sun & Zhang, 1994

Tribe Licinini Bonelli, 1810

Subtribe Dicaelina Laporte, 1834

Dicaelus Bonelli, 1813

Diplocheila Brullé, 1835

Subtribe Dicrochilina Ball, 1992

Dicrochile Guérin-Méneville, 1846

Subtribe Lestignathina Ball, 1992

Atrotus Péringuey, 1896

Dilonchus Andrewes, 1936

Hormacrus Sloane, 1898

Lacordairia Laporte, 1867

Lestignathus Erichson, 1842

Microferonia Blackburn, 1890

Microzargus Sciaky & Facchini, 1997

Platylytron W.J.MacLeay, 1873

Siagonyx W.J.MacLeay, 1871

Zargus Wollaston, 1854

Subtribe Licinina Bonelli, 1810

Badister Clairville, 1806

Colpostoma Semenov, 1889

Derostichus Motschulsky, 1860

Eurygnathus Wollaston, 1854

Eutogeneius Solier, 1849

Licinus Latreille, 1802

Omestes Andrewes, 1933

Physolaesthus Chaudoir, 1850

Tribe Oodini LaFerté-Sénectère, 1851

Subtribe Dercylina Sloane, 1923

Dercylus Laporte, 1832

Subtribe Geobaenina Péringuey, 1896

Geobaenus Dejean, 1829

Subtribe Melanchitonina Jeannel, 1948

Dicaelindus W.S.MacLeay, 1825

Melanchiton Andrewes, 1940

Melanchrous Andrewes, 1940

Subtribe Oodina LaFerté-Sénectère, 1851

Acanthoodes Basilewsky, 1953

Acutosternus Lecordier & Girard, 1988

Adelopomorpha Heller, 1916

Anatrichis LeConte, 1853

Bamaroodes Gueorguiev, 2014

Brachyodes Jeannel, 1949

Chaetocrepis Chaudoir, 1857

Coptocarpus Chaudoir, 1857

Dercylinus Chaudoir, 1883

Evolenes LeConte, 1853

Holcocoleus Chaudoir, 1883

Hoplolenus LaFerté-Sénectère, 1851

Lobatodes Basilewsky, 1956

Lonchosternus LaFerté-Sénectère, 1851

Macroprotus Chaudoir, 1878

Megaloodes Lesne, 1896

Microodes Jeannel, 1949

Miltodes Andrewes, 1922

Nanodiodes Bousquet, 1996

Neoodes Basilewsky, 1953

Nothoodes Gueorguiev & Liang, 2020

Oodes Bonelli, 1810

Oodinus Motschulsky, 1865

Orthocerodus Basilewsky, 1946

Polychaetus Chaudoir, 1882

Prionognathus LaFerté-Sénectère, 1851

Protopidius Basilewsky, 1949

Pseudoodes Gueorguiev & Liang, 2020

Pseudosphaerodes Jeannel, 1949

Simous Chaudoir, 1882

Sphaerodinus Jeannel, 1949

Sphoerodes Chaudoir, 1883

Stenocrepis Chaudoir, 1857

Sundaoodes Gueorguiev & Liang, 2020

Systolocranius Chaudoir, 1857

Thaioodes Gueorguiev, 2014

Thryptocerus Chaudoir, 1878

Trichopalpoodes B.Gueorguiev & J.Schmidt, 2016

Subfamily Loricerinae Bonelli, 1810

Loricera Latreille, 1802

Subfamily Melaeninae Csiki, 1933

Cymbionotum Baudi di Selve, 1864

Melaenus Dejean, 1831

Subfamily Migadopinae Chaudoir, 1861

Tribe Amarotypini Erwin, 1985

Amarotypus Bates, 1872

Migadopiella Baehr, 2009

Tribe Migadopini Chaudoir, 1861

Antarctonomus Chaudoir, 1861

Aquilex Moret, 1989

Calathosoma Jeannel, 1938

Calyptogonia Sloane, 1920

Decogmus Sloane, 1915

Dendromigadops Baehr, 2013

Lissopterus G.R.Waterhouse, 1843

Loxomerus Chaudoir, 1842

Migadopidius Jeannel, 1938

Migadops G.R.Waterhouse, 1842

Monolobus Solier, 1849

Nebriosoma Laporte, 1867

Pseudomigadops Jeannel, 1938

Rhytidognathus Chaudoir, 1861

Stichonotus Sloane, 1910

Taenarthrus Broun, 1914

Subfamily Nebriinae Laporte, 1834

Tribe Cicindini Csiki, 1927

Archaeocindis Kavanaugh & Erwin, 1991

Cicindis Bruch, 1908

Tribe Nebriini Laporte, 1834

Leistus Frölich, 1799

Nebria Latreille, 1802

Nippononebria Ueno, 1955

†Archaeonebria Kavanaugh & J.Schmidt, 2019

†Ledouxnebria Deuve, 1998

Tribe Notiokasiini Kavanaugh & Nègre, 1983

Notiokasis Kavanaugh & Nègre, 1983

Tribe Notiophilini Motschulsky, 1850

Notiophilus Duméril, 1805

Tribe Opisthiini Dupuis, 1912

Opisthius Kirby, 1837

Paropisthius Casey, 1920

Tribe Pelophilini Kavanaugh, 1996

Pelophila Dejean, 1821

Subfamily Nototylinae Bänninger, 1927

Nototylus Gemminger & Harold, 1868

Subfamily Omophroninae Bonelli, 1810

Omophron Latreille, 1802

Subfamily Orthogoniinae Schaum, 1857

Tribe Amorphomerini Sloane, 1923

Amorphomerus Sloane, 1923

Tribe Idiomorphini Bates, 1891

Subtribe Idiomorphina Bates, 1891

Idiomorphus Chaudoir, 1846

Rathymus Dejean, 1831

Strigia Brullé, 1835

Subtribe Perochnoristhina Basilewsky, 1973

Perochnoristhus Basilewsky, 1973

Tribe Orthogoniini Schaum, 1857

Subtribe Glyptina G.Horn, 1881

Glyptus Brullé, 1837

Neoglyptus Basilewsky, 1953

Subtribe Orthogoniina Schaum, 1857

Actenoncus Chaudoir, 1872

Anoncopeucus Chaudoir, 1872

Hexachaetus Chaudoir, 1872

Neoorthogonius Tian & Deuve, 2006

Nepalorthogonius Habu, 1979

Orthogonius W.S.MacLeay, 1825

Subfamily Panagaeinae Bonelli, 1810

Tribe Brachygnathini Basilewsky, 1946

Brachygnathus Perty, 1830

Tribe Panagaeini Bonelli, 1810

Subtribe Bascanina Basilewsky, 1953

Bascanus Péringuey, 1896

Subtribe Panagaeina Bonelli, 1810

Adischissus Fedorenko, 2015

Calathocosmus Emden, 1928

Cintaroa Kasahara, 1989

Cintaromorpha Häckel & Anichtchenko, 2016

Coptia Brullé, 1835

Craspedophorus Hope, 1838

Dischissus Bates, 1873

Epigraphodes Basilewsky, 1967

Epigraphus Chaudoir, 1869

Euschizomerus Chaudoir, 1850

Geobius Dejean, 1831

Micrixys LeConte, 1854

Microschemus Andrewes, 1940

Panagaeus Latreille, 1802

Paregraphus Basilewsky, 1967

Peronomerus Schaum, 1854

Psecadius Alluaud, 1911

Tefflus Leach in Samouelle, 1819

Tinoderus Chaudoir, 1879

Tinognathus Chaudoir, 1879

Trichisia Motschulsky, 1865

Tribe Peleciini Chaudoir, 1880

Subtribe Agonicina Sloane, 1920

Agonica Sloane, 1920

Pseudagonica B.Moore, 1960

Subtribe Peleciina Chaudoir, 1880

Ardistomopsis Straneo & Ball, 1989

Disphericus G.R.Waterhouse, 1842

Dyschiridium Chaudoir, 1861

Eripus Dejean, 1829

Pelecium Kirby, 1819

Stricteripus Straneo & Ball, 1989

Subfamily Patrobinae Kirby, 1837

Tribe Lissopogonini Zamotajlov, 2000

Lissopogonus Andrewes, 1923

Tribe Patrobini Kirby, 1837

Subtribe Deltomerina Chaudoir, 1872

Deltomerus Motschulsky, 1850

Subtribe Deltomerodina Zamotajlov, 2002

Deltomerodes Deuve, 1992

Subtribe Patrobina Kirby, 1837

Apatrobus Habu & Baba, 1960

Apenetretus Kurnakov, 1960

Archipatrobus Zamotajlov, 1992

Caelopenetretus Zamotajlov & Ito, 2000

Chaetapatrobus Lafer, 1996

Chinapenetretus Kurnakov, 1963

Dimorphopatrobus Casale & Sciaky, 1994

Diplous Motschulsky, 1850

Grandipenetretus Zamotajlov & Sciaky, 1999

Himalopenetretus Zamotajlov, 2002

Indopatrobus Zamotajlov & Wrase, 2006

Ledouxius Zamotajlov, 1992

Minipenetretus Zamotajlov, 2002

Minypatrobus Ueno, 1955

Naxipenetretus Zamotajlov, 1999

Parapatrobus Zamotajlov, 1992

Parapenetretus Kurnakov, 1960

Patanitretus Zamotajlov, 2002

Patrobus Dejean, 1821

Penetretus Motschulsky, 1865

Platidiolus Chaudoir, 1878

Platypatrobus Darlington, 1938

Prodiplous Zamotajlov & Sciaky, 2006

Qiangopatrobus Zamotajlov, 2002

Quasipenetretus Zamotajlov, 2002

Tibetopenetretus Zamotajlov & Sciaky, 1999

Subfamily Paussinae Latreille, 1806

Tribe Metriini LeConte, 1853

Subtribe †Kryzhanovskianina Deuve, 2019

†Kryzhanovskiana Kataev & Kirejtshuk, 2019

Metrius Eschscholtz, 1829

Sinometrius Wrase & J.Schmidt, 2006

Tribe Ozaenini Hope, 1838

Subtribe Microzaenina Deuve, 2019

Microzaena Fairmaire, 1901

Subtribe Mystropomina G.Horn, 1881

Mystropomus Chaudoir, 1848

Subtribe Ozaenina Hope, 1838

Crepidozaena Deuve, 2001

Entomoantyx Ball & McCleve, 1990

Filicerozaena Deuve, 2001

Gibbozaena Deuve, 2001

Goniotropis G.R.Gray, 1831

Inflatozaena Deuve, 2001

Mimozaena Deuve, 2001

Ozaena Olivier, 1812

Pachyteles Perty, 1830

Physea Brullé, 1835

Physeomorpha Ogueta, 1963

Platycerozaena Bänninger, 1927

Proozaena Deuve, 2001

Serratozaena Deuve, 2001

Tachypeles Deuve, 2001

Tropopsis Solier, 1849

Subtribe Pseudozaenina Sloane, 1905

Anentmetus Andrewes, 1924

Dhanya Andrewes, 1919

Eustra Schmidt-Goebel, 1846

Itamus Schmidt-Goebel, 1846

Pseudozaena Laporte, 1834

Sphaerostylus Chaudoir, 1848

Tribe Paussini Latreille, 1806

Subtribe Carabidomemnina Wasmann, 1928

Carabidomemnus Kolbe, 1924

Eohomopterus Wasmann, 1919

Subtribe Cerapterina Billberg, 1820

Arthropterus W.S.MacLeay, 1838

Cerapterus Swederus, 1788

Megalopaussus Lea, 1906

Mesarthropterus Wasmann, 1926

Subtribe Heteropaussina Janssens, 1950

Heteropaussus J.Thomson, 1860

Subtribe Homopterina Wasmann, 1920

Homopterus Westwood, 1841

Subtribe Paussina Latreille, 1806

Allorhopalus Nagel, 2017

Ceratoderus Westwood, 1841

Euplatyrhopalus Desneux, 1905

Lebioderus Westwood, 1838

Leleupaussus Luna de Carvalho, 1962

Melanospilus Westwood, 1847

Paussomorphus Raffray, 1885

Paussus Linnaeus, 1775

Platyrhopalides Wasmann, 1918

Platyrhopalopsis Desneux, 1905

Platyrhopalus Westwood, 1833

Pterorhopalus Maruyama, 2011

Subtribe Pentaplatarthrina Jeannel, 1946

Hexaplatarthrus Jeannel, 1955

Pentaplatarthrus Westwood, 1833

Subtribe †Arthropteritina Luna de Carvalho, 1961

†Arthropterites Wasmann, 1926

†Cerapterites Wasmann, 1926

†Protocerapterus Wasmann, 1926

Subtribe †Eopaussina Luna de Carvalho, 1951

†Eopaussus Wasmann, 1926

Tribe Protopaussini Gestro, 1892

Protopaussus Gestro, 1892

Subfamily Platyninae Bonelli, 1810

Tribe Omphreini Ganglbauer, 1891

Omphreus Dejean, 1828

Tribe Platynini Bonelli, 1810

Subtribe Enoicina Basilewsky, 1985

Abacetodes Straneo, 1939

Enoicus Péringuey, 1896

Subtribe Platynina Bonelli, 1810

Achaetocephala Habu, 1975

Achaetoprothorax Habu, 1978

Aepsera Chaudoir, 1874

Agelaea Gené, 1839

Agonidium Jeannel, 1948

Agonobembix Jeannel, 1948

Agonoriascus Basilewsky, 1985

Agonorites Jeannel, 1951

Agonum Bonelli, 1810

Altagonum Darlington, 1952

Anchomenus Bonelli, 1810

Andinocolpodes Perrault, 1991

Andrewesius Andrewes, 1939

Aparupa Andrewes, 1930

Archagonum Basilewsky, 1953

Archicolpodes J.Schmidt, 2001

Arhytinus Bates, 1889

Atranodes Jedlicka, 1953

Atranus LeConte, 1847

Austroglyptolenus Roig-Juñent, 2003

Beckeria Jedlicka, 1931

Blackburnia Sharp, 1878

Bothrocolpodes Basilewsky, 1985

Bruskespar Morvan, 1998

Bruskmoal Morvan, 1998

Bryanites Valentine, 1987

Callidagonum Lorenz, 1998

Cardiomera Bassi, 1834

Catacolpodes Basilewsky, 1985

Celaenagonum Habu, 1978

Chaetagonum Burgeon, 1933

Chaetosaurus J.Schmidt, 2001

Cinctagonum Baehr, 2012

Cistelagonum Baehr, 2012

Collagonum Baehr, 1995

Colpodes W.S.MacLeay, 1825

Colpoides Jedlicka, 1931

Colpomimus Basilewsky, 1985

Colposphodrus Jedlicka, 1953

Ctenognathus Fairmaire, 1843

Cymenopterus Jeannel, 1948

Cyrtopilus Basilewsky, 1985

Dalatagonum Fedorenko, 2011

Deliaesianum Morvan, 1999

Deltocolpodes Morvan, 1992

Dendragonum Jeannel, 1948

Diacanthostylus Habu, 1978

Dicranoncoides Habu, 1978

Dicranoncus Chaudoir, 1850

Dinocolpodes J.Schmidt, 2001

Dirotus W.S.MacLeay, 1825

Dister Morvan, 2006

Dolichocolpodes Basilewsky, 1985

Dyscolus Dejean, 1831

Elliptoleus Bates, 1882

Epicolpodes Basilewsky, 1985

Eucolpodes Jeannel, 1948

Euleptus Klug, 1833

Euplynes Schmidt-Goebel, 1846

Feroniascus Jeannel, 1951

Fortagonum Darlington, 1952

Galiciotyphlotes Assmann, 1999

Gastragonum Darlington, 1952

Glaucagonum Habu, 1978

Glyptolenoides Perrault, 1991

Glyptolenus Bates, 1878

Gyrochaetostylus Habu, 1978

Habragonum Ueno, 1964

Hannaphota Landin, 1955

Haplocolpodes Jeannel, 1951

Haplopeza Boheman, 1848

Helluocolpodes Liebherr, 2005

Hemiplatynus Casey, 1920

Henvelik Morvan, 1999

Herculagonum Baehr, 2002

Hikosanoagonum Habu, 1954

Idiagonum Darlington, 1952

Idiastes Andrewes, 1931

Idiocolpodes Basilewsky, 1985

Incagonum Liebherr, 1994

Iridagonum Darlington, 1952

Ischnagonum Kasahara & Satô, 1997

Jocqueius Basilewsky, 1988

Jujiroa Ueno, 1952

Kalchdigor Morvan, 1999

Kar Morvan, 1998

Karnes Morvan, 2010

Kaszabellus Jedlicka, 1954

Kiwiplatynus Larochelle & Larivière, 2021

Klapperichella Jedlicka, 1956

Kuceraianum Morvan, 2002

Kupeplatynus Larochelle & Larivière, 2021

Laevagonum Darlington, 1952

Lassalleianum Morvan, 1999

Lepcha Andrewes, 1930

Leptagonum Kolbe, 1897

Leptocolpodes Basilewsky, 1985

Letouzeya Bruneau de Miré, 1982

Liagonum Jeannel, 1948

Liamegalonychus Basilewsky, 1963

Liocolpodes Basilewsky, 1985

Lissagonum Habu, 1978

Lithagonum Darlington, 1952

Lobocolpodes Basilewsky, 1985

Lorostema Motschulsky, 1865

Loxocrepis Eschscholtz, 1829

Lucicolpodes J.Schmidt, 2000

Maculagonum Darlington, 1952

Maoriplatynus Larochelle & Larivière, 2021

Meleagros Kirschenhofer, 1999

Mesocolpodes Basilewsky, 1985

Metacolpodes Jeannel, 1948

Mexisphodrus Barr, 1965

Montagonum Darlington, 1952

Mooreagonum Baehr, 2016

Morimotoidius Habu, 1954

Nebriagonum Darlington, 1952

Negreum Habu, 1958

Neobatenus Jeannel, 1948

Neocolpodes Jeannel, 1948

Neodendragonum Basilewsky, 1953

Neomegalonychus Jeannel, 1948

Nesiocolpodes Jeannel, 1948

Nipponagonum Habu, 1978

Notagonum Darlington, 1952

Notocolpodes Basilewsky, 1985

Notoplatynus B.Moore, 1985

Nymphagonum Habu, 1978

Olisthopus Dejean, 1828

Oncostylus Habu, 1978

Onotokiba Alluaud, 1927

Onycholabis Bates, 1873

Onypterygia Dejean, 1831

Orophicus Alluaud, 1925

Orthotrichus Peyron, 1856

Oxygonium Basilewsky, 1951

Oxypselaphus Chaudoir, 1843

Pachybatenus Basilewsky, 1973

Pachyferonia Jeannel, 1951

Paracolpodes Basilewsky, 1985

Paraliagonum Basilewsky, 1957

Paramegalonychus Basilewsky, 1953

Paranchodemus Habu, 1978

Paranchus Lindroth, 1974

Paraplatynus Baehr, 2016

Pawgammm Morvan, 1999

Platyagonum Habu, 1978

Platynus Bonelli, 1810

Plaumannium Liebke, 1939

Plicagonum Darlington, 1952

Ponapagonum Darlington, 1970

Potamagonum Darlington, 1952

Praepristus Kirschenhofer, 1999

Promecoptera Dejean, 1831

Promegalonychus Basilewsky, 1953

Prophenorites Basilewsky, 1985

Prosphodrus Britton, 1959

Protocolpodes Basilewsky, 1985

Pseudanchomenus Tarnier, 1860

Pseudobatenus Basilewsky, 1951

Pseudomegalonychus Basilewsky, 1950

Rhadine LeConte, 1846

Rupa Jedlicka, 1935

Scotagonum Habu, 1978

Sericoda Kirby, 1837

Shibataia Habu, 1978

Sinocolpodes J.Schmidt, 2001

Skoeda Morvan, 1995

Skorlagad Morvan, 1999

Skouedhirraad Morvan, 1999

Sophroferonia Alluaud, 1933

Speagonum B.Moore, 1977

Speocolpodes Barr, 1974

Speokokosia Alluaud, 1932

Sperkanhir Morvan, 2010

Stenocnemus Mannerheim, 1837

Straneoa Basilewsky, 1953

Syletor Tschitscherine, 1899

Takasagoagonum Habu, 1977

Tanystoma Motschulsky, 1845

Tarsagonum Darlington, 1952

Tetraleucus Casey, 1920

Tostkar Morvan, 1998

Trogloagonum Casale, 1982

Tuiplatynus Larochelle & Larivière, 2021

Vitagonum B.Moore, 1999

Vulcanophilus Heller, 1898

Xestagonum Habu, 1978

†Praeanchodemus J.Schmidt; Göpel & Will, 2017

Tribe Sphodrini Laporte, 1834

Subtribe Atranopsina Baehr, 1982

Amaroschema Jeannel, 1943

Amphimasoreus Piochard de la Brûlerie, 1875

Atranopsis Baehr, 1982

Broter Andrewes, 1923

Gomerina Bolivar y Pieltain, 1940

Paraeutrichopus Mateu, 1954

Platyderus Stephens, 1827

Pseudomyas Uyttenboogaart, 1929

Pseudoplatyderus Bolivar y Pieltain, 1940

Subtribe Calathina Laporte, 1834

Calathus Bonelli, 1810

Thermoscelis Putzeys, 1873

Subtribe Dolichina Brullé, 1834

Acalathus Semenov, 1889

Anchomenidius Heyden, 1880

Casaleius Sciaky & Wrase, 1998

Dolichus Bonelli, 1810

Doliodactyla Sciaky & Wrase, 1998

Morphodactyla Semenov, 1889

Xestopus Andrewes, 1937

Subtribe Pristosiina Lindroth, 1956

Pristosia Motschulsky, 1865

Subtribe Sphodrina Laporte, 1834

Calathidius Putzeys, 1873

Cephalosdrophus Lassalle & Marcilhac, 1999

Eosphodrus Casale, 1988

Eremosphodrus Semenov, 1909

Himalosphodrus Casale, 1988

Hystricosphodrus Casale & Giachino, 2004

Ifridytes Deuve & Queinnec, 1994

Laemostenus Bonelli, 1810

Licinopsis Bedel, 1899

Miquihuana Barr, 1982

Pseudotaphoxenus L.Schaufuss, 1865

Reflexisphodrus Casale, 1988

Sphodropsis Seidlitz, 1887

Sphodrus Clairville, 1806

Stenolepta Semenov, 1889

Taphoxenus Motschulsky, 1850

Subtribe Synuchina Lindroth, 1956

Nipponosynuchus Morita, 1998

Parabroscus Lindroth, 1956

Synuchus Gyllenhal, 1810

Trephionus Bates, 1883

Subfamily Promecognathinae LeConte, 1853

Tribe Axinidiini Basilewsky, 1963

Axinidium Sturm, 1843

Holaxinidium Basilewsky, 1963

Metaxinidium Basilewsky, 1963

Paraxinidium Basilewsky, 1963

Tribe Dalyatini Mateu, 2002

Dalyat Mateu, 2002

Tribe Promecognathini LeConte, 1853

Promecognathus Chaudoir, 1846

Tribe †Palaeoaxinidiini McKay, 1991

†Palaeoaxinidium McKay, 1991

Subfamily Pseudomorphinae Hope, 1838

Adelotopus Hope, 1836

Cainogenion Notman, 1925

Cryptocephalomorpha Ritsema, 1875

Guyanemorpha Erwin, 2013

Manumorpha Erwin & Geraci, 2008

Notopseudomorpha Baehr, 1997

Paussotropus C.O.Waterhouse, 1877

Pseudomorpha Kirby, 1823

Samiriamorpha Erwin & Geraci, 2008

Sphallomorpha Westwood, 1840

Tuxtlamorpha Erwin & Geraci, 2008

Yasunimorpha Erwin & Geraci, 2008

Subfamily Psydrinae LeConte, 1853

Tribe Gehringiini Darlington, 1933

Subtribe Gehringiina Darlington, 1933

Gehringia Darlington, 1933

Subtribe Helenaeina Deuve, 2007

Afrogehringia Baehr; Schüle & Lorenz, 2009

Helenaea Schatzmayr & Koch, 1934

Tribe Moriomorphini Sloane, 1890

Subtribe Amblytelina Blackburn, 1892

Amblytelus Erichson, 1842

Dystrichothorax Blackburn, 1892

Epelyx Blackburn, 1892

Mecyclothorax Sharp, 1903

Meonis Laporte, 1867

Paratrichothorax Baehr, 2004

Pseudamblytelus Baehr, 2004

Raphetis B.Moore, 1963

Selenochilus Chaudoir, 1878

Trichamblytelus Baehr, 2004

Subtribe Moriomorphina Sloane, 1890

Celanida Laporte, 1867

Melisodera Westwood, 1835

Molopsida White, 1846

Moriodema Laporte, 1867

Moriomorpha Laporte, 1867

Neonomius B.Moore, 1963

Pterogmus Sloane, 1920

Rhaebolestes Sloane, 1903

Rossjoycea Liebherr, 2011

Sitaphe B.Moore, 1963

Spherita Liebherr, 2020

Tarastethus Sharp, 1883

Teraphis Laporte, 1867

Theprisa B.Moore, 1963

Trephisa B.Moore, 1963

Trichopsida Larochelle & Larivière, 2013

Subtribe Tropopterina Sloane, 1898

Pharetis Liebherr, 2020

Tropopterus Solier, 1849

Tribe Psydrini LeConte, 1853

Laccocenus Sloane, 1890

Nomius Laporte, 1835

Psydrus LeConte, 1846

Subfamily Pterostichinae Bonelli, 1810

Tribe Chaetodactylini Tschitscherine, 1903

Chaetodactyla Tschitscherine, 1897

Tribe Cnemalobini Germain, 1911

Cnemalobus Guérin-Méneville, 1838

Tribe Cratocerini Lacordaire, 1854

Subtribe Catapieseina Bates, 1882

Catapiesis Solier, 1835

Homalomorpha Brullé, 1837

Subtribe Cratocerina Lacordaire, 1854

Brachidius Chaudoir, 1852

Cratocerus Dejean, 1829

Oxyglychus Straneo, 1938

Subtribe Drimostomatina Chaudoir, 1873

Abacaelostus Straneo, 1952

Andrewesinulus Straneo, 1938

Apsidocnemus Alluaud, 1936

Barylaus Liebherr, 1985

Caecocaelus Straneo, 1949

Caelostomus W.S.MacLeay, 1825

Camptogenys Tschitscherine, 1899

Capabatus Csiki, 1930

Crenulostrigus Straneo, 1942

Cyrtolaus Bates, 1882

Dactyleurys Tschitscherine, 1899

Dactylinius Straneo, 1941

Diachipteryx Alluaud, 1925

Diceromerus Chaudoir, 1873

Dromistomus Jeannel, 1948

Drymonaxus Straneo, 1941

Feostoma Straneo, 1941

Hemitelestus Alluaud, 1895

Hoplizomenus Chaudoir, 1873

Leleuporites Straneo, 1960

Madapelmus Dajoz, 1985

Monodryxus Straneo, 1942

Pachycaecus Straneo, 1971

Pachyroxochus Straneo, 1942

Platyxythrius Lorenz, 1998

Stegazopteryx Will, 2004

Stomonaxellus Tschitscherine, 1901

Strigomerodes Straneo, 1939

Strigomerus Chaudoir, 1873

Trichillinus Straneo, 1938

Tribe Microcheilini Jeannel, 1948

Microcheila Brullé, 1835

Tribe Morionini Brullé, 1837

Buderes Murray, 1857

Hyperectenus Alluaud, 1935

Hyperion Laporte, 1834

Megamorio Chaudoir, 1880

Morion Latreille, 1810

Morionidius Chaudoir, 1880

Moriosomus Motschulsky, 1855

Platynodes Westwood, 1847

Stereostoma Murray, 1857

Tribe Pterostichini Bonelli, 1810

Subtribe Abacetina Chaudoir, 1873

Abacaecus Allegro & Giachino, 2020

Abacetus Dejean, 1828

Aristopus LaFerté-Sénectère, 1853

Celioinkosa Straneo, 1951

Chlaeminus Motschulsky, 1865

Colpodichius Straneo, 1952

Cosmodiscus Sloane, 1907

Cyrtomoscelis Chaudoir, 1874

Distrigidius Jeannel, 1948

Ecnomolaus Bates, 1892

Haptoderidius Straneo, 1975

Haptoderodes Straneo, 1986

Holconotus Schmidt-Goebel, 1846

Inkosa Péringuey, 1926

Mateuellus Deuve, 1990

Metabacetus Bates, 1892

Metaxellus Straneo, 1960

Metaxys Chaudoir, 1857

Novillidius Straneo, 1941

Oodinkosa Straneo, 1939

Ophonichius Straneo, 1942

Pediomorphus Chaudoir, 1878

Pioprosopus Tschitscherine, 1899

Pollicobius Vinson, 1939

Prostalomus Basilewsky, 1950

Pseudabacetus Burgeon, 1935

Pterostillichus Straneo, 1949

Rhagadillius Straneo, 1951

Tiferonia Darlington, 1962

Trachelocyphoides Straneo, 1942

Trachelocyphus Tschitscherine, 1900

Subtribe Euchroina Chaudoir, 1874

Abacillius Straneo, 1949

Abacillodes Straneo, 1988

Abaris Dejean, 1831

Apsaustodon Tschitscherine, 1901

Argutoridius Chaudoir, 1876

Blennidus Motschulsky, 1866

Blenniventer Straneo, 1986

Bothynoproctus Tschitscherine, 1900

Cedrorum Borges & A.Serrano, 1993

Cephalostichus Straneo, 1977

Cynthidia Chaudoir, 1874

Euchroa Brullé, 1835

Eumara Tschitscherine, 1901

Gastrogmus Sloane, 1915

Haplobothynus Tschitscherine, 1901

Litarthrum Sloane, 1915

Lobobrachus Sharp, 1885

Marsyas Putzeys, 1845

Meropalpus Tschitscherine, 1900

Microcephalus Dejean, 1828

Neotalus Will, 2002

Oribazus Chaudoir, 1874

Orthomus Chaudoir, 1838

Pachythecus Chaudoir, 1874

Paniestichus Will, 2011

Parorthomus B.Gueorguiev; Wrase & Farkac, 2014

Phaenaulax Tschitscherine, 1898

Prosopogmus Chaudoir, 1865

Pseudabarys Chaudoir, 1874

Setalimorphus Sloane, 1895

Setalis Laporte, 1867

Simodontus Chaudoir, 1843

Trirammatus Chaudoir, 1835

Subtribe Loxandrina Erwin & Sims, 1984

Cerabilia Laporte, 1867

Oxycrepis Reiche, 1843

Zeodera Laporte, 1867

Subtribe Metiina Straneo, 1951

Abropus G.R.Waterhouse, 1842

Antarctiola Straneo, 1977

Feroniola Tschitscherine, 1900

Kuschelinus Straneo, 1963

Metius Curtis, 1839

Subtribe Pterostichina Bonelli, 1810

Abacidus LeConte, 1863

Abacoleptus Fauvel, 1903

Abacomorphus Chaudoir, 1878

Abacophrastus Will, 2011

Abacops Tschitscherine, 1902

Abax Bonelli, 1810

Allotriopus Bates, 1882

Analoma Darlington, 1971

Apophylon B.Gueorguiev & Sciaky, 2015

Aristochroa Tschitscherine, 1898

Aristochroodes Marcilhac, 1993

Aulacopodus Britton, 1940

Basilewskya Straneo, 1948

Camptoscelis Dejean, 1828

Castelnaudia Tschitscherine, 1891

Catadromus W.S.MacLeay, 1825

Chaetauchenium Tschitscherine, 1900

Chalcochrous Chaudoir, 1838

Cophosomorpha Tschitscherine, 1891

Cratoferonia Tschitscherine, 1902

Cratogaster Blanchard, 1843

Cuneipectus Sloane, 1907

Cyclotrachelus Chaudoir, 1838

Cyrtoderus Hope, 1842

Darodilia Laporte, 1867

Delinius Westwood, 1864

Eucamptognathus Chaudoir, 1837

Eudromus Klug, 1835

Euryabax Fauvel, 1903

Euryaptus Bates, 1892

Eurypercus Jeannel, 1948

Eurystomis Chaudoir, 1878

Gastrellarius Casey, 1918

Gourlayia Britton, 1964

Harpostomus Chaudoir, 1856

Henrotiochoromus Busulini, 1958

Henrotius Jeannel, 1953

Holcaspis Chaudoir, 1865

Hybothecus Chaudoir, 1874

Leiolesticus Roux; Lassalle & Dubault, 2016

Lesticus Dejean, 1828

Licentius Jedlicka, 1939

Liopasa Tschitscherine, 1901

Lophoglossus LeConte, 1853

Loxogenius Sloane, 1907

Mecynognathus W.J.MacLeay, 1873

Megadromus Motschulsky, 1866

Molopidius Jeannel, 1942

Molopinus Jeannel, 1948

Molops Bonelli, 1810

Myas Sturm, 1826

Neoferonia Britton, 1940

Nesites Andrewes, 1939

Nirmala Andrewes, 1930

Notabax B.Moore, 1976

Notolestus Sloane, 1894

Notonomus Chaudoir, 1865

Nurus Motschulsky, 1866

Oberthueria Vuillet, 1911

Ogmophora Tschitscherine, 1896

Onawea Johns, 2007

Oscadytes Lagar Mascaro, 1975

Pachymorphus Chaudoir, 1838

Paranurus Tschitscherine, 1901

Pareuryaptus Dubault; Lassalle & Roux, 2008

Parhypates Motschulsky, 1866

Pedius Motschulsky, 1850

Percolaus Bates, 1882

Percus Bonelli, 1810

Peyrieraselus Deuve, 1981

Piesmus LeConte, 1846

Platycoelus Blanchard, 1843

Platysmodes Fauvel, 1903

Plocamostethus Britton, 1940

Poecilinus Jeannel, 1948

Poecilus Bonelli, 1810

Pseudoceneus Tschitscherine, 1891

Pterostichus Bonelli, 1810

Rhabdotus Chaudoir, 1865

Rhytiferonia Darlington, 1962

Rhytisternus Chaudoir, 1865

Sarticus Motschulsky, 1866

Secatophus Laporte, 1867

Setalidius Chaudoir, 1878

Speluncarius Reitter, 1886

Speomolops Patrizi, 1955

Sphodrosomus Perroud, 1864

Stenochoromus L.Miller, 1866

Stereocerus Kirby, 1837

Sthenocranion Tschitscherine, 1899

Stomis Clairville, 1806

Straneostichus Sciaky, 1994

Styracoderus Chaudoir, 1874

Tanythrix Schaum, 1858

Taphoxomimus Straneo, 1975

Tapinopterus Schaum, 1858

Teratotarsa Tschitscherine, 1893

Teropha Laporte, 1867

Trichosternus Chaudoir, 1865

Trigonaptus Fedorenko, 2020

Trigonotoma Dejean, 1828

Troglorites Jeannel, 1919

Tropidocerus Chaudoir, 1878

Typhlochoromus Moczarski, 1913

Wahlbergiana Bousquet, 2002

Xenion Tschitscherine, 1902

Zariquieya Jeannel, 1924

Zeopoecilus Sharp, 1886

Tribe Zabrini Bonelli, 1810

Subtribe Amarina Zimmermann, 1832

Amara Bonelli, 1810

Pseudamara Lindroth, 1968

Subtribe Zabrina Bonelli, 1810

Zabrus Clairville, 1806

Subfamily Rhysodinae Laporte, 1840

Tribe Clinidiini R.T. & J.R.Bell, 1978

Clinidium Kirby, 1830

Grouvellina R.T. & J.R.Bell, 1978

Rhyzodiastes Fairmaire, 1895

Tribe Dhysorini R.T. & J.R.Bell, 1978

Dhysores Grouvelle, 1903

Neodhysores R.T. & J.R.Bell, 1978

Tangarona R.T. & J.R.Bell, 1982

Tribe Leoglymmiini R.T. & J.R.Bell, 1978

Leoglymmius R.T. & J.R.Bell, 1978

Tribe Medisorini R.T. & J.R.Bell, 1987

Medisores R.T. & J.R.Bell, 1987

Tribe Omoglymmiini R.T. & J.R.Bell, 1978

Arrowina R.T. & J.R.Bell, 1978

Omoglymmius Ganglbauer, 1891

Plesioglymmius R.T. & J.R.Bell, 1978

Shyrodes Grouvelle, 1903

Srimara R.T. & J.R.Bell, 1978

Xhosores R.T. & J.R.Bell, 1978

Yamatosa R.T. & J.R.Bell, 1979

Tribe Rhysodini Laporte, 1840

Kaveinga R.T. & J.R.Bell, 1978

Kupeus R.T. & J.R.Bell, 1982

Rhysodes Germar, 1822

Tribe Sloanoglymmiini R.T. & J.R.Bell, 1991

Sloanoglymmius R.T. & J.R.Bell, 1991

Subfamily Scaritinae Bonelli, 1810

Tribe Clivinini Rafinesque, 1815

Subtribe Androzelmina R.T.Bell, 1998

Androzelma Dostal, 1993

Subtribe Ardistomina Putzeys, 1867

Ardistomis Putzeys, 1846

Aspidoglossa Putzeys, 1846

Kearophus Dajoz, 2004

Semiardistomis Kult, 1950

Subtribe Clivinina Rafinesque, 1815

Afrosyleter Basilewsky, 1959

Ancus Putzeys, 1867

Basilewskyana Kult, 1959

Brachypelus Putzeys, 1867

Cameroniola Baehr, 2000

Climax Putzeys, 1861

Clivina Latreille, 1802

Clivinarchus Sloane, 1896

Cryptomma Putzeys, 1846

Lachenus Putzeys, 1846

Leleuporella Basilewsky, 1956

Nannoryctes Baehr, 2000

Nyctosyles Putzeys, 1867

Orictites Andrewes, 1931

Platysphyrus Sloane, 1905

Psilidius Jeannel, 1957

Pyramoides Bousquet, 2002

Rhysocara Sloane, 1916

Rubidiclivina Baehr, 2015

Rugiluclivina Balkenohl, 1996

Semiclivina Kult, 1947

Sinesetosa Balkenohl, 1996

Syleter Andrewes, 1941

Thliboclivina Kult, 1959

Trilophidius Jeannel, 1957

Trilophus Andrewes, 1927

Trogloclivina Deuve, 2003

Whiteheadiana Perrault, 1994

Subtribe Forcipatorina Bänninger, 1938

Camptidius Putzeys, 1867

Camptodontus Dejean, 1826

Forcipator Maindron, 1904

Kultianella Perrault, 1994

Mesus Chevrolat, 1858

Obadius Burmeister, 1875

Oxygnathopsis Louwerens, 1953

Oxygnathus Dejean, 1826

Scolyptus Putzeys, 1861

Stratiotes Putzeys, 1846

Subtribe Reicheiina Jeannel, 1957

Alpiodytes Jeannel, 1957

Asioreicheia Bulirsch & Magrini, 2014

Catalanodytes Sciaky, 1989

Dalmatoreicheia Magrini & Bulirsch, 2005

Dimorphoreicheia Magrini; Fancello & Leo, 2002

Galicioreicheia Felix & Bulirsch, 2015

Guiodytes Tian, 2013

Gymnetoreicheia Magrini; Fancello & Casale, 2019

Iberodytes Jeannel, 1949

Ichnusodytes Magrini; Fancello & Onnis, 2019

Italodytes G.Müller, 1938

Kenyoreicheia Bulirsch & Magrini, 2007

Laoreicheia Balkenohl, 2005

Madagascareicheia Magrini & Bulirsch, 2009

Malagasyreicheia Magrini; Bulirsch & Fancello, 2021

Orientoreicheia Bulirsch & Hurka, 1994

Oxydrepanus Putzeys, 1867

Parareicheia Jeannel, 1957

Reicheadella Reitter, 1913

Reicheia Saulcy, 1862

Reicheidius Jeannel, 1957

Sikelioreicheia Magrini & Fancello, 2019

Spelaeodytes L.Miller, 1863

Typhloreicheia Holdhaus, 1924

Subtribe Schizogeniina Dostal, 2017

Baehrogenius Dostal, 2017

Coryza Putzeys, 1867

Halocoryza Alluaud, 1919

Lophocoryza Alluaud, 1941

Paracoryza Basilewsky, 1952

Psammocoryza Hogan, 2006

Schizogenius Putzeys, 1846

Subtribe Sparostesina Dostal, 2017

Bohemaniella Bousquet, 2002

Pseudoclivina Kult, 1947

Sparostes Putzeys, 1867

Tribe Corintascarini Basilewsky, 1973

Corintascaris Basilewsky, 1952

Tribe Dyschiriini Kolbe, 1880

Akephorus LeConte, 1852

Caledyschirius Bulirsch, 2010

Clivinopsis Bedel, 1895

Cribrodyschirius Bruneau de Miré, 1952

Dyschirius Bonelli, 1810

Neodyschirius Kult, 1954

Reicheiodes Ganglbauer, 1891

Setodyschirius Fedorenko, 1996

Striganoviella Fedorenko, 2012

Torretassoa Schatzmayr & Koch, 1933

†Dyschiriomimus Iablokoff-Khnzorian, 1960

Tribe Salcediini Alluaud, 1930

Subtribe Salcediina Alluaud, 1930

Salcedia Fairmaire, 1899

Subtribe Solenogenyina R.T.Bell, 1998

Holoprizus Putzeys, 1867

Solenogenys Westwood, 1859

Tribe Scaritini Bonelli, 1810

Subtribe Carenina W.J.MacLeay, 1887

Carenidium Chaudoir, 1868

Carenum Bonelli, 1813

Epilectus Blackburn, 1888

Euryscaphus W.J.MacLeay, 1865

Laccopterum W.J.MacLeay, 1878

Monocentrum Chaudoir, 1868

Mouhotia Laporte, 1862

Neocarenum Laporte, 1867

Neoscaphus Sloane, 1888

Philoscaphus W.J.MacLeay, 1871

Scaraphites Westwood, 1842

Trichocarenum Blackburn, 1892

Subtribe Pasimachina Putzeys, 1867

Pasimachus Bonelli, 1813

Subtribe Scapterina Putzeys, 1867

Acanthoscelis Dejean, 1825

Oxylobus Chaudoir, 1855

Parathlibops Basilewsky, 1958

Passalidius Chaudoir, 1863

Scapterus Dejean, 1826

Steganomma W.J.MacLeay, 1887

Thlibops Putzeys, 1867

Subtribe Scaritina Bonelli, 1810

Anomophaenus Fauvel, 1882

Antilliscaris Lorenz, 1998

Baenningeria Reichardt, 1976

Coptolobus Chaudoir, 1857

Crepidopterus Chaudoir, 1855

Cryptoscaphus Chaudoir, 1855

Dinoscaris Alluaud, 1902

Distichus Motschulsky, 1858

Dyscaris Bänninger, 1940

Dyscherinus Jeannel, 1955

Dyscherus Chaudoir, 1855

Geoscaptus Chaudoir, 1855

Glyptogrus Bates, 1875

Gnaphon Andrewes, 1920

Haplogaster Chaudoir, 1879

Haplotrachelus Chaudoir, 1855

Macromorphus Chaudoir, 1857

Madascaris Bänninger, 1938

Mamboicus Bates, 1886

Mecynoscaris Alluaud, 1930

Menigius Chaudoir, 1879

Neochryopus Bänninger, 1932

Ochyropus Schiödte, 1847

Pachyodontus Chaudoir, 1879

Paradyscherus Basilewsky, 1971

Pilades Heyne, 1895

Prodyscherodes Jeannel, 1955

Prodyscherus Jeannel, 1946

Scarites Fabricius, 1775

Storthodontus Chaudoir, 1855

Tapinoscaris Jeannel, 1946

Tibioscarites Bänninger, 1929

Tonkinoscaris Bänninger, 1956

Typhloscaris Kuntzen, 1914

Subfamily Siagoninae Bonelli, 1813

Tribe Enceladini G.Horn, 1881

Enceladus Bonelli, 1813

Tribe Siagonini Bonelli, 1813

Luperca Laporte, 1840

Siagona Latreille, 1804

Subfamily Trechinae Bonelli, 1810

Tribe Bembidarenini Maddison et al., 2019

Andinodontis Erwin & Toledano, 2010

Argentinatachoides Sallanave; Erwin & Roig-Juñent, 2008

Bembidarenas Erwin, 1972

Tasmanitachoides Erwin, 1972

Tribe Bembidiini Stephens, 1827

Subtribe Anillina Jeannel, 1937

Afranillus Giachino, 2015

Afrodipnus Giachino, 2015

Angustanillus Baehr & Main, 2016

Anillaspis Casey, 1918

Anillinus Casey, 1918

Anillodes Jeannel, 1963

Anillopsidius Coiffait, 1969

Anillopsis Jeannel, 1937

Anillotarsus Mateu, 1980

Anillus Jacquelin du Val, 1851

Argiloborus Jeannel, 1937

Austranillus Giachino, 2005

Bafutyphlus Bruneau de Miré, 1986

Bhutanillus M.E.Schmid, 1975

Bulirschia Giachino, 2008

Bylibaraphanus Giachino; Eberhard & Perina, 2021

Caeconannus Jeannel, 1963

Caecoparvus Jeannel, 1937

Carayonites Bruneau de Miré, 1986

Cryptocharidius M.Etonti & Mateu, 1992

Cryptorites Jeannel, 1950

Dicropterus Ehlers, 1883

Elgonotyphlus Sciaky & Zaballos, 1993

Erwinanillus Giachino; Eberhard & Perina, 2021

Externanillus Baehr & Main, 2016

Geocharidius Jeannel, 1963

Geocharis Ehlers, 1883

Gilesdytes Giachino; Eberhard & Perina, 2021

Gracilanillus Baehr & Main, 2016

Gregorydytes Giachino; Eberhard & Perina, 2021

Hesperanillus Baehr & Main, 2016

Honduranillus Zaballos, 1997

Hygranillus B.Moore, 1980

Hypodipnites Jeannel, 1963

Hypotyphlus Jeannel, 1937

Iason Giachino & Vailati, 2011

Iberanillus Español, 1971

Illaphanus W.J.MacLeay, 1865

Kimberleytyphlus Giachino; Eberhard & Perina, 2021

Leleupanillus Basilewsky, 1976

Magnanillus Baehr, 2017

Malagasydipnus Giachino, 2008

Malagasytyphlus Giachino, 2008

Megastylulus Giachino & Sciaky, 2003

Mexanillus Vigna Taglianti, 1973

Microdipnites Jeannel, 1957

Microdipnodes Basilewsky, 1960

Microdipnus Jeannel, 1937

Microtyphlus Linder, 1863

Mystroceridius Reichardt, 1972

Neodipnus Jeannel, 1957

Neoillaphanus Giachino; Eberhard & Perina, 2021

Nesamblyops Jeannel, 1937

Nothanillus Jeannel, 1962

Orthotyphlus Mateu & Zaballos, 1998

Paranillopsis Cicchino & Roig-Juñent, 2001

Paranillus Jeannel, 1949

Parillaphanus Baehr, 2018

Parvocaecus Coiffait, 1956

Pelocharis Jeannel, 1960

Pelodiaetodes B.Moore, 1980

Pelodiaetus Jeannel, 1937

Pelonomites Jeannel, 1963

Perucharidius Mateu & M.Etonti, 2002

Pilbaradytes Giachino; Eberhard & Perina, 2021

Pilbaranillus Baehr & Main, 2016

Pilbaraphanus Giachino; Eberhard & Perina, 2021

Prioniomus Jeannel, 1937

Pseudanillus Bedel, 1896

Pseudillaphanus Giachino, 2005

Rhegmatobius Jeannel, 1937

Scotodipnus Schaum, 1860

Selenodipnus Jeannel, 1963

Serranillus Barr, 1995

Serratotyphlus Giachino, 2008

Stylulus L.Schaufuss, 1882

Tasmanillus Giachino, 2005

Typhlocharis Dieck, 1869

Typhlonesiotes Jeannel, 1937

Winklerites Jeannel, 1937

Zapotecanillus Sokolov, 2013

Zeanillus Jeannel, 1937

Zoianillus Sciaky, 1994

Subtribe Bembidiina Stephens, 1827

Amerizus Chaudoir, 1868

Asaphidion Gozis, 1886

Bembidion Latreille, 1802

Caecidium Ueno, 1971

Lionepha Casey, 1918

Ocys Stephens, 1828

Orzolina Machado, 1987

Sinechostictus Motschulsky, 1864

Subtribe Horologionina Jeannel, 1949

Horologion Valentine, 1932

Subtribe Lovriciina Giachino; B.Gueorguiev & Vailati, 2011

Lovricia Pretner, 1979

Neolovricia Lakota; Jalzic & J.Moravec, 2009

Paralovricia Giachino; B.Gueorguiev & Vailati, 2011

Subtribe Tachyina Motschulsky, 1862

Aenigmatachys Baehr, 2020

Afrotachys Basilewsky, 1958

Anomotachys Jeannel, 1946

Costitachys Erwin, 1974

Elaphropus Motschulsky, 1839

Kiwitachys Larochelle & Larivière, 2007

Liotachys Bates, 1871

Lymnastis Motschulsky, 1862

Meotachys Erwin, 1974

Micratopus Casey, 1914

Moirainpa Erwin, 1984

Nothoderis Boyd & Erwin, 2016

Pericompsus LeConte, 1852

Polyderidius Jeannel, 1962

Porotachys Netolitzky, 1914

Setitachys Baehr, 2016

Stigmatachys Boyd & Erwin, 2016

Straneoites Basilewsky, 1947

Tachys Dejean, 1821

Tachysbembix Erwin, 2004

Tachyta Kirby, 1837

Tachyxysta Boyd & Erwin, 2016

Subtribe Xystosomina Erwin, 1994

Erwiniana Paulsen & A.Smith, 2003

Geballusa Erwin, 1994

Gouleta Erwin, 1994

Inpa Erwin, 1978

Mioptachys Bates, 1882

Philipis Erwin, 1994

Xystosomus Schaum, 1863

Tribe Pogonini Laporte, 1834

Bedeliolus Semenov, 1900

Cardiaderus Dejean, 1828

Diodercarus Lutshnik, 1931

Diplochaetus Chaudoir, 1872

Ochtozetus Chaudoir, 1872

Olegius Komarov, 1996

Pogonistes Chaudoir, 1872

Pogonopsis Bedel, 1898

Pogonus Dejean, 1821

Sirdenus Dejean, 1828

Syrdenoidius Baehr & Hudson, 2001

Thalassotrechus Van Dyke, 1918

Tribe Sinozolini Deuve, 1997

Chaltenia Roig-Juñent & Cicchino, 2001

Phrypeus Casey, 1924

Sinozolus Deuve, 1997

Tribe Trechini Bonelli, 1810

Subtribe Aepina Fowler, 1887

Aepopsis Jeannel, 1922

Aepus Leach in Samouelle, 1819

Kenodactylus Broun, 1909

Kiwitrechus Larochelle & Larivière, 2007

Maoritrechus Brookes, 1932

Neanops Britton, 1962

Oarotrechus Townsend, 2010

Temnostega Enderlein, 1905

Thalassobius Solier, 1849

Waiputrechus Townsend, 2010

Subtribe Perileptina Sloane, 1903

Apoplotrechus Alluaud, 1915

Neoblemus Jeannel, 1923

Perileptus Schaum, 1860

Subtribe Trechina Bonelli, 1810

Accoella Ueno, 1990

Acheroniotes Lohaj & Lakota, 2010

Adriaphaenops Noesske, 1928

Aepiblemus Belousov & Kabak, 1993

Agonotrechus Jeannel, 1923

Agostinia Jeannel, 1928

Alanorites Belousov, 1998

Albanotrechus Casale & V.B.Gueorguiev, 1994

Allegrettia Jeannel, 1928

Allotrechiama Ueno, 1970

Ameroduvalius Valentine, 1952

Anchotrechus Jeannel, 1927

Andinorites Mateu & Belles, 1980

Andinotrechus Mateu, 1981

Anillidius Jeannel, 1928

Anophthalmus Sturm, 1844

Aphaenopidius G.Müller, 1913

Aphaenops Bonvouloir, 1862

Aphaenopsis G.Müller, 1913

Apocimmerites Belousov, 1998

Apoduvalius Jeannel, 1953

Aputrechisibus Trezzi, 2007

Arctaphaenops Meixner, 1925

Aspidaphaenops Ueno, 2006

Austrotrechus B.Moore, 1972

Awatrechus Ueno, 1955

Balazucellus Deuve, 2001

Bathytrechus Ueno, 2005

Beronaphaenops B.V.Gueorguiev, 2012

Birmaphaenops Deuve, 2017

Blemus Dejean, 1821

Boldoriella Jeannel, 1928

Boreaphaenops Ueno, 2002

Bothynotrechus B.Moore, 1972

Casaleaphaenops Tian; Huang & Ma, 2021

Cathaiaphaenops Deuve, 1996

Caucasaphaenops Belousov, 1999

Caucasorites Belousov & Zamotajlov, 1997

Chaetoduvalius Jeannel, 1928

Chaetotrechiama Ueno, 1982

Chiapadytes Vigna Taglianti, 1977

Chu Tian & He, 2020

Cimbrodytes Piva, 2021

Cimmerites Jeannel, 1928

Cimmeritodes Deuve, 1996

Ckacus Tian & Huang, 2021

Columbitrechus Mateu, 1982

Coreoblemus Ueno, 1969

Croatotrechus Casale & Jalzic, 1999

Dabatrechus Ueno, 2004

Dactylotrechus Belousov & Kabak, 2003

Daiconotrechus Ueno, 1971

Dalmataphaenops Monguzzi, 1993

Darlingtonea Valentine, 1952

Derossiella Queinnec, 2008

Deuveaphaenops Tian & Huang, 2017

Dianotrechus Tian, 2016

Doderotrechus Vigna Taglianti, 1968

Dongoblemus Deuve & Tian, 2016

Dongodytes Deuve, 1993

Dracotrechus Ueno, 2010

Duvalioblemus Deuve, 1995

Duvaliomimus Jeannel, 1928

Duvaliopsis Jeannel, 1928

Duvalius Delarouzée, 1859

Eocnides Jeannel, 1954

Epaphiopsis Ueno, 1953

Epaphiotrechus Deuve & Kavanaugh, 2016

Epaphius Leach in Samouelle, 1819

Erebotrechus Britton, 1964

Escolatrechus Mateu, 2003

Eutrechopsis B.Moore, 1972

Eutrechus B.Moore, 1972

Geotrechus Jeannel, 1919

Giraffaphaenops Deuve, 2002

Glabroduvalius Vrbica; Curcic; Antic & Curcic, 2013

Goedetrechus B.Moore, 1972

Gotoblemus Ueno, 1970

Graciliblemus Deuve & Tian, 2016

Guatemalatrechus Trezzi, 2003

Guiaphaenops Deuve, 2002

Guizhaphaenops Vigna Taglianti, 1997

Guizhaphaenopsoides Tian; Huang & Li, 2021

Gulaphaenops Ueno, 1987

Himalaphaenops Ueno, 1980

Himalotrechus Belousov; Kabak & J.Schmidt, 2019

Homaloderodes Jeannel, 1962

Huoyanodytes Tian & Huang, 2016

Iberotrechus Jeannel, 1920

Iga Ueno, 1953

Incatrechus Mateu & Belles, 1982

Inotrechus Dolzhanski & Ljovushkin, 1989

Ishidatrechus Ueno, 1956

Ishikawatrechus Habu, 1950

Italaphaenops Ghidini, 1964

Iyotrechus Ueno & Naito, 2009

Jalzicaphaenops Lohaj & Lakota, 2010

Jeannelius Kurnakov, 1959

Jiangxiaphaenops Ueno & Clarke, 2007

Jiulongotrechus Tian; Huang & Wang, 2015

Junaphaenops Ueno, 1997

Junnanotrechus Ueno & Yin, 1993

Kettlotrechus Townsend, 2010

Kosswigia Jeannel, 1947

Kozlovites Jeannel, 1935

Kupetrechus Larochelle & Larivière, 2007

Kurasawatrechus Yoshida & Nomura, 1952

Kusumia Ueno, 1952

Lanxangaphaenops Deuve, 2012

Laoblemus Ueno, 2006

Laosaphaenops Deuve, 2000

Lessinodytes Vigna Taglianti, 1982

Libotrechus Ueno, 1998

Luoxiaotrechus Tian & Yin, 2013

Luyatrechus M.Etonti & Mateu, 2000

Luzonotrechus Ueno, 1979

Mamesdytes Trezzi, 2003

Masuzoa Ueno, 1960

Masuzonoblemus Ueno, 1989

Mayaphaenops Vigna Taglianti, 1977

Meganophthalmus Kurnakov, 1959

Mexaphaenops Bolivar y Pieltain, 1942

Mexitrechus Barr, 1982

Miaotrechus Tian; Chen & Ma, 2020

Microblemus Ueno, 2007

Mimanillus B.Moore, 1972

Mimotrechus B.Moore, 1972

Minimaphaenops Deuve, 2000

Minosaphaenops Queinnec, 2008

Minutotrechus Deuve & Kavanaugh, 2016

Nannotrechus Winkler, 1926

Nanotrechiama Belousov & Kabak, 2018

Neaphaenops Jeannel, 1920

Nelsonites Valentine, 1952

Neotrechus G.Müller, 1913

Nesaeoduvalius Casale & Giachino, 2016

Nesiotrechus Ueno, 1995

Nipponaphaenops Ueno, 1971

Nototrechus B.Moore, 1972

Nunbergites Pawlowski & Stachowiak, 1991

Omalodera Blanchard, 1842

Oodinotrechus Ueno, 1998

Oroblemites Ueno & Pawlowski, 1981

Oroblemus Ueno & Yoshida, 1966

Orotrechus G.Müller, 1913

Oxytrechus Jeannel, 1927

Panaphaenops Tian; Huang & Ma, 2021

Paracimmerites Belousov, 1998

Paraphaenops Jeannel, 1916

Paratrechiotes Ueno, 1995

Paratrechus Jeannel, 1920

Petraphaenops Delic; Kapla & Colla, 2019

Pheggomisetes Knirsch, 1923

Pilosaphaenops Deuve & Tian, 2008

Plesioaphaenops Deuve & Tian, 2011

Pogonoschema Jeannel, 1927

Pontodytes Casale & Giachino, 1989

Porocimmerites Belousov, 1998

Protrechiama Belousov & Kabak, 2003

Pseudanophthalmus Jeannel, 1920

Pseudaphaenops Winkler, 1912

Pseudocnides Jeannel, 1927

Pseudotrechisibus Mateu & Belles, 1982

Puertrechus Belousov & Kabak, 2014

Putzeysius Jeannel, 1962

Qianaphaenops Ueno, 2000

Qianotrechus Ueno, 2000

Queinnectrechus Deuve, 1992

Rakantrechus Ueno, 1951

Ryugadous Habu, 1950

Sardaphaenops Cerruti & Henrot, 1956

Satotrechus Ueno, 2006

Sbordoniella Vigna Taglianti, 1980

Scotoplanetes Absolon, 1913

Scototrechus Britton, 1962

Shenaphaenops Ueno, 1999

Shennongotrechus Ueno, 2004

Shenoblemus Tian & Fang, 2020

Shilinotrechus Ueno, 2003

Shiqianaphaenops Tian, 2016

Shuangheaphaenops Tian, 2017

Shuaphaenops Ueno, 1999

Sichuanotrechus Deuve, 2005

Sidublemus Tian & Yin, 2013

Sinaphaenops Ueno & Wang, 1991

Sinocimmerites Deuve, 2007

Sinotrechiama Ueno, 2000

Sinotroglodytes Deuve, 1996

Sloanella Jeannel, 1927

Speotrechus Jeannel, 1922

Stevensius Jeannel, 1923

Stygiotrechus Ueno, 1958

Subilsia Español, 1967

Sumatrechus Deuve, 2005

Superbotrechus Deuve & Tian, 2009

Suzuka Ueno, 1956

Taiwanotrechus Ueno, 1987

Taniatrechus Belousov & Dolzhanski, 1994

Tasmanorites Jeannel, 1927

Tasmanotrechus B.Moore, 1972

Taurocimmerites Belousov, 1998

Thalassoduvalius Ueno, 1956

Tianeotrechus Tian & Tang, 2016

Tianzhuaphaenops Zhao & Tian, 2016

Tienmutrechus Suenson, 1957

Tonkinaphaenops Deuve, 2013

Toshiaphaenops Ueno, 1999

Trechepaphiama Deuve & Kavanaugh, 2016

Trechepaphiopsis Deuve & Kavanaugh, 2016

Trechiama Jeannel, 1927

Trechiamiotes Deuve, 1998

Trechicomimus Mateu & Nègre, 1972

Trechiella Jeannel, 1927

Trechimorphus Jeannel, 1927

Trechinotus Jeannel, 1962

Trechiotes Jeannel, 1954

Trechisibus Motschulsky, 1862

Trechistus B.Moore, 1972

Trechoblemus Ganglbauer, 1891

Trechus Clairville, 1806

Trichaphaenops Jeannel, 1916

Troglocimmerites Ljovuschkin, 1970

Tropidotrechus Jeannel, 1927

Typhlotrechus G.Müller, 1913

Uenoaphaenops Tian & He, 2020

Uenoites Belousov & Kabak, 2016

Uenotrechus Deuve & Tian, 1999

Ushijimaella Ueno, 1980

Velebitaphaenops Casale & Jalzic, 2012

Velesaphaenops S.Curcic et al., 2018

Vietotrechus Ueno, 1995

Wanhuaphaenops Tian & Wang, 2016

Wanoblemus Tian & Fang, 2016

Wulongoblemus Ueno, 2007

Xenotrechus Barr & Krekeler, 1967

Xiangxius Tian; Huang & Li, 2021

Xuedytes Tian & Huang, 2017

Yalongaphaenops Belousov & Kabak, 2021

Yamautidius Ueno, 1957

Yanzaphaenops Ueno, 2010

Yunotrechus Tian & Huang, 2014

Zhijinaphaenops Ueno & Ran, 2002

Subtribe Trechodina Jeannel, 1926

Amblystogenium Enderlein, 1905

Canarobius Machado, 1987

Cnides Motschulsky, 1862

Cothresia Jeannel, 1964

Cyphotrechodes Jeannel, 1926

Eotrechodes Ueno; Lafer & Sundukov, 1995

Himalotrechodes Ueno, 1981

Iberotrechodes Faille et al., 2021

Pachydesus Motschulsky, 1864

Pachytrechodes Jeannel, 1960

Paratrechodes Jeannel, 1926

Scaurotrechodes Geginat, 2006

Spelaeovulcania Machado, 1987

Sporades Fauvel, 1882

Thalassophilus Wollaston, 1854

Trechobembix Jeannel, 1926

Trechodes Blackburn, 1901

Trechosia Jeannel, 1926

Trechosiella Jeannel, 1960

†Antarctotrechus Ashworth & Erwin, 2016

†Trechinites Heer, 1862

†Trechoides Motschulsky, 1856

Tribe Zolini Sharp, 1886

Subtribe Merizodontina Sloane, 1920

Idacarabus Lea, 1910

Maungazolus Larochelle & Larivière, 2017

Merizodus Solier, 1849

Percodermus Sloane, 1920

Pseudoopterus Csiki, 1928

Pterocyrtus Sloane, 1920

Sloaneana Csiki, 1933

Synteratus Broun, 1909

Thayerella Baehr, 2016

Subtribe Zolina Sharp, 1886

Oopterus Guérin-Méneville, 1841

Zolus Sharp, 1886

Subfamily Xenaroswellianinae Erwin, 2007

Xenaroswelliana Erwin, 2007

Subfamily †Conjunctiinae Ponomarenko, 1977

†Conjunctia Ponomarenko, 1977

Subfamily †Protorabinae Ponomarenko, 1977

†Aethocarabus Ren; Lu & Guo, 1995

†Atrirabus Hong & Wang, 1990

†Cordorabus Ponomarenko, 1977

†Cretorabus Ponomarenko, 1977

†Denudirabus Ren; Lu & Guo, 1995

†Lirabus Hong, 1992

†Lithorabus Ponomarenko, 1977

†Magnirabus Hong & Wang, 1990

†Mesorabus Ponomarenko, 1977

†Nebrorabus Ponomarenko, 1989

†Ovrabites Ponomarenko, 1977

†Penecupes Ren, 1995

†Protorabus Ponomarenko, 1977

Not Assigned to Subfamily

†Agatoides Motschulsky, 1856

†Amphoxyne Bode, 1953

†Carabites Heer, 1852

†Cavicarabus Hong, 1991

†Conexicoxa Lin, 1986

†Cymatopterus Lomnicki, 1894

†Fangshania Hong, 1981

†Glenopterus Heer, 1847

†Hebeicarabus Hong, 1983

†Megacarabus Hong, 1983

†Meileyingia Hong, 1987

†Miocarabus Hong, 1983

†Neothanes Scudder, 1890

†Procarabus Oppenheim, 1888

†Prosynactus Bode, 1953

†Shanwangicarabus Hong, 1985

†Sinis Heer, 1862

†Sinocalosoma Hong & Wang, 1986

†Sinocaralosoma Hong, 1984

†Sunocarabus Hong, 1987

†Tauredon Handlirsch, 1910

†Wuchangicarabus Hong, 1991

†Xishanocarabus Hong, 1984

†Yunnanocarabus Lin, 1977