Biology:Volgadraco

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Volgadraco ("Volga River dragon") is a genus of pteranodontian pterosaur from the Upper Cretaceous of European Russia.

Discovery and naming

Volgadraco is known from lower beak (holotype SGU, no. 46/104a) and postcranial fragments from the early Campanian-age Rybushka Formation of Saratov, Russia. The size of this animal, and the development of blood supply in the lower jaw are intermediate between older Santonian or Turonian azhdarchids like Azhdarcho and Bakonydraco and later Maastrichtian azhdarchids like Quetzalcoatlus. Volgadraco was described in 2008 by Averianov, Arkhangelsky, and Pervushov. The type species is V. bogolubovi, the specific name honouring Russian paleontologist Nikolai Nikolaevich Bogolubov.[1]

Classification

Averianov and colleagues initially assigned Volgadraco to the family Azhdarchidae, and considered the earlier named genus Bogolubovia to be a nomen dubium that is potentially identical to Volgadraco.[1] In contrast, Longrich and colleagues in 2018 had recovered it within the family Nyctosauridae, the cladogram of which can be seen below:[2]

Pteranodontoidea
 

Ornithocheiromorpha

 
Pteranodontia
 
Pteranodontidae
 
 

Pteranodon sternbergi

 
 

Pteranodon longiceps

 
 
 

Tethydraco regalis

 
 
 
Nyctosauridae
 

Alamodactylus byrdi

 
 

Volgadraco bogolubovi

 
 
 

Cretornis hlavaci

 
 
 
 

Alcione elainus

 
 

Simurghia robusta

 
 
 
 

Muzquizopteryx coahuilensis

 
 
 
 

Barbaridactylus grandis

 
 

Nyctosaurus lamegoi

 
 
 
 

Nyctosaurus nanus

 
 

Nyctosaurus gracilis

 
 
 
 
 
 
 
 
 

In 2020, Averianov and Arkhangelsky reconsidered both Bogolubovia and Volgadraco as members of the family Pteranodontidae.[3] In 2022, Averianov and Kurin considered both Volgadraco and Bogolubovia as valid pteranodontids based on a new specimen of Bogolubovia.[4] In the same year, however, Volgadraco was considered to be a pteranodontian both outside the Nyctosauridae and Pteranodontidae based on phylogenetic analysis, the cladogram of which can be seen below:[5]

Pteranodontia
 
 

Tethydraco

 
 
 

Pteranodon longiceps

 
 

Pteranodon sternbergi (Geosternbergia)

 
 
 
 
 

Alamodactylus

 
 

Volgadraco

 
 
 

Cretornis

 
Aponyctosauria
 
 

Epapatelo

 
 

Simurghia

 
 

Alcione

 
 
Nyctosauridae
 

Muzquizopteryx

 
 
 
 

Nyctosaurus lamegoi

 
 

Nyctosaurus grandis (Barbaridactylus)

 
 
 
 

Nyctosaurus nanus

 
 

Nyctosaurus gracilis

 
 
 
 
 
 
 
 

See also

References

  1. ↑ 1.0 1.1 Averianov, A.O.; Arkhangelsky, M.S.; Pervushov, E.M. (October 2008). "A New Late Cretaceous Azhdarchid (Pterosauria, Azhdarchidae) from the Volga Region". Paleontological Journal 42 (6): 634–642. doi:10.1134/S0031030108060099. Bibcode: 2008PalJ...42..634A. 
  2. ↑ Longrich, Nicholas R.; Martill, David M.; Andres, Brian; Penny, David (2018). "Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary". PLOS Biology 16 (3). doi:10.1371/journal.pbio.2001663. PMID 29534059. 
  3. ↑ Alexander O. Averianov; Maxim S. Arkhangelsky (2020). "A large pteranodontid pterosaur from the Late Cretaceous of Eastern Europe". Geological Magazine 158 (7): 1143–1155. doi:10.1017/S0016756820001119. https://www.cambridge.org/core/journals/geological-magazine/article/abs/large-pteranodontid-pterosaur-from-the-late-cretaceous-of-eastern-europe/1D369709DD7ABE8EABAF492AB55B5E06. 
  4. ↑ "A new specimen of pteranodontid pterosaur Bogolubovia orientalis from the Upper Cretaceous of Penza Province, Russia". Historical Biology: An International Journal of Paleobiology 35 (8): 1288–1296. 2022. doi:10.1080/08912963.2022.2087522. 
  5. ↑ Fernandes, Alexandra E.; Mateus, Octávio; Andres, Brian; Polcyn, Michael J.; Schulp, Anne S.; Gonçalves, António Olímpio; Jacobs, Louis L. (2022). "Pterosaurs from the Late Cretaceous of Angola". Diversity 14 (9): 741. doi:10.3390/d14090741. 

Wikidata ☰ Q2210361 entry