Biology:Hymenasplenium

From HandWiki

Hymenasplenium is one of three genera of ferns in the Aspleniaceae (spleenwort family), in the eupolypods II clade of the order Polypodiales.[1][2] The others are Hemidictyum and Asplenium. Hymenasplenium was segregated because it is a natural grouping with differing rhizome morphology – dorsiventral v. radial for the rest of Asplenium, differing chromosome count – x=39 v. x=36 for the rest of Asplenium, and a clear monophyletic grouping based on genetic analysis. It was confirmed as a sister group to Asplenium in a 2015 molecular study of the genera.[3]

List of species

As of July 2025, the Checklist of Ferns and Lycophytes of the World recognized the sixty-seven species and three hybrids below.[4]

  • Hymenasplenium trapeziforme (Wall. ex Roxb.) comb.ined., Asplenium trapeziforme Wall. ex Roxb.


Hybrids

  • Hymenasplenium × incisoserratum (Rosenst.) comb.ined., Asplenium × incisoserratum (Rosenst.) N.Murak. & R.C.Moran

Phylogeny

Phylogeny from Fern Tree of Life[5][6]
Hymenasplenium
 
 
 

H. delitescens (Maxon) Regalado & Prada

 
 

H. hoffmannii (Hieronymus) Regalado & Prada

 
 
 
 
 

H. laetum (Swartz) Regalado & Prada

 
 

H. praestans (Copeland) Testo, Riibe & Sundue

 
 
 
 

H. triquetrum (N.Murak. & Moran) Regalado & Prada

 
 
 

H. riparium (Liebmann) Regalado & Prada

 
 

H. volubile (N.Murak. & Moran) Regalado & Prada

 
 
 
 
 
 
 
 
 

H. phamhoanghoi Zhang, Xu & Luong

 
 
 
 

H. excisum (Presl) S.Linds. 2009 (Scalloped spleenwort)

 
 

H. obscurum (Blume) Tagawa

 
 
 
 

H. tholiforme Zhang, Ju & Xu

 
 
 

H. pseudobscurum Viane

 
 

H. obtusidentatum Chang & Zhang

 
 
 
 
 
 
 

H. cheilosorum

 
 
 
 

H. amoenum (Presl ex Mettenius) Ranil

 
 
 

H. filipes (Copeland) Sugim.

 
 

H. obliquissimum (Hayata) Sugim.

 
 
 
 
 

H. szechuanense (Ching) Viane & Dong

 
 
 

H. furfuraceum (Ching) Viane & Dong

 
 
 

H. latidens (Ching) Viane & Dong

 
 

H. quercicola (Ching) Viane & Dong

 
 
 
 
 
 
 
 
 
 

H. subnormale (Copeland) Nakaike

 
 

H. wildii (Bail.) Ohlsen

 
 
 
 
 

H. distans Zhang, Xu & Zhang

 
 
 

H. apogamum (N.Murak. & Hatan.) Nakaike

 
 
 

H. ngheanense Zhang, Xu & Lu

 
 
 

H. laterepens N.Murak. & Cheng ex Chang & Hori

 
 

H. unilaterale (Lam.) Hayata

 
 
 
 
 
 
 
 

H. cardiophyllum (Hance) Nakaike

 
 
 

H. quangnamense Zhang, Xu & Zhang

 
 
 

H. hastifolium Xu, Zhang & Liao

 
 

H. pubirhizoma (Ching & Liu) Xu & Zhang

 
 
 
 
 
 

H. chingii Xu, Zhang & Liao

 
 
 

H. wangpeishanii Zhang & Xu

 
 
 

H. hondoense (N.Murak. & Hatan.) Nakaike

 
 

H. speluncicola Zhang, Xu & He

 
 
 
 
 
 
 
 
 

References

  1. ↑ Alan R. Smith; Kathleen M. Pryer; Eric Schuettpelz; Petra Korall; Harald Schneider; Paul G. Wolf (2006). "A classification for extant ferns". Taxon 55 (3): 705–731. doi:10.2307/25065646. Bibcode: 2006Taxon..55..705S. http://www.pryerlab.net/publication/fichier749.pdf. 
  2. ↑ Maarten J. M. Christenhusz; Xian-Chun Zhang; Harald Schneider (2011). "A linear sequence of extant families and genera of lycophytes and ferns". Phytotaxa 19: 7–54. doi:10.11646/phytotaxa.19.1.2. Bibcode: 2011Phytx..19....7C. http://www.mapress.com/phytotaxa/content/2011/f/pt00019p054.pdf. 
  3. ↑ "Phylogeny of the fern family Aspleniaceae in Australasia and the south-western Pacific". Australian Systematic Botany 27 (6): 355–71. 2015. doi:10.1071/sb14043. 
  4. ↑ Template:Cite WF
  5. ↑ Nitta, Joel H.Expression error: Unrecognized word "et". (2022). "An Open and Continuously Updated Fern Tree of Life". Frontiers in Plant Science 13. doi:10.3389/fpls.2022.909768. PMID 36092417. Bibcode: 2022FrPS...1309768N. 
  6. ↑ "Tree viewer: interactive visualization of FTOL". 2024. https://fernphy.github.io/viewer.html. 

Template:Fern classification Wikidata ☰ Q5956439 entry



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