Biology:Theretairus

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Short description: Extinct genus of reptiles


Theretairus
Temporal range: Late Jurassic
Illustration of the Theretairus antiquus holotype mandible.
Scientific classification
Kingdom:
Animalia
Phylum:
Class:
Reptilia
Superorder:
Order:
Genus:
Theretairus
Type species
†Theretairus antiquus
Simpson, 1926

Theretairus is a Late Jurassic genus of sphenodont reptile from the Morrison Formation of western North America,[1] present in stratigraphic zones 5 and 6.[2]

History and naming

The holotype consists of a right mandible and several in-socket teeth from Quarry 9 at Como Bluff, Wyoming where it was excavated by paleontologist William Harlow Reed, who then was working for Othniel Charles Marsh,[1][3] and it was deposited at the Yale Peabody Museum under YPM VP 13764.[3] The locality comes from strata of the Brushy Basin Member of the Morrison Formation, which also bears other sphenodontians Opisthias and Eilenodon.[4][3] It was not named until George Gaylord Simpson described it as a new genus and species in a 1926 paper in the American Journal of Science. The generic name means "mammal companion", due to its association with the jaws of mammals in the Como Bluff quarry, and the specific name means "ancient", due to its old age as a sphenodontian.[3]

Description and classification

The type specimen preserves a very strange morphology, with masseteric fossae in the posterior end similar to those in mammals. The teeth progressively get larger as they get closer to the anterior end, a trait also similar to mammals.[3] Recently, the symphysis and two (perhaps three) enlarged acrodont fangs at the anterior end of the jaw, but separated one from the other to a lesser degree of the type have linked the taxon to the Mexican taxon Sphenovipera. The anterior border of the symphysis is steeply inclined in the holotype of Theretarius, suggesting it is an adult or older individual. Theretarius preserves five fossae along the posterior end of the mandible, making the mandible lighter, a trait only shared with Sphenovipera.[5]C

The following is a cladogram of Rhynchocephalia after DeMar et al. 2022.[6]

 
 

Younginia capensis

 
 

Prolacerta broomi

 
Lepidosauria
Pan-Squamata
 

Sophineta cracoviensis

 
 

Pristidactylus

 
 

Eichstaettisaurus schroederi

 
 
 

Megachirella wachtleri

 
 

Marmoretta oxoniensis

 
 
 
Rhynchocephalia
 

Gephyrosaurus bridensis

 
Sphenodontia
 

Diphydontosaurus avonis

 
Acrosphenodontia
 

Planocephalosaurus robinsonae

 
 
 

Rebbanasaurus jaini

 
 
 
 

Godavarisaurus lateefi

 
 

Theretairus antiquus

 
 
Eusphenodontia
 

Polysphenodon mulleri

 
 
 

Opisthiamimus gregori

 
 
Clevosauridae
 
 

Clevosaurus convallis

 
 
 

Clevosaurus brasiliensis

 
 

Clevosaurus hadroprodon

 
 
 
 
 

Clevosaurus bairdi

 
 
 

Clevosaurus hudsoni

 
 

Clevosaurus cambrica

 
 
 
 
Neosphenodontia
 

Brachyrhinodon taylori

 
 
 

Colobops noviportensis

 
 
Sphenodontidae
 

Sphenodon punctatus (tuatara)

 
 

Cynosphenodon huizachalensis

 
 

Sphenovipera jimmysjoyi

 
 
 

Kawasphenodon expectatus

 
 

Kawasphenodon peligrensis

 
 
 

Pelecymala robustus

 
Eilenodontinae
 
 

Fraserosphenodon latidens

 
 

Opisthias rarus

 
 
 
 

Eilenodon robustus

 
 
 

Sphenotitan leyesi

 
 
 

Toxolophosaurus cloudi

 
 

Priosphenodon avelasi

 
 
 
 
 
 
Leptorhynchia
 
 

Homoeosaurus maximiliani

 
 

Kallimodon pulchellus

 
 
 
 
 

Sigmala sigmala

 
 

Vadasaurus herzogi

 
 
Pleurosauridae
 

Palaeopleurosaurus posidonae

 
 
 

Pleurosaurus goldfussi

 
 

Pleurosaurus ginsburgi

 
 
 
 
 

Kallimodon cerinensis

 
Sapheosauridae
 

Sapheosaurus thiollierei

 
 
 

Ankylosphenodon pachyostosus

 
 

Oenosaurus muehlheimensis

 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 
 

See also

References

  1. ↑ 1.0 1.1 Foster, J. (2007). "Table 2.1: Fossil Vertebrates of the Morrison Formation." Jurassic West: The Dinosaurs of the Morrison Formation and Their World. Indiana University Press. pp. 58-59.
  2. ↑ Foster, J. (2007). "Appendix." Jurassic West: The Dinosaurs of the Morrison Formation and Their World. Indiana University Press. pp. 327-329.
  3. ↑ 3.0 3.1 3.2 3.3 3.4 Simpson, G. G. (1926-07-01). "American terrestrial Rhynchocephalia" (in en). American Journal of Science s5-12 (67): 12–16. doi:10.2475/ajs.s5-12.67.12. ISSN 0002-9599. http://www.ajsonline.org/cgi/doi/10.2475/ajs.s5-12.67.12. 
  4. ↑ Rasmussen, T. E.; Callison, George (1981). "A New Herbivorous Sphenodontid (Rhynchocephalia: Reptilia) from the Jurassic of Colorado". Journal of Paleontology. 55 (5): 1109–1116. JSTOR 1304536.
  5. ↑ Reynoso, Víctor-Hugo (2005-09-30). [0646:PEOAVA2.0.CO;2 "Possible evidence of a venom apparatus in a Middle Jurassic Sphenodontian from the Huizachal red beds of Tamaulipas, México"]. Journal of Vertebrate Paleontology 25 (3): 646–654. doi:10.1671/0272-4634(2005)025[0646:PEOAVA2.0.CO;2]. ISSN 0272-4634. https://doi.org/10.1671/0272-4634(2005)025[0646:PEOAVA]2.0.CO;2. 
  6. ↑ DeMar, David G.; Jones, Marc E. H.; Carrano, Matthew T. (2022-12-31). "A nearly complete skeleton of a new eusphenodontian from the Upper Jurassic Morrison Formation, Wyoming, USA, provides insight into the evolution and diversity of Rhynchocephalia (Reptilia: Lepidosauria)" (in en). Journal of Systematic Palaeontology 20 (1): 1–64. doi:10.1080/14772019.2022.2093139. ISSN 1477-2019. https://www.tandfonline.com/doi/full/10.1080/14772019.2022.2093139. 

Wikidata ☰ Q7782944 entry