Biology:Endomychidae
Endomychidae, also commonly known as handsome fungus beetles, is a family of coccinelloid beetles with representatives found in all biogeographic realms. As the common name suggests, members of this family feed on fungi.
Description
They have a tarsal formal of 4-4-4 or 3-3-3 and the wings lack a closed radial cell. The second antennal segment has a sensory appendage that is as long as the third antennal segment.
Taxonomy
The family was established based on the type genus Endomychus, a genus erected in 1795 by Panzer which was applied to a species that Linnaeus called Chrysomela coccinea.
Crowson, in his influential treatment of the beetles, placed the family within the Cucujoidea. The family has also been grouped with the Coccinellidae in a group called the Trimera for having pseudotrimerous tarsi.[1] A 2015 molecular phylogeny study found that the Cucujoidea were found to be non-monophyletic and the Endomychidae was refined with the removal of the Anamorphinae from within the family and elevated to the status of a full family, Anamorphidae. Mycetaeinae and Eupsilobiinae were also found not to belong within the clades of the core Endomychidae, and likewise reclassified into the families Mycetaeidae and Eupsilobiidae.[2]
This family contains around 120 genera and 1300 species. A list of subfamilies can be seen below:
The subfamilies that are included:
- Danascelinae
- Endomychinae (including Stenotarsinae)
- Epipocinae
- Leiestinae
- Lycoperdininae
- Merophysiinae
- Pleganophorinae
- Xenomycetinae
Genera
These 70 genera belong to the family Endomychidae as defined in the past (due to revisions, this may not be entirely accurate):
- Acinaces Gerstaecker, 1858 g
- Aclemmysa Reitter, 1904 g
- Agaricophilus Motschulsky, 1838 g
- Amphisternus Germar, 1843 g
- Amphistethus Strohecker, 1964 g
- Ancylopus Costa, 1854 g
- Anidrytus Gerstaecker, 1858 i c g
- Aphorista Gorham, 1873 i c g b
- Archipines Strohecker, 1953 i c g
- Asymbius Gorham, 1896 g
- Atopomychus Tomaszewska & Szawaryn, 2013 g
- Atrichonota Arrow, 1925 g
- Avencymon Strohecker, 1971 g
- Beccariola Arrow, 1943 g
- Bolbomorphus Gorham, 1888 g
- Brachytrycherus Arrow, 1920 g
- Bystodes Strohecker, 1953 g
- Cholovocera Motschulsky, 1838 g
- Chondria Gorham, 1888 g
- Corynomalus Chevrolat in Dejean, 1836 i c g
- Cyclotoma Mulsant, 1851 i g
- Danae Reiche, 1847 i c g b
- Danascelis Tomaszewska, 1999 i c g
- Dapsa Latreille, 1829 g
- Dexialia Sasaji, 1970 g
- Dialexia Gorham, 1891 g
- Discolomopsis Shockley, 2006 g
- Displotera Reitter, 1877 g
- Ectomychus Gorham, 1888 g
- Eidorus g
- Endomychus Panzer, 1795 i c g b
- Ephebus Chevrolat in Dejean, 1836 i c g
- Epipocus Germar, 1843 i c g b
- Epopterus Chevrolat in Dejean, 1836 i c g
- Eucteanus Gerstaecker, 1857 g
- Eucymon g
- Eumorphus Weber, 1801 g
- Geoendomychus Lea, 1922 g
- Glesirhanis Shockley & Alekseev, 2014 g
- Hadromychus Bousquet & Leschen, 2002 i c g b
- Holoparamecus Curtis, 1833 i c g b
- Hylaia Guerin, 1857 g
- Hylaperdina Tomaszewska, 2012 g
- Idiophyes Blackburn, 1896 g
- Indalmus Gerstaecker, 1858 g
- Leiestes Chevrolat, 1836 g
- Lycoperdina Latreille, 1807 i c g b
- Meilichius Gerstaecker, 1857 g
- Merophysia Lucas, 1852 g
- Microxenus Wallaston, 1861 g
- Mycetina Mulsant, 1846 i c g b
- Mychothenus Strohecker, 1953 g
- Natalinus Tomaszewska, 2011 g
- Ohtaius Chûjô, 1938 g
- Palaeoestes Kirejtshuk & Nel, 2009 g
- Paniegena Heller, 1916 g
- Parasymbius Arrow, 1920 g
- Phymaphora Newman, 1838 i c g b
- Phymaphoroides Motschoulsky, 1857 g
- Pleganophorus Hampe, 1855 g
- Polymus Mulsant, 1846 g
- Pseudindalmus Arrow, 1920 g
- Rhanidea Strohecker, 1953 i c g b
- Saula Gerstaecker, 1858 i c g
- Sinocymbachus Strohecker & Chujo, 1970 g
- Sinopanamomus Esser, 2019
- Stenotarsus Perty, 1832 i c g b
- Stethorhanis Blaisdell, 1931 i c g
- Trochoideus Westwood, 1833 i c g b
- Trycherus Gerstaecker, 1857 g
- Xenomycetes Horn, 1880 i c g b
Data sources: i = ITIS,[3] c = Catalogue of Life,[4] g = GBIF,[5] b = Bugguide.net[6]
Fossil genera
- Burmalestes Tomaszewska and Ślipiński, 2018[7] Burmese amber, Myanmar, Late Cretaceous (Cenomanian)
- Cretolestes Tomaszewska, Ślipiński and Ren, 2018[7] Burmese amber, Myanmar, Cenomanian
- Cretaparamecus Tomaszewska, Ślipiński, Bai and Zhang, 2018[7] Burmese amber, Myanmar, Cenomanian
- Palaeomycetes Tomaszewska, Ślipiński and Ren, 2018[7] Burmese amber, Myanmar, Cenomanian
- Zemyna Tomaszewska, 2018[7] (replaced name Laima Alekseev and Tomaszewska, 2018): Baltic amber, Eocene
References
- ↑ Tomaszewska, K.Wioletta (2000). "Morphology, phylogeny and classification of adult endomychidae (Coleoptera: Cucujoidea)". Annales Zoologici 50 (4): 449–558. http://rcin.org.pl/Content/56456/WA058_74047_P255-T50_Annal-Zool-nr-4-1.pdf.
- ↑ Robertson, James A; Ślipiński, Adam; Moulton, Matthew; Shockley, Floyd W; Giorgi, Adriano; Lord, Nathan P; McKenna, Duane D; Tomaszewska, Wioletta et al. (2015). "Phylogeny and classification of Cucujoidea and the recognition of a new superfamily Coccinelloidea (Coleoptera: Cucujiformia)". Systematic Entomology 40 (4): 745. doi:10.1111/syen.12138. Bibcode: 2015SysEn..40..745R. https://repository.si.edu/bitstream/handle/10088/21810/ent_Shockley_Tomaszewska_McHugh_2009b.pdf.
- ↑ "Endomychidae Report". https://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=114369.
- ↑ "Browse Endomychidae". http://www.catalogueoflife.org/col/browse/tree/id/a4a75dd67401cc5ba261577dec2e7526.
- ↑ "Endomychidae". https://www.gbif.org/species/7828.
- ↑ "Endomychidae Family Information". https://bugguide.net/node/view/13860.
- ↑ 7.0 7.1 7.2 7.3 7.4 Tomaszewska, Wioletta; Ślipiński, Adam; Bai, Ming; Zhang, Weiwei; Ren, Dong (November 2018). "The oldest representatives of Endomychidae (Coleoptera: Coccinelloidea) from the Upper Cretaceous Burmese amber" (in en). Cretaceous Research 91: 287–298. doi:10.1016/j.cretres.2018.07.001. Bibcode: 2018CrRes..91..287T. https://linkinghub.elsevier.com/retrieve/pii/S0195667118301605.
External links
Wikidata ☰ Q1076939 entry
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