Biology:Seldsienean

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Seldsienean is an extinct genus of teleosauroid thalattosuchian from the Middle Jurassic of England and France.[1] The only known species, S. megistorhynchus, is known from remains from the Calcaire de Caen and the Cornbrash Formation.[1] In life, Seldsienean was a marine reptile that would have fed mostly on fish and other small animals.[2]


Taxonomy

The type species, S. megistorhynchus, was originally named as a species Steneosaurus in 1866.[3] The genus Steneosaurus was used as a longtime wastebasket taxon for various teleosauroid specimens and had more than a dozen species. The type species, Steneosaurus rostromajor is undiagnostic, making the genus and species dubious and invalid.[1] Additionally, many species of so-called "Steneosaurus" were found to be quite different and unrelated to one another, thus needing new generic names.[1]

In 2020, Michela Johnson and colleagues reassigned "Steneosaurus" megistorhynchus to the new genus Seldsienean, resulting in the new combination Seldsienean megistorhynchus.[1] The generic name is derived from Old English and translates to "seldom seen one" in reference to its fossils being rarer compared to other Bathonian teleosauroids.[1]

Classification

Seldsienean was initially recovered as a basal member of Machimosauridae, a family of large-bodied, marine-adapted reptiles. Members of this group are characterized by robust skulls, powerful jaws, and heavily built bodies, adaptations that positioned them among the apex predators within their respective marine ecosystems. The cladogram below is from the 2020 analysis by Johnson and colleagues.[1]

Machimosauridae
 

Macrospondylus bollensis

 
 
 

Clovesuurdameredeor stephani

 
 
 

Charitomenosuchus leedsi

 
 
 

Seldsienean megistorhynchus

 
 
 

Deslongchampsina larteti

 
Machimosaurinae
 
 

Proexochokefalos hebertii

 
 

cf. Proexochokefalos bouchardi

 
 
 
 

Neosteneosaurus edwardsi

 
 

Andrianavoay baroni

 
Machimosaurini
 

Lemmysuchus obtusidens

 
 

Yvridiosuchus boutilieri

 
 
 

Machimosaurus buffetauti

 
 
 

Machimosaurus mosae

 
 
 

Machimosaurus rex

 
 

Machimosaurus hugii

 
 
 
 
 
 
 
 
 
 
 
 

A later study in 2024 by Mark Young and colleagues found Seldsienean to be part of a different teleosauroid group called Teleosauridae. It was found to be most closely related to the subgroup Aeolodontini.[4]

Teleosauroidea
 

Machimosauridae

 
Teleosauridae
 
 

Mystriosaurus laurillardi

 
 
 

Indosinosuchus potamosiamensis

 
 

Chinese teleosauroid (IVPP V 10098)

 
 
 
 
Teleosaurinae
 

Platysuchus multiscrobiculatus

 
 

Teleosaurus cadomensis

 
 
 
 

Seldsienean megistorhynchus

 
Aeolodontini
 

Mycterosuchus nasutus

 
 
 

Aeolodon priscus

 
 
 

Sericodon jugleri

 
 

Bathysuchus megarhinus

 
 
 
 
 
 
 
 

Young et al., 2024 also submitted a proposal to the International Commission on Zoological Nomenclature to designate Seldsienean megistorhynchus as the new type species for Steneosaurus and to adopt this revised type species as the basis for defining the family Steneosauridae.[4] However, as of 2025 this issue is not fully resolved and Seldsienean remains valid.

Ecology

Based on its jaw morphology, Seldsienean is thought to have been a longirostrine generalist. It possessed long thin jaws that had a fast but relatively weak bite. These adaptations point to a lifestyle of catching and eating fish and other smaller marine animals.[2]

References

  1. ↑ 1.0 1.1 1.2 1.3 1.4 1.5 1.6 Johnson, Michela M.; Young, Mark T.; Brusatte, Stephen L. (2020). "The phylogenetics of Teleosauroidea (Crocodylomorpha, Thalattosuchia) and implications for their ecology and evolution" (in en). PeerJ 8. doi:10.7717/peerj.9808. ISSN 2167-8359. PMID 33083104. 
  2. ↑ 2.0 2.1 Johnson, M. M.; Foffa, D.; Young, M. T.; Brusatte, S. L. (2022). "The ecological diversification and evolution of Teleosauroidea (Crocodylomorpha, Thalattosuchia), with insights into their mandibular biomechanics". Ecology and Evolution 12 (11). doi:10.1002/ece3.9484. PMID 36415878. Bibcode: 2022EcoEv..12E9484J. 
  3. ↑ Eudes-Deslongchamps, J. A. "Sur la découverte d'une mâchoire inférieure entière de Steneosaurus megistorhynchus (GEOFF.) trouvée à Allemagne et comparaison de cette espèce avec le Teleosaurus larteti (DESL.)." Bulletin de la Société linnéenne de Normandie 10 (1866): 80-85.
  4. ↑ 4.0 4.1 Young, Mark T.; Wilberg, Eric W.; Johnson, Michela M.; Herrera, Yanina; De Andrade, Marco Brandalise; Brignon, Arnaud; Sachs, Sven; Abel, Pascal et al. (2024). "The history, systematics, and nomenclature of Thalattosuchia (Archosauria: Crocodylomorpha)". Zoological Journal of the Linnean Society 200 (2): 547–617. doi:10.1093/zoolinnean/zlad165. 

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