Biology:Agraphydrus

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Short description: Genus of beetles

Agraphydrus
Dorsal view of Agraphydrussp.
Scientific classification e
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Coleoptera
Family: Hydrophilidae
Subfamily: Acidocerinae
Genus: Agraphydrus
Régimbart, 1903
Diversity
205 species
Synonyms
  • Gymnhelochares d’Orchymont, 1932
  • Pseudohelochares Satô, 1960
  • Horelophopsis Hansen, 1997
  • Pseudopelthydrus Jia, 1998
  • Megagraphydrus Hansen, 1999

Agraphydrus is a genus of water scavenger beetle in the family Hydrophilidae represented by 205 described species.[1] It is distributed across the Afrotropical, Australasian, and Indomalayan realms.

Taxonomy

The genus Agraphydrus was described for the first time by Régimbart in 1903.[2]

It currently contains 205 described species, partly thanks to a series revisionary work developed by Albretch Komarek.[3][4][5][6][7]

Description

Small beetles (1.4–4.8 mm), pale/yellowish to dark brown in coloration, eyes not emarginate, labrum exposed, maxillary palps moderately long. Elytra without sutural striae, not laterally explanate; the elytral punctation ranges from very fine and subtle to coarse.

A complete diagnosis was presented by Girón and Short,[1] but also see Komarek and collaborators.[3][4][5][6][7]

Habitat

According to Girón and Short,[1] based on annotations by Komarek and collaborators:[3][4][5][6][7]

Agraphydrus can be found in an extremely broad range of habitats, from rivers, streams and forest pools, to hygropetric environments around waterfalls or seepages over rocks; a few species have been collected in terrestrial habitats by sifting moss and leaves from near water bodies, or in the gravel along the bank of a river; in many cases specimens have been found associated with floating vegetation, mosses and algae.[1]

Species

  1. Agraphydrus abrasus Komarek and Freitag, 2020 [6]
  2. Agraphydrus activus Komarek and Hebauer, 2018 [7]
  3. Agraphydrus acutus Komarek, 2020 [5]
  4. Agraphydrus aethiopicus Komarek, 2020 [5]
  5. Agraphydrus agilis Komarek and Hebauer, 2018 [7]
  6. Agraphydrus albescens (Régimbart, 1903) [8]
  7. Agraphydrus ampullatus Komarek and Freitag, 2020 [6]
  8. Agraphydrus anacaenoides Komarek, 2019 [4]
  9. Agraphydrus anatinus Komarek, 2018 [3]
  10. Agraphydrus andamanicus Komarek, 2018 [3]
  11. Agraphydrus andringitra Komarek, 2020 [5]
  12. Agraphydrus angulatus Komarek, 2019 [4]
  13. Agraphydrus angustatus Komarek, 2020 [5]
  14. Agraphydrus angustipenis Komarek, 2018 [3]
  15. Agraphydrus anhuianus (Hebauer, 2000) [9]
  16. Agraphydrus annapurnensis Komarek, 2018 [3]
  17. Agraphydrus arduus Komarek and Hebauer, 2018 [7]
  18. Agraphydrus ater Komarek, 2018 [3]
  19. Agraphydrus atripalpis Komarek, 2020 [5]
  20. Agraphydrus attenuatus (Hansen, 1999) [10]
  21. Agraphydrus audax Komarek and Hebauer, 2018 [7]
  22. Agraphydrus avita (Hansen, 1997) [11]
  23. Agraphydrus bacchusi Komarek, 2019 [4]
  24. Agraphydrus balkeorum Komarek, 2019 [4]
  25. Agraphydrus batak Komarek and Freitag, 2020 [6]
  26. Agraphydrus bhutanensis Komarek, 2018 [3]
  27. Agraphydrus bicoloratus Komarek, 2020 [5]
  28. Agraphydrus bilardoi Komarek, 2020 [5]
  29. Agraphydrus biltoni Komarek, 2020 [5]
  30. Agraphydrus biprojectus Minoshima, Komarek, and Ôhara, 2015 [12]
  31. Agraphydrus borneensis Komarek, 2019 [4]
  32. Agraphydrus boukali Komarek, 2018 [3]
  33. Agraphydrus brevilobatus Komarek and Freitag, 2020 [6]
  34. Agraphydrus brevipenis Komarek, 2019 [4]
  35. Agraphydrus burmensis Komarek, 2019 [4]
  36. Agraphydrus calvus Komarek and Hebauer, 2018 [7]
  37. Agraphydrus camerunensis Komarek, 2020 [5]
  38. Agraphydrus cantonensis Komarek and Hebauer, 2018 [7]
  39. Agraphydrus carinatulus Komarek, 2019 [4]
  40. Agraphydrus cervus Komarek, 2019 [4]
  41. Agraphydrus ceylonensis Komarek, 2018 [3]
  42. Agraphydrus chinensis Komarek and Hebauer, 2018 [7]
  43. Agraphydrus cinnamum Komarek, 2018 [3]
  44. Agraphydrus clarus Komarek, 2019 [4]
  45. Agraphydrus comes Komarek and Hebauer, 2018 [7]
  46. Agraphydrus communis Komarek, 2018 [3]
  47. Agraphydrus confusus Komarek and Hebauer, 2018 [7]
  48. Agraphydrus congolensis Komarek, 2020 [5]
  49. Agraphydrus conicus Komarek and Hebauer, 2018 [7]
  50. Agraphydrus connexus Komarek and Hebauer, 2018 [7]
  51. Agraphydrus constrictus Komarek, 2018 [3]
  52. Agraphydrus contractus Komarek and Hebauer, 2018 [7]
  53. Agraphydrus coomani (d'Orchymont, 1927) [13]
  54. Agraphydrus coronarius Minoshima, Komarek, and Ôhara, 2015 [12]
  55. Agraphydrus crassipenis Komarek, 2018 [3]
  56. Agraphydrus dapengensis Yang & Jia, 2021 [14]
  57. Agraphydrus decipiens Minoshima, Komarek, and Ôhara, 2015 [12]
  58. Agraphydrus delineatus Komarek, 2019 [4]
  59. Agraphydrus elongatus Ribera, Hernando, and Cieslak, 2019 [15]
  60. Agraphydrus engkari Komarek, 2019 [4]
  61. Agraphydrus excisus Komarek, 2019 [4]
  62. Agraphydrus exedis (d'Orchymont, 1937) [16]
  63. Agraphydrus exiguus Komarek, 2019 [4]
  64. Agraphydrus falcatus Komarek, 2018 [3]
  65. Agraphydrus fasciatus Komarek and Hebauer, 2018 [7]
  66. Agraphydrus fikaceki Komarek and Hebauer, 2018 [7]
  67. Agraphydrus flavescens Komarek, 2020 [5]
  68. Agraphydrus flavipes Komarek, 2020 [5]
  69. Agraphydrus flavonotus Komarek, 2018 [3]
  70. Agraphydrus floresinus Komarek, 2019 [4]
  71. Agraphydrus fontis Komarek, 2020 [5]
  72. Agraphydrus forcipatus Komarek and Hebauer, 2018 [7]
  73. Agraphydrus fortis Komarek, 2018 [3]
  74. Agraphydrus fujianensis Komarek and Hebauer, 2018 [7]
  75. Agraphydrus geminus (d'Orchymont, 1932) [17]
  76. Agraphydrus gereckei Komarek, 2020 [5]
  77. Agraphydrus gilvus Komarek, 2018 [3]
  78. Agraphydrus glaber Komarek, 2018 [3]
  79. Agraphydrus globipenis Komarek and Hebauer, 2018 [7]
  80. Agraphydrus goldschmidti Komarek, 2020 [5]
  81. Agraphydrus gracilipalpis Komarek, 2018 [3]
  82. Agraphydrus hamatus Komarek, 2019 [4]
  83. Agraphydrus hanseni (Satô and Yoshitomi, 2004) [18]
  84. Agraphydrus heinrichi Komarek, 2018 [3]
  85. Agraphydrus helicopter Komarek, 2019 [4]
  86. Agraphydrus hendrichi Komarek, 2019 [4]
  87. Agraphydrus heterochromatus Komarek, 2019 [4]
  88. Agraphydrus hortensis Komarek, 2019 [4]
  89. Agraphydrus hygropetricus Komarek, 2018 [3]
  90. Agraphydrus igneus Komarek and Hebauer, 2018 [7]
  91. Agraphydrus imitans Komarek, 2019 [4]
  92. Agraphydrus indicus (d'Orchymont, 1932) [17]
  93. Agraphydrus inflatus Komarek, 2018 [3]
  94. Agraphydrus infuscatus Komarek, 2019 [4]
  95. Agraphydrus insidiator Minoshima, Komarek, and Ôhara, 2015 [12]
  96. Agraphydrus ishiharai (Matsui, 1994) [19]
  97. Agraphydrus jaechi (Hansen, 1999) [10]
  98. Agraphydrus jankodadai Komarek, 2019 [4]
  99. Agraphydrus jilanzhui Komarek and Hebauer, 2018 [7]
  100. Agraphydrus kallar Komarek, 2018 [3]
  101. Agraphydrus kathapa Komarek, 2019 [4]
  102. Agraphydrus kempi (d'Orchymont, 1922)
  103. Agraphydrus khasiensis Komarek, 2018 [3]
  104. Agraphydrus kodaguensis Komarek, 2018 [3]
  105. Agraphydrus komareki Yang & Jia, 2021 [14]
  106. Agraphydrus laocaiensis Komarek, 2019 [4]
  107. Agraphydrus latus Komarek, 2019 [4]
  108. Agraphydrus longipalpus (Jia, 1998) [20]
  109. Agraphydrus longipenis Komarek and Hebauer, 2018 [7]
  110. Agraphydrus lunaris Komarek, 2019 [4]
  111. Agraphydrus luteilateralis (Minoshima and Fujiwara, 2009) [21]
  112. Agraphydrus madagascarensis Komarek, 2020 [5]
  113. Agraphydrus maehongsonensis Komarek, 2019 [4]
  114. Agraphydrus malayanus (Hebauer, 2000) [9]
  115. Agraphydrus malkini Komarek, 2020 [5]
  116. Agraphydrus manfredjaechi Komarek, 2019 [4]
  117. Agraphydrus masatakai Minoshima, Komarek, and Ôhara, 2015 [12]
  118. Agraphydrus matoposensis Komarek, 2020 [5]
  119. Agraphydrus mazzoldii Komarek, 2019 [4]
  120. Agraphydrus meghalayanus Komarek, 2018 [3]
  121. Agraphydrus microphthalmus Komarek, 2019 [4]
  122. Agraphydrus minutissimus (Kuwert, 1890) [22]
  123. Agraphydrus mirabilis Komarek, 2019 [4]
  124. Agraphydrus montanus Minoshima, Komarek, and Ôhara, 2015 [12]
  125. Agraphydrus muluensis Komarek, 2019 [4]
  126. Agraphydrus musculus Komarek, 2019 [4]
  127. Agraphydrus namthaensis Komarek, 2019 [4]
  128. Agraphydrus nanus Komarek, 2018 [3]
  129. Agraphydrus narusei (Satô, 1960) [23]
  130. Agraphydrus nemorosus Komarek, 2019 [4]
  131. Agraphydrus nepalensis Komarek, 2018 [3]
  132. Agraphydrus niger Komarek, 2018 [3]
  133. Agraphydrus nigroflavus Komarek, 2019 [4]
  134. Agraphydrus obesus Komarek, 2019 [4]
  135. Agraphydrus obscuratus Komarek, 2018 [3]
  136. Agraphydrus obsoletus Komarek, 2018 [3]
  137. Agraphydrus occultus Komarek and Freitag, 2020 [6]
  138. Agraphydrus ogatai Minoshima, 2016 [24]
  139. Agraphydrus orbicularis Komarek, 2019 [4]
  140. Agraphydrus orientalis (d'Orchymont, 1932) [17]
  141. Agraphydrus palawanensis Komarek and Freitag, 2020 [6]
  142. Agraphydrus pallidus Komarek, 2019 [4]
  143. Agraphydrus papuanus Komarek, 2019 [4]
  144. Agraphydrus pauculus (Knisch, 1924) [25]
  145. Agraphydrus pauper Komarek, 2020 [5]
  146. Agraphydrus pelingeni Komarek and Freitag, 2020 [6]
  147. Agraphydrus penangensis Komarek, 2019 [4]
  148. Agraphydrus piceus Komarek, 2019 [4]
  149. Agraphydrus politus (Hansen, 1999) [10]
  150. Agraphydrus praecipuus (d'Orchymont, 1937) [26]
  151. Agraphydrus protentus Komarek, 2018 [3]
  152. Agraphydrus pseudoniger Yang & Jia, 2021 [14]
  153. Agraphydrus pullus Komarek, 2018 [3]
  154. Agraphydrus punctatellus Régimbart, 1903 [2]
  155. Agraphydrus punctulatus Komarek, 2018 [3]
  156. Agraphydrus puzhelongi (Jia, 2010) [27]
  157. Agraphydrus pygmaeus (Knisch, 1924) [25]
  158. Agraphydrus raucus Komarek, 2019 [4]
  159. Agraphydrus reductus Komarek and Hebauer, 2018 [7]
  160. Agraphydrus regularis (Hansen, 1999) [10]
  161. Agraphydrus reticulatus Komarek, 2019 [4]
  162. Agraphydrus reticuliceps Komarek and Hebauer, 2018 [7]
  163. Agraphydrus rhodesiensis Komarek, 2020 [5]
  164. Agraphydrus rhomboideus Komarek, 2019 [4]
  165. Agraphydrus rivalis Komarek, 2020 [5]
  166. Agraphydrus robustus Komarek and Hebauer, 2018 [7]
  167. Agraphydrus rostratus Komarek, 2018 [3]
  168. Agraphydrus rugosus Komarek, 2018 [3]
  169. Agraphydrus sabulosus Yang & Jia, 2021 [14]
  170. Agraphydrus sarawakensis Komarek, 2019 [4]
  171. Agraphydrus schoedli Komarek, 2019 [4]
  172. Agraphydrus schoenmanni Komarek and Hebauer, 2018 [7]
  173. Agraphydrus scintillans Komarek, 2019 [4]
  174. Agraphydrus scutifer Komarek, 2020 [5]
  175. Agraphydrus setifer Komarek and Hebauer, 2018 [7]
  176. Agraphydrus shaverdoae Komarek, 2019 [4]
  177. Agraphydrus siamensis (Hansen, 1999) [10]
  178. Agraphydrus sipekorum Komarek, 2018 [3]
  179. Agraphydrus skalei Komarek, 2019 [4]
  180. Agraphydrus spadix Komarek, 2019 [4]
  181. Agraphydrus spinosus Komarek, 2019 [4]
  182. Agraphydrus splendens Komarek and Hebauer, 2018 [7]
  183. Agraphydrus stagnalis (d'Orchymont, 1937)
  184. Agraphydrus stramineus Komarek, 2019 [4]
  185. Agraphydrus sucineus Komarek, 2019 [4]
  186. Agraphydrus sundaicus Komarek, 2019 [4]
  187. Agraphydrus tamdao Komarek, 2019 [4]
  188. Agraphydrus taprobanensis Komarek, 2018 [3]
  189. Agraphydrus tenuipalpis Komarek and Freitag, 2020 [6]
  190. Agraphydrus thaiensis Minoshima, Komarek, and Ôhara, 2015 [12]
  191. Agraphydrus tristis Komarek, 2019 [4]
  192. Agraphydrus tulipa Komarek, 2019 [4]
  193. Agraphydrus tumidus Komarek, 2020 [5]
  194. Agraphydrus tumulosus Komarek, 2018 [3]
  195. Agraphydrus umbrosus Komarek and Hebauer, 2018 [7]
  196. Agraphydrus uncinatus Komarek and Hebauer, 2018 [7]
  197. Agraphydrus usambaraensis Komarek, 2020 [5]
  198. Agraphydrus uvaensis (Hebauer, 2000) [9]
  199. Agraphydrus vadoni Komarek, 2020 [5]
  200. Agraphydrus variabilis Komarek and Hebauer, 2018 [7]
  201. Agraphydrus vietnamensis Komarek, 2019 [4]
  202. Agraphydrus villiersi (Balfour-Browne, 1958) [28]
  203. Agraphydrus wangmiaoi Komarek and Hebauer, 2018 [7]
  204. Agraphydrus yunnanensis Komarek and Hebauer, 2018 [7]
  205. Agraphydrus zetteli Komarek and Freitag, 2020 [6]

References

  1. ↑ 1.0 1.1 1.2 1.3 Girón, Jennifer C.; Short, Andrew Edward Z. (2021-06-18). "The Acidocerinae (Coleoptera, Hydrophilidae): taxonomy, classification, and catalog of species". ZooKeys (1045): 1–236. doi:10.3897/zookeys.1045.63810. ISSN 1313-2970. PMID 34228772. PMC 8233300. https://zookeys.pensoft.net/article/63810/. 
  2. ↑ 2.0 2.1 Régimbart, MA (1903). "Coléoptères aquatiques (Haliplidae, Dytiscidae, Gyrinidae et Hydrophilidae) recueillis dans le Sud de Madagascar par M. Ch. Alluaud (Juillet 1900 – Mai 1901)". Annales de la Société Entomologique de France 72: 1–51. https://biodiversitylibrary.org/page/8247889. 
  3. ↑ 3.00 3.01 3.02 3.03 3.04 3.05 3.06 3.07 3.08 3.09 3.10 3.11 3.12 3.13 3.14 3.15 3.16 3.17 3.18 3.19 3.20 3.21 3.22 3.23 3.24 3.25 3.26 3.27 3.28 3.29 3.30 3.31 3.32 3.33 3.34 3.35 3.36 3.37 3.38 3.39 3.40 Komarek, A. (2018). "Taxonomic revision of AgraphydrusRégimbart, 1903 II. The Indian Subcontinent". Koleopterologische Rundschau 88: 103–204. http://www.coleoptera.at/uploads/www.coleoptera.at/KOR_88_2018_0103-0204.pdf. Retrieved 2022-01-07. 
  4. ↑ 4.00 4.01 4.02 4.03 4.04 4.05 4.06 4.07 4.08 4.09 4.10 4.11 4.12 4.13 4.14 4.15 4.16 4.17 4.18 4.19 4.20 4.21 4.22 4.23 4.24 4.25 4.26 4.27 4.28 4.29 4.30 4.31 4.32 4.33 4.34 4.35 4.36 4.37 4.38 4.39 4.40 4.41 4.42 4.43 4.44 4.45 4.46 4.47 4.48 4.49 4.50 4.51 4.52 4.53 4.54 4.55 4.56 4.57 4.58 4.59 4.60 4.61 4.62 Komarek, A. (2019). "Taxonomic revision of AgraphydrusRégimbart, 1903 III. Southeast Asia (except Philippines) and Australian Region (Coleoptera: Hydrophilidae: Acidocerinae)". Koleopterologische Rundschau 89: 151–316. http://www.coleoptera.at/uploads/www.coleoptera.at/KOR_89_2019_0151-0316.pdf. Retrieved 2022-01-07. 
  5. ↑ 5.00 5.01 5.02 5.03 5.04 5.05 5.06 5.07 5.08 5.09 5.10 5.11 5.12 5.13 5.14 5.15 5.16 5.17 5.18 5.19 5.20 5.21 5.22 5.23 5.24 5.25 5.26 5.27 Komarek, A. (2020). "Taxonomic revision of AgraphydrusRégimbart, IV. Africa, Western Asia, and redescription of the genus (Coleoptera: Hydrophilidae: Acidocerinae)". Koleopterologische Rundschau 90: 127–200. http://www.coleoptera.at/uploads/www.coleoptera.at/KOR_90_2020_0127-0200.pdf. Retrieved 2022-01-07. 
  6. ↑ 6.00 6.01 6.02 6.03 6.04 6.05 6.06 6.07 6.08 6.09 6.10 6.11 Komarek, A.; Freitag, H. (2020). "Taxonomic revision of AgraphydrusRégimbart, 1903 V. Philippine species and their first DNA barcodes". Koleopterologische Rundschau 90: 201–242. http://www.coleoptera.at/uploads/www.coleoptera.at/KOR_90_2020_0127-0200.pdf. Retrieved 2022-01-07. 
  7. ↑ 7.00 7.01 7.02 7.03 7.04 7.05 7.06 7.07 7.08 7.09 7.10 7.11 7.12 7.13 7.14 7.15 7.16 7.17 7.18 7.19 7.20 7.21 7.22 7.23 7.24 7.25 7.26 7.27 7.28 7.29 7.30 7.31 7.32 7.33 Komarek, A.; Hebauer, F. (2018-07-30). "Taxonomic revision of Agraphydrus Régimbart, 1903 I. China and Taiwan (Coleoptera: Hydrophilidae: Acidocerinae)". Zootaxa 4452 (1): 1–101. doi:10.11646/zootaxa.4452.1.1. ISSN 1175-5334. PMID 30314249. https://doi.org/10.11646/zootaxa.4452.1.1. 
  8. ↑ Régimbart, M. (1903). "Coléoptères aquatiques (Haliplidae, Dytiscidae, Gyrinidae et Hydrophi-lidae) recueillis dans le Sud de Madagascar par M. Ch. Alluaud (Juillet 1900 – Mai 1901)". Annales de la Société Entomologique de France 72: 1–51. https://biodiversitylibrary.org/page/8247889. 
  9. ↑ 9.0 9.1 9.2 Hebauer, F. (2000). "The genus Megagraphydrus Hansen, 1999, with description of new species (Coleoptera: Hydrophilidae)". Acta Coleopterologica 16: 14–22. 
  10. ↑ 10.0 10.1 10.2 10.3 10.4 Hansen, Michael (1999-01-01). "Fifteen new genera of Hydrophilidae (Coleoptera), with remarks on the generic classification of the family" (in en). Insect Systematics & Evolution 30 (2): 121–172. doi:10.1163/187631200X00228. ISSN 1876-312X. https://brill.com/view/journals/ise/30/2/article-p121_1.xml. 
  11. ↑ Hansen, M. (1997). "A new subfamily for a remarkable new genus and species of Hydrophilidae from New Guinea (Coleoptera: Hydrophilidae)". Annales Zoologici 47: 107–110. 
  12. ↑ 12.0 12.1 12.2 12.3 12.4 12.5 12.6 Minoshima, Y. N.; Komarek, A.; Ôhara, M. (2015). "A revision of Megagraphydrus Hansen (Coleoptera, Hydrophilidae): synonymization with Agraphydrus Régimbart and description of seven new species". Zootaxa 3930: 1–63. PMID 25781810. https://www.biotaxa.org/Zootaxa/article/view/1%E2%80%9363. 
  13. ↑ d’Orchymont, A. (1927). "Papers on Malayan aquatic biology. V. Notes on the Hydrophilidae in the Federated Malay States Museums". Journal of the Federated Malay States Museums 13 (4): 246–252. 
  14. ↑ 14.0 14.1 14.2 14.3 Yang, Zhen-ming; Jia, Fenglong; Jiang, Lu; Guo, Qiang (2021-05-18). "Four new species of Agraphydrus Régimbart, 1903 with additional faunistic record from China (Coleoptera, Hydrophilidae, Acidocerinae)". Deutsche Entomologische Zeitschrift 68 (1): 189–205. doi:10.3897/dez.68.66200. ISSN 1860-1324. https://dez.pensoft.net/article/66200/. 
  15. ↑ Ribera, I.; Hernando, C.; Cieslak, A. (2019). "Aquatic Coleoptera of North Oman, with description of new species of Hydraenidae and Hydrophilidae". Acta Entomologica Musei Nationalis Pragae 59: 253–272. doi:10.2478/aemnp-2019-0021. http://aemnp.eu/PDF/59_1/59_1_253.pdf. 
  16. ↑ d’Orchymont, A. (1937). "Descriptions of three new Hydrophilidae from India". Records of the Indian Museum 39: 29–31. 
  17. ↑ 17.0 17.1 17.2 d’Orchymont, A. (1932). "Zur Kenntnis der Kolbenwasserkäfer (Palpicornia) von Sumatra, Java und Bali". Archiv für Hydrobiologie Supplement Band IX: 623–714. [pl. XIV–XVII.]. 
  18. ↑ Satô, M.; Yoshitomi, H. (2004). "Discovery of a second representative of the genus Horelophopsis (Coleoptera, Hydrophilidae) from the Ryukyu archipelago, Japan". Elytra 32: 41–49. 
  19. ↑ Matsui, E. (1994). "Three new species of the genus Enochrus from Japan and Taiwan (Coleoptera: Hydrophilidae)". Transactions of the Shikoku Entomological Society 20: 215–220. 
  20. ↑ Jia, F. (1998). "A new genus Pseudopelthydrus gen. n. from Hainan Island, China (Coleoptera: Hydrophilidae: Hydrophilinae)". Chinese Journal of Entomology 18: 225–230. 
  21. ↑ Minoshima, Y. N.; Fujiwara, J. (2009). "Discovery of water scavenger beetle genus Megagraphydrus Hansen (Coleoptera, Hydrophilidae) from Japan, with description of a new species". Elytra 37: 53–64. http://coleoptera.sakura.ne.jp/Elytra/37(1)053MinoshimaY_and_FujiwaraJ.pdf. 
  22. ↑ Kuwert, A. (1890). "Bestimmungs-Tabellen der europäischen Coleopteren. XX. Heft. Hydrophilidae. II. Abteilung: Sphaeridiini und Helophorini". Verhandlungen des naturforschenden Vereins in Brünn 28: 159–328. https://biodiversitylibrary.org/page/43763542. 
  23. ↑ Satô, M. (1960). "One new genus and two new species of the subtribe Helocharae from Japan (Coleoptera: Hydrophilidae)". Transactions of the Shikoku Entomological Society 6: 76–80. 
  24. ↑ Minoshima, Yûsuke N. (2016). "Taxonomic review of Agraphydrus from Japan (Coleoptera: Hydrophilidae: Acidocerinae)". Entomological Science 19 (4): 351–366. doi:10.1111/ens.12213. ISSN 1479-8298. https://onlinelibrary.wiley.com/doi/abs/10.1111/ens.12213. 
  25. ↑ 25.0 25.1 Knisch, A. (1924). "Neue neotropische Palpicornier. (Col. Hydrophilidae. - Op. 16.)". Wiener Entomologische Zeitung 41: 114–140. 
  26. ↑ d’Orchymont, A. (1937). "Contribution à l'étude des Palpicornia IX". Bulletin et Annales de la Société Entomologique de Belgique 77: 213–255. 
  27. ↑ Jia, Fenglong (2010-06-08). "Megagraphydrus puzhelongi, sp. n., a new water scavenger beetle from China (Coleoptera: Hydrophilidae: Hydrophilinae)". Zootaxa 2498 (1): 65. doi:10.11646/zootaxa.2498.1.6. ISSN 1175-5334. https://biotaxa.org/Zootaxa/article/view/zootaxa.2498.1.6. 
  28. ↑ Balfour-Browne, J. (1958). "La Reserve naturelle integrale du Mont Nimba, Guinea. VI. Coléoptères Hydrophiloides". Mémoires de l'Institut français d'Afrique noire 53: 169–190. 

Wikidata ☰ Q21215758 entry



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