Earth:Rybushka Formation

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Rybushka Formation
Stratigraphic range: Campanian
~84–71 Ma
TypeGeological formation
Lithology
PrimarySandstone
OtherPhosphorite
Location
Coordinates [ ⚑ ] : 51°18′N 45°00′E / 51.3°N 45.0°E / 51.3; 45.0
Paleocoordinates [ ⚑ ] 42°36′N 40°12′E / 42.6°N 40.2°E / 42.6; 40.2
RegionPenza, Saratov
CountryRussia
Rybushka Formation is located in Russia
Rybushka Formation
Rybushka Formation (Russia)

The Rybushka Formation is a upper Campanian geologic formation in the Penza and Saratov Oblasts of European Russia. The largest location of fossils is located near the village of Malaya Serdoba in the Penza region.

This formation is one of the largest in Europe in terms of its abundance of marine reptile and pterosaur remains, and the largest marine formation in Russia. One of the formation's unique features is the discovery of many species of fish, sharks, and marine reptiles previously known only from specific regions, such as North America and Western Europe. Pterosaur, fish and invertebrate fossils have been recovered from the formation.[1][2]

Fossil content

The following fossils have been reported from the formation:

See also

  • List of pterosaur-bearing stratigraphic units

References

  1. ↑ Sullivan, Robert M.; Lucas, Spencer G.; Spielmann, Justin A. (2011-01-01) (in en). Fossil Record 3: Bulletin 53. New Mexico Museum of Natural History and Science. https://books.google.com/books?id=fugoCgAAQBAJ. 
  2. ↑ Averianov, Alexander (2014-08-11). "Review of taxonomy, geographic distribution, and paleoenvironments of Azhdarchidae (Pterosauria)". ZooKeys (432): 1–107. doi:10.3897/zookeys.432.7913. ISSN 1313-2989. PMID 25152671. PMC 4141157. Bibcode: 2014ZooK..432....1A. https://archive.org/details/pubmed-PMC4141157. 
  3. ↑ Kanarkina, A.; Zverkov, Nikolay G.; Popov, Evgeny V. (2025). "The first record of the pachycormid fish Bonnerichthys in Eurasia marks a global distribution of Late Cretaceous suspension-feeding giants". Journal of Palaeogeography 14 (3): 100255. doi:10.1016/j.jop.2025.100255. 
  4. ↑ 4.0 4.1 "Современная палеонтология: классические и новейшие методы" (in ru). https://www.paleo.ru/upload/iblock/4ed/6jsb5wyqt65bcjkxvi3i77sbc4mb46b4.pdf. 
  5. ↑ [1]
  6. ↑ "New record of Hesperoricus rossicus" (PDF). http://cretaceous.ru/files/pub/people/popov/panteleev_et_al2004-cmp-hesperornis-saratov.pdf. 
  7. ↑ "MARINE FISHES (CHONDRICHTHYES, HOLOCEPHALI, ACTINOPTERYGII) FROM THE UPPER CRETACEOUS (CAMPANIAN) RYBUSHKA FORMATION NEAR BELOE OZERO, SARATOV OBLAST, RUSSIA" (PDF). https://www.researchgate.net/publication/359625149_MARINE_FISHES_CHONDRICHTHYES_HOLOCEPHALI_ACTINOPTERYGII_FROM_THE_UPPER_CRETACEOUS_CAMPANIAN_RYBUSHKA_FORMATION_NEAR_BELOE_OZERO_SARATOV_OBLAST_RUSSIA. 
  8. ↑ "RYBUSHKA, Russia". https://www.mindat.org/paleo_strat.php?id=14459. 
  9. ↑ "Fig. 8 -Teleost remains from the Beloe Ozero locality in the Saratov...". https://www.researchgate.net/figure/Teleost-remains-from-the-Beloe-Ozero-locality-in-the-Saratov-Oblast-Russia-A-B-MMO-1-No_fig4_359625149. 
  10. ↑ "Правая верхнечелюстная кость мозазавра - Кампан". https://www.ammonit.ru/foto/78964.htm. 
  11. ↑ "Новые данные о кампанской ихтиофауне Нижнего Поволжья" (in ru). http://cretaceous.ru/files/pub/2022/solonin_arkhangelsky_priroda2022.pdf#:~:text=%D1%81%D1%82%D0%B0%D0%B2%D0%BB%D0%B5%D0%BD%D1%8B%20%D0%B7%D0%B4%D0%B5%D1%81%D1%8C%20%D1%81%D0%BB%D0%B5%D0%B4%D1%83%D1%8E%D1%89%D0%B8%D0%BC%D0%B8%20%D1%82%D0%B0%D0%BA%D1%81%D0%BE%D0%BD%D0%B0%D0%BC%D0%B8:%20Pachyrhi%2D%20zodus%20sp.%2C%20Saurocephalus%20lanciformis%2C%20Paralbula%20casei%2C. 
  12. ↑ "Позднемеловые плезиозавры-поликотилиды Европейской России" (in ru). http://www.ginras.ru/p-science/files/Zverkov_Arkhangelsky_polycotylids_Priroda_10_2024.pdf. 
  13. ↑ "О находке clidastes propython cope (squamata, mosasauridae) в верхнем мелу саратовской области" (in ru). http://cretaceous.ru/files/pub/temp3/grigorevidr.pdf. 
  14. ↑ Grigoriev, D. V.; Arkhangelsky, M. S.; Zverkov, N. G.; Gubarev, D. I. (2025). "First record of a rare plioplatecarpine mosasaur Latoplatecarpus in Europe as further evidence of a semi-global Campanian marine vertebrate fauna". Cretaceous Research: 106102. doi:10.1016/j.cretres.2025.106102. 

Further reading

  • M. S. Arkhangelsky, A. O. Averianov, and E. M. Pervushov. 2007. Short-necked plesiosaurs of the Family Polycotylidae from the Campanian of the Saratov Region. Paleontological Journal 41(6):656-660
  • A. V. Averianov and E. V. Popov. 2014. A pterosaurian vertebra from the Upper Cretaceous of the Saratov Region. Paleontological Journal 48(3):326-329
  • A. O. Averianov, M. S. Arkhangelsky, and E. M. Pervushov. 2008. A new Late Cretaceous azhdarchid (Pterosauria, Azhdarchidae) from the Volga region. Paleontological Journal 42(6):634-642
  • A. O. Averianov. 2007. New records of azhdarchids (Pterosauria, Azhdarchidae) from the Late Cretaceous of Russia, Kazakhstan, and Central Asia. Paleontological Journal 41(2):189-197
  • N. N. Bogoliubow. 1914. O ozvonkye pterodaktilya iz verkhnemelovykh otlozheniy Saratovskoy gubernii [A pterodactyl vertebra from the Upper Cretaceous deposits of Saratov province]. Ezhegodnikie po Geologii i Mineralogii Rossii/Annuaire Géologique et Minéralogique de la Russie 16(1):1-7



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