Biology:Trichomycterus

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Trichomycterus is a genus of fish in the family Trichomycteridae, the largest genus of its family with over 170 species currently described. This genus is native to freshwater habitats in Central and South America.[1] These fish are generally small, usually about 5 to 15 cm (2–6 in) in standard length, although the largest, T. rivulatus, can reach more than twice this size.[1] Species differ from one another primarily in body proportions, fin ray counts and colouration.[2] Despite their relatively small size, some, such as T. punctulatus, support fisheries and are important in the local cuisine.[3]

Taxonomy

This genus is defined by the lack of specializations found in other trichomycterids and is certainly polyphyletic.[4][5] Although known to contain many species, Trichomycterus is poorly known with many of the known species based on brief descriptions.[4] Many species have been described recently and many more are waiting to be described.[6]

Some authors have tentatively defined putative monophyletic assemblages within the genus Trichomycterus based on the possession of unique morphological features.[4] The Trichomycterus brasiliensis species-complex includes T. brasiliensis, T. iheringi, T. maracaya, T. mimonha, T. pirabitira, T. potschi, T. vermiculatus and several undescribed species apparently endemic to the main river basins draining the Brazilian Shield.[7] However, an assemblage of species from south and southeastern Brazil is also supported that includes T. araxa,[8] T. castroi, T. davisi, T. guaraquessaba, T. immaculatus, T. itatiayae, T. mboycy, T. mirissumba, T. naipi, T. nigricans, T. papilliferus, T. plumbeus, T. stawiarski, T. taroba, T. triguttatus and T. zonatus,[5] T. aguarague, T. alterus, T. belensis, T. boylei and T. ramosus also form a diagnosable species assemblage.[2]

Distribution and habitat

Species of Trichomycterus inhabit a diversity of habitats throughout South and Central America from Costa Rica in the north to Patagonia in the south and from lowland Atlantic rainforest in the east to Andean highland streams in the west.[2] They are, together with Astroblepus and Orestias, among the very few native fish genera at high altitude in the Andes[9] and in some places, for example certain high-elevation localities in western Argentina, Trichomycterus are the only fishes.[10] About 60 nominal species are endemic to the river basins draining the Andes and hills of the Guianan Shield and about 30 species are endemic to river basins draining the Brazilian Shield.[11]

Despite the broad distribution of the genus, most species have limited distributions and usually are restricted to only one river.[12] Wide-ranging species are most likely complexes of species that are difficult to differentiate, such as the T. brasiliensis species-complex.[7] Trichomycterus gorgona, from a small stream on Gorgona Island located west of the Pacific coast of Colombia, is the first known trichomycterid to be endemic to an offshore island.[13] Many species are troglobitic.[14]

Species

There are currently 213 recognized species in this genus:

  • Trichomycterus adautoleitei Costa, Azevedo-Santos & Katz, 2023
  • Trichomycterus aguarague L. A. Fernández & Osinaga, 2006
  • Trichomycterus albinotatus W. J. E. M. Costa, 1992
  • Trichomycterus alternatus (C. H. Eigenmann, 1917)
  • Trichomycterus alterus (Marini, Nichols & La Monte, 1933)
  • Trichomycterus altipombensis Costa, Katz, Vilardo & Mattos, 2022
  • Trichomycterus anaisae Katz & Costa, 2021
  • Trichomycterus antiquus Costa, Feltrin, Mattos & Katz, 2024
  • Trichomycterus araxa W. J. E. M. Costa, Mattos, Sampaio, Giongo, Almeida, & Katz, 2022 [8]
  • Trichomycterus ardilai Ardila Rodríguez, 2023
  • Trichomycterus areolatus Valenciennes, 1846
  • Trichomycterus argos Lezama, Triques & P. S. Santos, 2012[15]
  • Trichomycterus arhuaco Ardila Rodríguez, 2016[16]
  • Trichomycterus arleoi (Fernández-Yépez, 1972)
  • Trichomycterus astromycterus Reis, de Pinna & Pessali, 2019
  • Trichomycterus atochae (W. R. Allen, 1942)
  • Trichomycterus auroguttatus W. J. E. M. Costa, 1992
  • Trichomycterus bahianus W. J. E. M. Costa, 1992
  • Trichomycterus ballesterosi Ardila Rodríguez, 2011[17]
  • Trichomycterus banneaui (C. H. Eigenmann, 1912)
  • Trichomycterus barbouri (C. H. Eigenmann, 1911)
  • Trichomycterus barrocus Reis & de Pinna, 2022
  • Trichomycterus belensis L. A. Fernández & Vari, 2002
  • Trichomycterus berthalutzae Costa, Barbosa & Katz 2024
  • Trichomycterus bogotensis (C. H. Eigenmann, 1912)
  • Trichomycterus bomboizanus (Tortonese, 1942)
  • Trichomycterus borellii Boulenger, 1897
  • Trichomycterus boylei (Nichols, 1956)
  • Trichomycterus brasiliensis Lütken, 1874[18]
  • Trichomycterus brigadeirensis Costa, Katz & Vilardo, 2023
  • Trichomycterus brucutu Reis & de Pinna, 2022
  • Trichomycterus brunoi M. A. Barbosa & W. J. E. M. Costa, 2010[18]
  • Trichomycterus cachiraensis Ardila Rodríguez, 2008
  • Trichomycterus caipora S. M. Q. Lima, Lazzarotto & W. J. E. M. Costa, 2008
  • Trichomycterus calai Ardila Rodríguez, 2019
  • Trichomycterus caliensis (C. H. Eigenmann, 1912)
  • Trichomycterus candidus (P. Miranda-Ribeiro, 1949)
  • Trichomycterus caparaoensis Costa, Barbosa & Katz, 2023
  • Trichomycterus caratinguensis Costa, Katz & Vilardo, 2023
  • Trichomycterus caribensis Ardila Rodríguez, 2024
  • Trichomycterus casitaensis Ardila Rodríguez, 2017[19]
  • Trichomycterus castelensis Costa, Katz & Vilardo, 2023
  • Trichomycterus catamarcensis L. A. Fernández & Vari, 2000
  • Trichomycterus caudofasciatus Alencar & W. J. E. M. Costa, 2004
  • Trichomycterus celsae Lasso A. & Provenzano, 2003
  • Trichomycterus cepitaensis Ardila Rodríguez, 2025
  • Trichomycterus cerritoensis Ardila Rodríguez, 2018
  • Trichomycterus chaberti Durand, 1968
  • Trichomycterus chapmani (C. H. Eigenmann, 1912)
  • Trichomycterus chiltoni (C. H. Eigenmann, 1920)
  • Trichomycterus choroloensis Ardila Rodríguez, 2024
  • Trichomycterus chungaraensis Arratia, 1983
  • Trichomycterus claudiae M. A. Barbosa & W. J. E. M. Costa, 2010[18]
  • Trichomycterus coelhorum Costa, Azevedo-Santos & Katz, 2023
  • Trichomycterus conradi (C. H. Eigenmann, 1912)
  • Trichomycterus corduvensis Weyenbergh (de), 1877
  • Trichomycterus curitiensis Ardila Rodríguez, 2024
  • Trichomycterus dali Rizzato, E. P. D. Costa, Trajano & Bichuette, 2011[14]
  • Trichomycterus diamantinensis Costa, Feltrin, Mattos & Katz, 2024
  • Trichomycterus dispar (Tschudi, 1846)
  • Trichomycterus donascimientoi Castellanos-Morales, 2018
  • Trichomycterus dorsostriatus (C. H. Eigenmann, 1917)
  • Trichomycterus duellmani Arratia & Menu-Marque, 1984
  • Trichomycterus emanueli (L. P. Schultz, 1944)
  • Trichomycterus espinhacensis Costa & Katz, 2023
  • Trichomycterus fabioheppi Costa, Barbosa & Katz, 2024
  • Trichomycterus fassli (Steindachner, 1915)
  • Trichomycterus ferreri Ardila Rodríguez, 2018
  • Trichomycterus florensis (P. Miranda-Ribeiro, 1943)
  • Trichomycterus fuliginosus M. A. Barbosa & W. J. E. M. Costa, 2010[18]
  • Trichomycterus funebris Katz & Costa, 2021
  • Trichomycterus gabrieli (G. S. Myers, 1926)
  • Trichomycterus gairaensis Ardila Rodríguez, 2018
  • Trichomycterus galvisi Ardila Rodríguez, 2024
  • Trichomycterus gambitaensis Ardila Rodríguez, 2023
  • Trichomycterus garbei Costa, Azevedo-Santos & Katz, 2023
  • Trichomycterus garciamarquezi Ardila Rodríguez, 2016[16]
  • Trichomycterus gasparinii M. A. Barbosa, 2013[20]
  • Trichomycterus giarettai M. A. Barbosa & Katz, 2016[21]
  • Trichomycterus giganteus S. M. Q. Lima & W. J. E. M. Costa, 2004
  • Trichomycterus goeldii Boulenger, 1896
  • Trichomycterus gorgona L. A. Fernández & Schaefer, 2005
  • Trichomycterus guianensis (C. H. Eigenmann, 1909)
  • Trichomycterus hatoensis Ardila Rodríguez, 2025
  • Trichomycterus heterodontus (C. H. Eigenmann, 1917)
  • Trichomycterus hualco L. A. Fernández & Vari, 2009
  • Trichomycterus humboldti Costa & Katz, 2021
  • Trichomycterus illuvies Reis & de Pinna, 2022
  • Trichomycterus immaculatus (C. H. Eigenmann & R. S. Eigenmann, 1889)
  • Trichomycterus ingaiensis Katz & Costa, 2021
  • Trichomycterus ipatinga Reis & de Pinna, 2022
  • Trichomycterus itacambirussu Triques & Vono, 2004
  • Trichomycterus itacarambiensis Trajano & de Pinna, 1996
  • Trichomycterus itatiayae A. Miranda-Ribeiro, 1906
  • Trichomycterus jacupiranga Wosiacki & Oyakawa, 2005
  • Trichomycterus jequitinhonhae Triques & Vono, 2004
  • Trichomycterus kankuamo Ardila Rodríguez, 2016[16]
  • Trichomycterus knerii Steindachner, 1882
  • Trichomycterus landinga Triques & Vono, 2004
  • Trichomycterus largoperculatus Costa & Katz, 2022
  • Trichomycterus latidens (C. H. Eigenmann, 1917)
  • Trichomycterus latistriatus (C. H. Eigenmann, 1917)
  • Trichomycterus laucaensis Arratia, 1983
  • Trichomycterus lauryi Donin, Ferrer & Carvalho, 2020[22]
  • Trichomycterus lauzannii Miranda & Fernández, 2020
  • Trichomycterus lewi Lasso A. & Provenzano, 2003
  • Trichomycterus listruroides Costa, Katz & Azevedo-Santos, 2023
  • Trichomycterus longibarbatus W. J. E. M. Costa, 1992
  • Trichomycterus luetkeni Katz & Costa, 2021
  • Trichomycterus macareguaensis
  • Trichomycterus macrophthalmus M. A. Barbosa & W. J. E. M. Costa, 2012[23]
  • Trichomycterus macrotrichopterus M. A. Barbosa & W. J. E. M. Costa, 2010[18]
  • Trichomycterus maculosus M. A. Barbosa & W. J. E. M. Costa, 2010[24]
  • Trichomycterus maldonadoi Ardila Rodríguez, 2011[25]
  • Trichomycterus manaurensis Ardila Rodríguez, 2016[16]
  • Trichomycterus maracaiboensis (L. P. Schultz, 1944)
  • Trichomycterus maracaya Bockmann & Sázima (fr), 2004
  • Trichomycterus mariamole M. A. Barbosa & W. J. E. M. Costa, 2010[18]
  • Trichomycterus megantoni L. A. Fernández & Chuquihuamaní, 2007
  • Trichomycterus melanopygius Reis, Santos, Britto, Volpi & de Pinna, 2020
  • Trichomycterus meridae Regan, 1903
  • Trichomycterus migrans (Dahl, 1960)
  • Trichomycterus mimonha W. J. E. M. Costa, 1992
  • Trichomycterus mimosensis M. A. Barbosa, 2013[20]
  • Trichomycterus minus L. A. Fernández & Vari, 2012[26]
  • Trichomycterus mirissumba W. J. E. M. Costa, 1992
  • Trichomycterus mogotensis Ardila Rodríguez, 2017
  • Trichomycterus mondolfi (L. P. Schultz, 1945)
  • Trichomycterus montesi Ardila Rodríguez, 2016[16]
  • Trichomycterus motatanensis (L. P. Schultz, 1944)
  • Trichomycterus mutabilicolor Costa, 2022
  • Trichomycterus nabusimakensis Ardila Rodríguez, 2018
  • Trichomycterus nietoi Ardila Rodríguez, 2014[27]
  • Trichomycterus nigricans Valenciennes, 1832
  • Trichomycterus nigroauratus M. A. Barbosa & W. J. E. M. Costa, 2008
  • Trichomycterus nigromaculatus Boulenger, 1887
  • Trichomycterus novalimensis M. A. Barbosa & W. J. E. M. Costa, 2010[18]
  • Trichomycterus ocanaensis Ardila Rodríguez, 2011[28]
  • Trichomycterus oroyae (C. H. Eigenmann & R. S. Eigenmann, 1889)
  • Trichomycterus pantherinus Alencar & W. J. E. M. Costa, 2004
  • Trichomycterus paquequerensis (P. Miranda-Ribeiro, 1943)
  • Trichomycterus pauciradiatus Alencar & W. J. E. M. Costa, 2006
  • Trichomycterus pirabitira M. A. Barbosa & Azevedo-Santos, 2012[29]
  • Trichomycterus piratymbara Katz, M. A. Barbosa & W. J. E. M. Costa, 2013[30]
  • Trichomycterus piurae (C. H. Eigenmann, 1922)
  • Trichomycterus potschi M. A. Barbosa & W. J. E. M. Costa, 2003
  • Trichomycterus pradensis Sarmento-Soares, Martins-Pinheiro, Aranda & Chamon, 2005
  • Trichomycterus pseudosilvinichthys L. A. Fernández & Vari, 2004
  • Trichomycterus puna Fernández, Contrera & Bize, 2023
  • Trichomycterus punctatissimus Castelnau, 1855
  • Trichomycterus punctulatus Valenciennes, 1846
  • Trichomycterus puriventris M. A. Barbosa & W. J. E. M. Costa, 2012[31]
  • Trichomycterus quechuorus (Steindachner, 1900)
  • Trichomycterus quintus Costa, 2020
  • Trichomycterus ramosus L. A. Fernández, 2000
  • Trichomycterus regani (C. H. Eigenmann, 1917)
  • Trichomycterus reinhardti (C. H. Eigenmann, 1917)
  • Trichomycterus retropinnis Regan, 1903
  • Trichomycterus riojanus (C. Berg, 1897)
  • Trichomycterus rivulatus Valenciennes, 1846
  • Trichomycterus roigi Arratia & Menu-Marque, 1984
  • Trichomycterus romeroi (Fowler, 1941)
  • Trichomycterus rosablanca Mesa S., Lasso, Ochoa & DoNascimiento, 2018
  • Trichomycterus rubbioli Bichuette & Rizzato, 2012[32]
  • Trichomycterus rubiginosus M. A. Barbosa & W. J. E. M. Costa, 2010[18]
  • Trichomycterus ruficaudatus Costa, Mattos, Amorim, Gago & Katz, 2025
  • Trichomycterus ruitoquensis Ardila Rodríguez, 2007
  • Trichomycterus sainthilairei Katz & Costa, 2021
  • Trichomycterus sanchezi Ardila Rodríguez, 2024
  • Trichomycterus sandovali Ardila Rodríguez, 2006
  • Trichomycterus santaeritae (C. H. Eigenmann, 1918)
  • Trichomycterus santanderensis Castellanos-Morales, 2007
  • Trichomycterus santuarioensis Ardila Rodríguez, 2024
  • Trichomycterus saquarema Costa, Katz, Vilardo & Amorim, 2022
  • Trichomycterus saturatus Costa, Katz & Azevedo-Santos, 2023
  • Trichomycterus septemradiatus Katz, M. A. Barbosa & W. J. E. M. Costa, 2013[30]
  • Trichomycterus simacotaensis Ardila Rodríguez, 2024
  • Trichomycterus sketi Castellanos-Morales, 2011[33]
  • Trichomycterus spectrum DoNascimiento & Prada-Pedreros, 2020
  • Trichomycterus spegazzinii (C. Berg, 1897)
  • Trichomycterus spelaeus DoNascimiento, Villarreal & Provenzano, 2001
  • Trichomycterus spilosoma (Regan, 1913)
  • Trichomycterus steindachneri DoNascimiento, Prada-Pedreros & Guerrero-Kommritz, 2014[34]
  • Trichomycterus stellatus (C. H. Eigenmann, 1918)
  • Trichomycterus straminius (C. H. Eigenmann, 1917)
  • Trichomycterus striatus (Meek & Hildebrand, 1913)
  • Trichomycterus sucrensis Ardila Rodríguez, 2018
  • Trichomycterus taczanowskii Steindachner, 1882
  • Trichomycterus taenia Kner, 1863
  • Trichomycterus taeniops Fowler, 1954
  • Trichomycterus tantalus Reis, Vieira & de Pinna, 2022
  • Trichomycterus taroba Wosiacki & Garavello, 2004
  • Trichomycterus tenuis Weyenbergh (de), 1877
  • Trichomycterus tete M. A. Barbosa & W. J. E. M. Costa, 2011[35]
  • Trichomycterus tetuanensis L. J. García-Melo, Villa-Navarro & DoNascimiento, 2016[36]
  • Trichomycterus therma L. A. Fernández & Miranda-Chumacero, 2007
  • Trichomycterus tiraquae (Fowler, 1940)
  • Trichomycterus torcoromaensis Ardila Rodríguez, 2016[37]
  • Trichomycterus transandianus (Steindachner, 1915)
  • Trichomycterus travassosi (Miranda Ribeiro, 1949)
  • Trichomycterus trefauti Wosiacki, 2004
  • Trichomycterus uberabensis Costa, Azevedo-Santos & Katz, 2023
  • Trichomycterus uisae Castellanos-Morales, 2008
  • Trichomycterus unicolor (Regan, 1913)
  • Trichomycterus valleduparensis Ardila Rodríguez, 2018
  • Trichomycterus varii Fernández & Andreoli Bize, 2018
  • Trichomycterus vermiculatus (C. H. Eigenmann, 1917)
  • Trichomycterus vinnulus Reis & de Pinna, 2022
  • Trichomycterus vitalbrazili Paulo J. Vilardo, Katz & W. J. E. M. Costa[38]
  • Trichomycterus vittatus Regan, 1903
  • Trichomycterus weyrauchi (Fowler, 1945)
  • Trichomycterus wiwa Ardila Rodríguez, 2018
  • Trichomycterus yuska Fernández & Schaefer, 2003
  • Trichomycterus zapatocaensis Ardila Rodríguez, 2023

References

  1. ↑ 1.0 1.1 Froese, Rainer and Pauly, Daniel, eds. (2017). Species of Trichomycterus in FishBase. June 2017 version.
  2. ↑ 2.0 2.1 2.2 Fernández, L.; Osinaga, K. (2006). "A New Trichomycterus (Siluriformes: Trichomycteridae) from Aguarague National Park of the Bolivian Preandean Region, with Comments on Relationships within of the Genus". Environmental Biology of Fishes 75 (4): 385–393. doi:10.1007/s10641-005-5065-4. Bibcode: 2006EnvBF..75..385F. https://link.springer.com/article/10.1007/s10641-005-5065-4. 
  3. ↑ "Monsefú Life". FondazionesLowFood. 2017. https://www.fondazioneslowfood.com/en/ark-of-taste-slow-food/monsefu-life/. 
  4. ↑ 4.0 4.1 4.2 Alencar, A.R.; Costa, W.J.E.M. (2006). "Trichomycterus pauciradiatus, a new catfish species from the upper rio Paraná basin, southeastern Brazil (Siluriformes: Trichomycteridae)". Zootaxa 1269: 43–49. doi:10.11646/zootaxa.1269.1.3. http://www.mapress.com/zootaxa/2006f/zt01269p049.pdf. 
  5. ↑ 5.0 5.1 Wosiacki, W.B. (2005). "A new species of Trichomycterus (Siluriformes: Trichomycteridae) from south Brazil and redescription of T. iheringi (Eigenmann)". Zootaxa 1040: 49–64. doi:10.11646/zootaxa.1040.1.5. http://www.mapress.com/zootaxa/2005f/zt01040p064.pdf. 
  6. ↑ Lima, S.M.Q.; Lazzarotto, H.; Costa, W.J.E.M. (2008). "A new species of Trichomycterus (Siluriformes: Trichomycteridae) from lagoa Feia drainage, southeastern Brazil". Neotropical Ichthyology 6 (3): 315–322. doi:10.1590/S1679-62252008000300004. http://www.scielo.br/pdf/ni/v6n3/a04v6n3.pdf. 
  7. ↑ 7.0 7.1 Bockmann, F.A.; Sazima, I. (2004). "Trichomycterus maracaya, a new catfish from the upper rio Paraná, southeastern Brazil (Siluriformes: Trichomycteridae), with notes on the T. brasiliensis species-complex". Neotropical Ichthyology 2 (2): 61–74. doi:10.1590/S1679-62252004000200003. http://www.scielo.br/pdf/ni/v2n2/v2n2a03.pdf. 
  8. ↑ 8.0 8.1 Costa, W.J.E.M.; Mattos, J.L.O.; Sampaio, W.M.S.; Giongo, P.; de Almeida, F.B.; Katz, A.M. (2022). "Phylogenetic relationships of a new catfish of the genus Trichomycterus (Siluriformes, Trichomycteridae) from the Brazilian Cerrado, and the role of Cenozoic events in the diversification of mountain catfishes". Zoosystematics and Evolution 98 (1): 151–164. doi:10.3897/zse.98.83109. https://zse.pensoft.net/article/83109/. 
  9. ↑ Nelson, Joseph S.; Grande, Terry C.; Wilson, Mark V. H. (2016). Fishes of the World (5th ed.). Hoboken: John Wiley and Sons. p. 752. doi:10.1002/9781119174844. ISBN 978-1-118-34233-6. https://books.google.co.uk/books?id=E-MLDAAAQBAJ&pg=PP1. 
  10. ↑ Fernández, L.; Vari, R.P. (2004). "New Species of Trichomycterus from Midelevation Localities of Northwestern Argentina (Siluriformes: Trichomycteridae)". Copeia 2004 (4): 876–882. doi:10.1643/CI-04-094R1. http://www.bioone.org/doi/abs/10.1643/CI-04-094R1. 
  11. ↑ Alencar, A.R.; Costa, W.J.E.M. (2004). "Description of two new species of the catfish genus Trichomycterus from southeastern Brazil (Siluriformes: Trichomycteridae)". Zootaxa 744: 1–8. doi:10.11646/zootaxa.744.1.1. http://www.mapress.com/zootaxa/2004f/zt00744.pdf. 
  12. ↑ Lima, S.M.Q.; Costa, W.J.E.M. (2004). "Trichomycterus giganteus (Siluriformes: Loricarioidea: Trichomycteridae): a new catfish from the Rio Guandu basin, southeastern Brazil". Zootaxa 761: 1–6. doi:10.11646/zootaxa.761.1.1. http://www.mapress.com/zootaxa/2004f/zt00761.pdf. 
  13. ↑ Fernández, L.; Schaefer, S.A. (2005). "New Trichomycterus (Siluriformes: Trichomycteridae) from an Offshore Island of Colombia". Copeia 2005 (1): 68–76. doi:10.1643/CI-04-177R1. http://www.bioone.org/doi/abs/10.1643/CI-04-177R1. 
  14. ↑ 14.0 14.1 Rizzato, P.P.; Costa, E.P.D.Jr.; Trajano, E.; Bichuette, M.E. (2011). "Trichomycterus dali: a new highly troglomorphic catfish (Silurifomes: Trichomycteridae) from Serra da Bodoquena, Mato Grosso do Sul State, Central Brazil". Neotropical Ichthyology 9 (3): 477–491. doi:10.1590/S1679-62252011000300003. http://www.scielo.br/pdf/ni/v9n3/03.pdf. 
  15. ↑ Lezama, A.Q.; Triques, M.L.; Santos, P.S. (2012). "Trichomycterus argos (Teleostei: Siluriformes: Trichomycteridae), a new species from the Doce River Basin, Eastern Brazil". Zootaxa 3352: 60–68. doi:10.11646/zootaxa.3352.1.6. http://www.mapress.com/zootaxa/2012/f/z03352p068f.pdf. 
  16. ↑ 16.0 16.1 16.2 16.3 16.4 Ardila Rodríguez, C.A. (2016). "Cinco nuevas especies de peces Trichomycterus para la región Caribe - Colombia". Barranquilla, Departamento del Atlántico 2: 1–26. https://drive.google.com/file/d/0BznGz8v1UX8YdC1walo0RnFfRTA/view. 
  17. ↑ Ardila Rodríguez, C.A. (2011). "Trichomycterus ballesterosi (Siluriformes: Trichomyceridae), especie nueva de la cuenca alta del Río Sinú, Colombia". Dahlia 11: 3–12. http://www.acictios.org/dahlia/Revista-Dahlia-11.pdf. 
  18. ↑ 18.0 18.1 18.2 18.3 18.4 18.5 18.6 18.7 Barbosa, M.A.; Costa, W.J.E.M. (2010). "Seven new species of the catfish genus Trichomycterus (Teleostei: Siluriformes: Trichomycteridae) from Southeastern Brazil and redescription of T. brasiliensis". Ichthyological Exploration of Freshwaters 21 (2): 97–122. http://www.pfeil-verlag.de/wp-content/uploads/2015/05/ief21_2_01.pdf. 
  19. ↑ Ardila Rodríguez, C.A. (2017). "Trichomycterus casitaensis, sp. nov. (Siluriformes: Trichomycteridae) una nueva especie de pez del río Tarra, cuenca alta del río Catatumbo departamento de Norte de Santander – Colombia". Barranquilla, Departamento del Atlántico 3: 1–23. 
  20. ↑ 20.0 20.1 Barbosa, M.A. (2013). "Description of two new species of the catfish genus Trichomycterus (Teleostei: Siluriformes: Trichomycteridae) from the coastal river basins, southeastern Brazil". Vertebrate Zoology 63 (3): 269–275. doi:10.3897/vz.63.e31442. http://www.senckenberg.de/files/content/forschung/publikationen/vertebratezoology/vz63-3/04_vertebrate_zoology_63-3_barbosa_269-275.pdf. 
  21. ↑ Barbosa, M.A.; Katz, A.M. (2016). "A new species of the catfish genus Trichomycterus (Teleostei: Siluriformes: Trichomycteridae) from the Paranaíba basin, Central Brazil". Vertebrate Zoology 66 (3): 261–265. doi:10.3897/vz.66.e31564. http://www.senckenberg.de/files/content/forschung/publikationen/vertebratezoology/vz66-3/03_vertebrate_zoology_66-3_barbosa_261-265.pdf. 
  22. ↑ Donin, Laura M.; Ferrer, Juliano; Carvalho, Tiago P. (2020). "Taxonomical study of Trichomycterus (Siluriformes: Trichomycteridae) from the Ribeira de Iguape River basin reveals a new species recorded in the early 20th century" (in en). Journal of Fish Biology 96 (4): 886–904. doi:10.1111/jfb.14278. ISSN 1095-8649. PMID 32039475. Bibcode: 2020JFBio..96..886D. https://onlinelibrary.wiley.com/doi/abs/10.1111/jfb.14278. 
  23. ↑ Barbosa, M.A.; Costa, W.J.E.M. (2012). "Trichomycterus macrophthalmus (Teleostei: Siluriformes: Trichomycteridae), a new species of catfish from the Paraíba do Sul river basin, southeastern Brazil". Vertebrate Zoology 62 (1): 79–82. doi:10.3897/vz.62.e31368. http://www.senckenberg.de/files/content/forschung/publikationen/vertebratezoology/vz62-1/03_vertebrate_zoology_62-1_barbosa_79-82.pdf. 
  24. ↑ Barbosa, M.A.; Costa, W.J.E.M. (2010). "Description of a new species of the catfish genus Trichomycterus (Teleostei: Siluriformes: Trichomycteridae) from the rio Paraíba do Sul basin, southeastern Brazil". Vertebrate Zoology 60 (3): 193–197. doi:10.3897/vz.60.e31010. http://www.senckenberg.de/files/content/forschung/publikationen/vertebratezoology/vz60-3/02_vertebrate_zoology_60-3_barbosa.pdf. 
  25. ↑ Ardila Rodríguez, C.A. (2011). "Trichomycterus maldonadoi (Siluriformes: Trichomyceridae), especie nueva de la cuenca alta del Río Sinú, Colombia". Dahlia 11: 13–22. http://www.acictios.org/dahlia/Revista-Dahlia-11.pdf. 
  26. ↑ Fernández, L.; Vari, R.P. (2012). "New Species of Trichomycterus (Teleostei: Siluriformes) from the Andean Cordillera of Argentina and the Second Record of the Genus in Thermal Waters". Copeia 2012 (4): 631–636. doi:10.1643/CI-12-035. https://repository.si.edu/bitstream/handle/10088/19591/vz_Fernandez_and_Vari_2012.pdf. 
  27. ↑ Ardila Rodríguez, C.A. (2014). "Trichomycterus nietoi sp. nov. (Siluriformes: Trichomycteridae) una nueva especie de pez del río Guachaca Sierra Nevada de Santa Marta. Departamento del Magdalena – Colombia". Barranquilla, Departamento del Atlántico 1: 1–23. https://carlosardilarodriguez.files.wordpress.com/2013/06/trichomycterus-nietoi.pdf. 
  28. ↑ Ardila Rodríguez, C.A. (2011). "Trichomycterus ocanaensis sp. nov. (Siluriformes: Trichomycteridae) una nueva especie de pez del Río Tejo, cuenca alta del Río Catatumbo. Departamento de Norte de Santander – Colombia". Barranquilla, Departamento del Atlántico 1: 1–16. https://carlosardilarodriguez.files.wordpress.com/2013/07/revista-trichomycterus-ocanaensis.pdf. 
  29. ↑ Barbosa, M.A.; Azevedo-Santos, V.M. (2012). "A new species of the catfish genus Trichomycterus (Teleostei: Siluriformes: Trichomycteridae) from the rio Paraná basin, southeastern Brazil". Vertebrate Zoology 62 (3): 357–362. doi:10.3897/vz.62.e31395. http://www.senckenberg.de/files/content/forschung/publikationen/vertebratezoology/vz62-3/02_vertebrate_zoology_62-3_barbosa_357-362.pdf. 
  30. ↑ 30.0 30.1 Katz, A.M.; Barbosa, M.A.; Costa, W.J.E.M. (2013). "Two new species of the catfish genus Trichomycterus from the Paraná river basin, southeastern Brazil (Teleostei: Trichomycteridae)". Ichthyological Exploration of Freshwaters 23 (4): 359–366. http://pfeil-verlag.de/wp-content/uploads/2015/05/ief23_4_08.pdf. 
  31. ↑ Barbosa, M.A.; Costa, W.J.E.M. (2012). "Trichomycterus puriventris (Teleostei: Siluriformes: Trichomycteridae), a new species of catfish from the rio Paraíba do Sul basin, southeastern Brazil". Vertebrate Zoology 62 (2): 155–160. doi:10.3897/vz.62.e31379. http://www.senckenberg.de/files/content/forschung/publikationen/vertebratezoology/vz62-2/01_vertebrate_zoology_62-2_barbosa__costa_155-160.pdf. 
  32. ↑ Bichuette, M.E.; Rizzato, P.P. (2012). "A new species of cave catfish from Brazil, Trichomycterus rubbioli sp.n., from Serra do Ramalho karstic area, São Francisco River basin, Bahia State (Silurifomes: Trichomycteridae)". Zootaxa 3480: 48–66. doi:10.11646/zootaxa.3480.1.2. http://www.lesbio.ufscar.br/assets/bichuette_rizzato_2012_trichomycterus_rubbioli.pdf. 
  33. ↑ Castellanos-Morales, C.A. (2011). "Trichomycterus sketi: a new species of subterranean catfish (Siluriformes: Trichomycteridae) from the Andean Cordillera of Colombia". Biota Colombiana 11 (1–2): 33–41. https://www.researchgate.net/publication/266417002. 
  34. ↑ DoNascimiento, C.; Prada-Pedreros, S.; Guerrero-Kommritz, J. (2014). "A new catfish species of the genus Trichomycterus (Siluriformes: Trichomycteridae) from the río Orinoco versant of Páramo de Cruz Verde, Eastern Cordillera of Colombia". Neotropical Ichthyology 12 (4): 717–728. doi:10.1590/1982-0224-20140005. http://www.scielo.br/pdf/ni/v12n4/1679-6225-ni-1982-0224-20140005.pdf. 
  35. ↑ Barbosa, M.A.; Costa, W.J.E.M. (2011). "Description of a new species of the catfish genus Trichomycterus (Teleostei: Siluriformes: Trichomycteridae) from the rio de Contas basin, northeastern Brazil". Vertebrate Zoology 61 (3): 307–312. doi:10.3897/vz.61.e31157. http://www.senckenberg.de/files/content/forschung/publikationen/vertebratezoology/vz61-3/03_vertebrate_zoology_61-3_barbosa_costa_307.pdf. 
  36. ↑ García-Melo, L.J.; Villa-Navarro, F.A.; DoNascimiento, C. (2016). "A new species of Trichomycterus (Siluriformes: Trichomycteridae) from the upper río Magdalena basin, Colombia". Zootaxa 4117 (2): 226–240. doi:10.11646/zootaxa.4117.2.6. PMID 27395171. http://www.mapress.com/j/zt/article/view/zootaxa.4117.2.6. 
  37. ↑ Ardila Rodríguez, C.A. (2016). "Trichomycterus torcoromaensis sp. nov. (Siluriformes: Trichomycteridae) una nueva especie de pez de la quebrada Torcoroma, cuenca del río Magdalena. Departamento de Norte de Santander – Colombia". Barranquilla, Departamento del Atlántico 2: 1–22. https://drive.google.com/file/d/0B2G5RQpBHXijT0dYQ3NRdzFqcEE/view. 
  38. ↑ Vilardo PJ; Katz AM; Costa WJEM (2020). "Relationships and description of a new species of Trichomycterus (Siluriformes: Trichomycteridae) from the Rio Paraíba do Sul basin, south-eastern Brazil". Zool Stud 59 (59): 53. doi:10.6620/ZS.2020.59-53. PMID 33456550. 

Wikidata ☰ Q584577 entry



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