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Short description
Order of fungi

Scientific classification

Lichtw. & Manier (1978)

The Harpellales are an order of fungi classified in the subdivision Kickxellomycotina. Thalli are either unbranched or branched, producing basipetal series of trichospores. Zygospores are biconical. Species in the order are found attached to the gut lining of aquatic larvae of Insecta or (rarely) Isopoda. Harpellales are divided into two families, the Harpellaceae and the Legeriomycetaceae.[1] According to the Dictionary of the Fungi (10th edition, 2008), the order contains 38 genera and 200 species.[2] The order was formally described in 1978 Mycotaxon publication.[3] Harpellales has served as a model to study and understand the evolution, growth, and biodiversity of other such fungi found in the gut as species are plentiful around the world.[4]

Species include Allantomyces zopilotei, Bojamyces olmecensis, Gauthieromyces viviparus and Graminella ophiuroidea.[5]


  1. "Zygomycota". 
  2. Dictionary of the Fungi (10th ed.). Wallingford, UK: CABI. 2008. p. 305. ISBN 978-0-85199-826-8. 
  3. "Validation of the Harpellales and Asellariales". Mycotaxon 7 (3): 441–42. 1978. 
  4. Wang, Yan; White, Merlin M.; Moncalvo, Jean-Marc (October 2019). "Diversification of the gut fungi Smittium and allies (Harpellales) co-occurred with the origin of complete metamorphosis of their symbiotic insect hosts (lower Diptera)". Molecular Phylogenetics and Evolution 139: 106550. doi:10.1016/j.ympev.2019.106550. PMID 31279967. 
  5. "Harpellales in the digestive tracts of Ephemeroptera and Plecoptera nymphs from Veracruz, Mexico". Mycologia 100 (1): 149–62. 2008. doi:10.3852/mycologia.100.1.149. PMID 18488361. 

See also Wikidata entry Q1055276.