Biology:Umenocoleidae
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Umenocoleidae is an extinct family of dictyopteran insects known from the Cretaceous. They are considered to be closely related to the Alienopteridae.[1] They were originally considered to be beetles[2] due to their beetle-like morphology, with sclerotised elytra-like forewings. This was probably an adaptation for living under bark and in other tight spaces.[1][3]
Systematics
- †Umenocoleus Chen and Tan 1973 Dalazi Formation, China, Early Cretaceous (Aptian); Zhonggou Formation, China, Early Cretaceous (Albian); Jinju Formation, South Korea, Early Cretaceous (Albian)
- †Ponopterix Vršanský and Grimaldi 1999 Crato Formation, Brazil, Early Cretaceous (Aptian)
- †Blattapterix Vršanský 2003 Xiagou Formation, China, Early Cretaceous (Aptian); Jinju Formation, South Korea, Early Cretaceous (Albian)
- †Elytropterix Vršanský 2003 Dzun-Bain Formation, Mongolia, Aptian
- †Petropterix Vršanský 2003 Dzun-Bain Formation, Mongolia; Zaza Formation, Russia; Kitadani Formation, Japan, Early Cretaceous (Aptian)[6]; Zhonggou Formation, China, Early Cretaceous (Albian); Jinju Formation, South Korea, Early Cretaceous (Albian)
- †Compunctiotypus Kaddumi 2005 Jordanian amber, Early Cretaceous (Albian)
- †Umenopterix Lee 2016 Crato Formation, Brazil, Early Cretaceous (Aptian) Burmese amber, Myanmar, mid-Cretaceous (latest Albian-earliest Cenomanian)
- †Alienopterix Mlynský, Vršanský et Wang, 2018 Burmese amber, Myanmar, mid-Cretaceous (latest Albian-earliest Cenomanian)
- †Enervipraeala Luo, Xu et Jarzembowski, 2021 Burmese amber, Myanmar, mid-Cretaceous (latest Albian-earliest Cenomanian)
- †Laticephalana Luo, Beutel, Xu et Jarzembowski, 2021 Burmese amber, Myanmar, mid-Cretaceous (latest Albian-earliest Cenomanian)
- †Archaeospinapteryx Sendi et al. 2023 Burmese amber, Myanmar, mid-Cretaceous (latest Albian-earliest Cenomanian)[5]
- †Trapezionotum Sendi et al. 2023 Burmese amber, Myanmar, mid-Cretaceous (latest Albian-earliest Cenomanian)[5]
- †Poikiloprosopon Sendi et al. 2023 Burmese amber, Myanmar, mid-Cretaceous (latest Albian-earliest Cenomanian)[5]
- †?Vzrkadlenie Burmese amber, Myanmar, mid Cretaceous (latest Albian-earliest Cenomanian) (Also placed in Alienopteridae or Cratovitismidae)
- †Nigropterix Sendi et al. 2023 Kzyl-Zhar, Kazakhstan, Late Cretaceous (Turonian)[5]; Burmese amber, Myanmar, mid-Cretaceous (latest Albian-earliest Cenomanian)[7]
- †Umenotypus Sendi et al. 2023 Kzyl-Zhar, Kazakhstan, Late Cretaceous (Turonian)[5]
- †Pseudoblattapterix Lee et al. 2024 Jinju Formation, South Korea, Early Cretaceous (Albian)
- †Chunxiangus Luo et al. 2025 Burmese amber, Myanmar, mid-Cretaceous (latest Albian-earliest Cenomanian)[8]
Jantaropterix Vršanský and Grimaldi 2003, Perspicuus Koubová, 2020, Lepidopterix Sendi, 2020 and Antophiloblatta Sendi, 2020 have been transferred to Cratovitismidae, while Vitisma Vršanský 1999 and Permoponopterix Nel, Prokop et Kirejtshuk, 2014 have been excluded from the family.[3]
References
- ↑ 1.0 1.1 Liuo, C.-H.; Beutel, R. G.; Thomson, U. R; Zheng, D.-R.; Li, J.-H.; Zhao, X.-Y.; Zhang, H.-C.; Wang, B. (2021). "Beetle or roach: systematic position of the enigmatic Umenocoleidae based on new material from Zhonggou Formation in Jiuquan, Northwest China, and a morphocladistic analysis". Palaeoworld 31: 121–130. doi:10.1016/j.palwor.2021.01.003.
- ↑ Chen, S.; Tan, C. C. (1973). "A new family of Coleoptera from the Lower Cretaceous of Kansu". Acta Entomologica Sinica 16 (2): 169–179. http://www.insect.org.cn/CN/Y1973/V16/I2/169.
- ↑ 3.0 3.1 3.2 Luo, Cihang; Beutel, Rolf G.; Engel, Michael S.; Liang, Kun; Li, Liqin; Li, Jiahao; Xu, Chunpeng; Vršanský, Peter et al. (February 2022). "Life history and evolution of the enigmatic Cretaceous–Eocene Alienopteridae: A critical review" (in en). Earth-Science Reviews 225. doi:10.1016/j.earscirev.2021.103914. https://linkinghub.elsevier.com/retrieve/pii/S0012825221004153.
- ↑ "Fossilworks: Umenocoleidae". https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=227708.
- ↑ 5.0 5.1 5.2 5.3 5.4 5.5 Sendi, H.; Le Tirant, S.; Palková, H.; Chorvát, D.; Šurka, J.; Cumming, R. (2023). "Umenocoleidae (Insecta: Dictyoptera) from Turonian sediments of Kzyl-Zhar, Kazakhstan and Cenomanian northern Myanmar amber". Biologia. doi:10.1007/s11756-023-01356-z.
- ↑ Oyama, Nozomu; Yukawa, Hirokazu; Imai, Takuya (10 January 2022). "New cockroach assemblage from the Lower Cretaceous Kitadani Formation, Fukui, Japan". Palaeontographica Abteilung A: 37–52. doi:10.1127/pala/2021/0112.
- ↑ Sendi, Hemen (15 March 2024). "Nigropterix cummingi sp. n., an umenocoleid (Insecta: Dictyoptera) from mid-Cretaceous northern Myanmar amber". Zootaxa 5424 (4): 476–482. doi:10.11646/zootaxa.5424.4.6.
- ↑ Luo, Deqiong; Xiao, Chuantao; Luo, Cihang (29 July 2025). "Chunxiangus shii gen. et sp. nov., a new genus and species of Umenocoleidae (Dictyoptera) from mid-Cretaceous Kachin amber". Palaeontologia Electronica 28 (2): 1–10. doi:10.26879/1568.
Wikidata ☰ Q21366583 entry
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