Biology:List of mosasaur genera

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Tylosaurus proriger mounted skeleton in the Rocky Mountain Dinosaur Resource Center in Woodland Park, Colorado

This list of mosasaurs is a comprehensive listing of all genera that have ever been included in the family Mosasauridae or the parent clade Mosasauroidea, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful (nomen dubium), or were not formally published (nomen nudum), as well as junior synonyms of more established names, and genera that are no longer considered mosasauroid. Non-mosasaurid mosasauroids shall be noted as such. The list currently includes 92 genera, out of which 58 are considered valid (47 mosasaurids and 10 other mosasauroids).

Scope and terminology

There is no official, canonical list of mosasaur genera but one of the most thorough attempts can be found on the "Pythonomorpha" section of Mikko Haaramo's Phylogeny Archive.

Naming conventions and terminology follow the International Code of Zoological Nomenclature. Technical terms used include:

  • Junior synonym: A name which describes the same taxon as a previously published name. If two or more genera are formally designated and the type specimens are later assigned to the same genus, the first to be published (in chronological order) is the senior synonym, and all other instances are junior synonyms. Senior synonyms are generally used, except by special decision of the ICZN, but junior synonyms cannot be used again, even if deprecated. Junior synonymy is often subjective, unless the genera described were both based on the same type specimen.
  • Nomen nudum (Latin for "naked name"): A name that has appeared in print but has not yet been formally published by the standards of the ICZN. Nomina nuda (the plural form) are invalid, and are therefore not italicized as a proper generic name would be. If the name is later formally published, that name is no longer a nomen nudum and will be italicized on this list. Often, the formally published name will differ from any nomina nuda that describe the same specimen. In this case, these nomina nuda will be deleted from this list in favor of the published name.
  • Preoccupied name: A name that is formally published, but which has already been used for another taxon. This second use is invalid (as are all subsequent uses) and the name must be replaced. As preoccupied names are not valid generic names, they will also go unitalicized on this list.
  • Nomen dubium (Latin for "dubious name"): A name describing a fossil with no unique diagnostic features.

Mosasaur genera

Genus Author(s) Year Status Age Location Notes Images
Acteosaurus Meyer 1860 Valid Late Cretaceous  Slovenia Its vertebral column resembles that of extant varanid lizards
Adriosaurus Seeley 1881 Valid Late Cretaceous  Croatia
 Slovenia
The A. skrbinensis specimen preserves a phosphatic matter in its stomach area, likely remains of fish, which suggests it may have been piscivorous 200px
Aigialosaurus Kramberger 1892 Valid Late Cretaceous  Croatia Has been suggested to be the oldest known member of the lineage that led to the larger mosasaurids 200px
Amphekepubis Mehl 1930 Dubious Late Cretaceous  Mexico Possibly a junior synonym of Mosasaurus
Amphorosteus Gibbs 1851 Dubious Late Cretaceous  United States Only known from two, heavily weathered vertebrae
Ancylocentrum Schmidt 1927 Jr. synonym N/A N/A Junior synonym of Prognathodon
Angolasaurus Antunes 1964 Valid Late Cretaceous  Angola
 Brazil?
 Niger?
 United States?
Mostly similar to the related Platecarpus, but with a somewhat longer skull 200px
Aphanizocnemus Dal Sasso & Pinna 1997 Valid Late Cretaceous  Lebanon Although commonly regarded as a dolichosaurid, a few analyses suggest it may be outside of that group 200px
Baptosaurus Marsh 1870 Jr. synonym N/A N/A Junior synonym of Halisaurus
Baseodon Leidy 1865 Jr. synonym N/A N/A Junior synonym of Mosasaurus
Batrachiosaurus Harlan 1839 Jr. synonym N/A N/A Junior synonym of Mosasaurus
Bentiabasaurus Polcyn, Schulp & Gonçalves 2023 Valid Late Cretaceous  Angola Discovered as a stomach content of an adult Prognathodon 200px
Brachysaurana Strand 1926 Jr. synonym N/A N/A Junior synonym of Prognathodon
Brachysaurus Williston 1897 Preoccupied N/A N/A Preoccupied by a junior synonym of iguanian lizard genus Stenocercus;[1] referred to replacement names Brachysaurana and Ancylocentrum, of which the latter has priority as such, although both names are synonyms of Prognathodon
Carinodens Thurmond 1969 Valid Late Cretaceous  Belgium
 Denmark
 Jordan
 Morocco
 Netherlands
 Russia
 Ukraine
Closely related to Globidens, but can be distinguished by its compressed teeth 200px
Carsosaurus Kornhuber 1893 Valid Late Cretaceous  Slovenia Preserves skin impressions and sternal cartilage 200px
Clidastes Cope 1868 Valid Late Cretaceous  United States One of the earliest known hydropedal mosasaurs 200px
Compressidens Dollo 1924 Preoccupied N/A N/A Preoccupied by a tusk shell; later renamed Carinodens
Coniasaurus Owen 1850 Valid Late Cretaceous  Germany
 United Kingdom
 United States
Only known from incomplete remains, but they are enough to tell that it had an elongated skull containing specialized dentition 200px
Dallasaurus Polcyn & Bell 2005 Valid Late Cretaceous  United States Has been said to be a "missing link" uniting fully aquatic mosasaur taxa and their terrestrial ancestors 200px
Dolichosaurus Owen 1894 Valid Late Cretaceous  United Kingdom Possessed an exceptionally long neck 200px
Dollosaurus Yakovlev 1905 Jr. synonym N/A N/A Junior synonym of Prognathodon
Drepanodon Leidy 1856 Jr. synonym N/A N/A Junior synonym of Mosasaurus
Ectenosaurus Russell 1967 Valid Late Cretaceous  United States One of the few plioplatecarpines that was not exclusive to nearshore marine environments 200px
Edestosaurus Marsh 1871 Jr. synonym N/A N/A Junior synonym of Clidastes
Eidolosaurus Nopcsa 1923 Valid Late Cretaceous  Slovenia One of the oldest known mosasauroids
Elliptonodon Emmons 1858 Jr. synonym N/A N/A Junior synonym of Tylosaurus
Eonatator Bardet et al. 2005 Valid Late Cretaceous  Colombia
 Sweden
 United States
The type species was originally referred to Clidastes and Halisaurus 200px
Eremiasaurus LeBlanc et al. 2012 Valid Late Cretaceous  Brazil
 Israel
 Morocco
Characterised by its robust skull with pronounced heterodont dentition 200px
Gavialimimus Strong et al. 2020 Valid Late Cretaceous  Angola?
 Morocco
Unusually, its snout was elongated, convergent with the extant gharial, hence its genus name 200px
Globidens Gilmore 1912 Valid Late Cretaceous  Angola
 Brazil
 Colombia
 Jordan
 Morocco
 Syria
 United States
Known for its rounded teeth, an adaptation to crush shelled prey such as molluscs 200px
Gnathomortis Lively 2020 Valid Late Cretaceous  United States Previously identified as a species of Prognathodon, but it has been found to be different enough from that taxon to receive its own genus
Goronyosaurus Azzaroli et al. 1972 Valid Late Cretaceous  Niger
 Nigeria
Possessed straight teeth with rounded apices more well suited for smashing food 200px
Haasiasaurus Polcyn et al. 2003 Valid Late Cretaceous  Palestine Possibly chimeric as all referred remains were not found in association[2]
Hainosaurus Dollo 1885 Jr. synonym N/A N/A Junior synonym of Tylosaurus
Halisaurus Marsh 1869 Valid Late Cretaceous  Egypt
 Jordan?
 Morocco
 Peru?
 United States
May have been a poor swimmer due to the lack of hyperphalangy as seen in more derived genera 200px
Harranasaurus Kaddumi 2009 Valid Late Cretaceous  Jordan Only known from a single mandible 200px
Holcodus Gibbs 1851 Jr. synonym N/A N/A Junior synonym of

Platecarpus

Holosaurus Marsh 1880 Preoccupied N/A N/A Referred to Platecarpus
Hydrosaurus Kornhuber 1873 Preoccupied N/A N/A Preoccupied by agamid lizard genus Hydrosaurus; renamed Pontosaurus
Igdamanosaurus Lingham-Soliar 1991 Valid Late Cretaceous  Egypt
 Niger
Its dentary was massively built, similar to Prognathodon and its relative Globidens 200px
Jormungandr Zietlow, Boyd & van Vranken 2023 Valid Late Cretaceous  United States Shares features of its skeleton with both basal and derived mosasaurines 200px
Kaganaias Evans et al. 2006 Valid Early Cretaceous  Japan The oldest and basalmost known dolichosaurid 200px
Kaikaifilu Otero et al. 2017 Valid Late Cretaceous Antarctica Potentially a tylosaurine, although some researchers consider this assignment problematic 200px
Khinjaria[3] Longrich et al. 2024 Valid Late Cretaceous  Morocco May have probably hunted large prey items due to its large body size and blade-like teeth. Closely related to Goronyosaurus
Kolposaurus Camp 1942 Preoccupied N/A N/A Preoccupied by a junior synonym of the unrelated Nothosaurus;[4] referred to Plotosaurus
Komensaurus Caldwell & Palci 2007 Valid Late Cretaceous  Slovenia Before its formal description, it had been nicknamed the "Trieste aigialosaur"
Kourisodon Nicholls & Meckert 2002 Valid Late Cretaceous  Canada
 Japan
Coexisted with several elasmosaurids, turtles, and other mosasaurs 200px
Lakumasaurus Novas et al. 2002 Jr. synonym N/A N/A Junior synonym of Taniwhasaurus
Latoplatecarpus Konishi & Caldwell 2011 Valid Late Cretaceous  Canada
 Russia
 United States
One of the largest named plioplatecarpines 200px
Leiodon Owen 1841 Preoccupied N/A N/A Preoccupied by a fish; referred to Liodon
Lesticodus Leidy 1859 Jr. synonym N/A N/A Junior synonym of Mosasaurus
Lestosaurus Marsh 1872 Jr. synonym N/A N/A Junior synonym of Platecarpus
Liodon Agassiz 1846 Dubious Late Cretaceous  United Kingdom Several species have been referred to this genus, although they do not belong to a single taxon 200px
Macrosaurus Owen 1849 Jr. synonym N/A N/A Junior synonym of Mosasaurus
Megapterygius[5] Konishi et al. 2023 Valid Late Cretaceous  Japan The neural spines of its posterior dorsal vertebrae have an abrupt change in orientation, which would have supported a dolphin-like dorsal fin in life 200px
Mesoleptos Cornalia & Chiozza 1852 Valid Late Cretaceous  Croatia
 Palestine
 Slovenia
An early member of the Mosasauroidea 200px
Moanasaurus Wiffen 1980 Valid Late Cretaceous  New Zealand One of the largest known mosasaurines 200px
Mosasaurus Conybeare 1822 Valid Late Cretaceous Antarctica
 Argentina
 Belgium
 Brazil
 Canada
 Jordan
 Morocco
 Netherlands
 Russia
 South Africa
 Turkey
 United States
The first mosasaur genus ever named and described 200px
Nectoportheus Cope 1868 Jr. synonym N/A N/A Junior synonym of Mosasaurus
Oneirosaurus[6] Páramo-Fonseca et al. 2025 Valid Late Cretaceous  Colombia Known from a single, well-preserved skull 200px
Opetiosaurus Kornhuber 1901 Valid Late Cretaceous  Croatia Has been suggested to be synonymous with Aigialosaurus,[7] but this was not supported by subsequent research[8] 200px
Oterognathus Dollo 1889 Jr. synonym N/A N/A Junior synonym of Plioplatecarpus
Pannoniasaurus Makádi, Caldwell & Ősi 2012 Valid Late Cretaceous  Hungary Lived in a freshwater habitat unlike other members of its family, which were marine predators 200px
Phosphorosaurus Dollo 1889 Valid Late Cretaceous  Belgium
 Japan
Had large eye sockets, which imply it would have hunted its prey in deep water or at night 200px
Platecarpus Cope 1869 Valid Late Cretaceous  Australia?
 Belgium?
 United States
One specimen is so well-preserved it shows that mosasaurs were powerful, agile swimmers 200px
Plesioplatecarpus Konishi & Caldwell 2011 Valid Late Cretaceous  United States Six specimens are known to date 200px
Plesiotylosaurus Camp 1942 Valid Late Cretaceous  United States Some traits of its skull are seemingly convergent with tylosaurine mosasaurs 200px
Plioplatecarpus Dollo 1882 Valid Late Cretaceous  Canada
 Netherlands
 Sweden
 United States
Lived in a broad range as suggested by its fossil record 200px
Plotosaurus Camp 1951 Valid Late Cretaceous  United States Unusually, its overall morphology was more similar to that of ichthyosaurs than to other mosasaurs, which led to its quite derived position within the latter group 200px
Pluridens Lingham-Soliar 1998 Valid Late Cretaceous  Morocco
 Niger
 Nigeria
Some specimens preserve injuries to their jaws, which would indicate they may have engaged in intraspecific combat 200px
Pontosaurus Gorjanovic-Kramberger 1892 Valid Late Cretaceous  Croatia
 Lebanon
Two species are known 200px
Portunatasaurus Mekarski et al. 2019 Valid Late Cretaceous  Croatia Has been used as a subject for the understanding on the evolution of mosasauroid limb morphology
Primitivus Paparella et al. 2018 Valid Late Cretaceous  Italy The first dolichosaurid named from Italy 200px
Proaigialosaurus Kuhn 1958 Valid Late Jurassic  Germany Often regarded as an aigialosaurid, but it may have also been a pleurosaurid
Prognathodon Dollo 1889 Valid Late Cretaceous  Angola?
 Belgium
 Canada?
 Israel
 Jordan
 Mexico
 Netherlands?
 New Zealand?
 Spain
 Syria
 Ukraine
 United States?
Possibly paraphyletic as most assigned species may not belong to this genus 200px
Prognathosaurus Williston 1897 Jr. synonym N/A N/A Junior synonym of Prognathodon
Pterycollosaurus Dollo 1882 Jr. synonym N/A N/A Junior synonym of Mosasaurus
Rhamphosaurus Cope 1872 Preoccupied N/A N/A Preoccupied by another lizard genus; referred to Tylosaurus
Rhinosaurus Marsh 1872 Preoccupied N/A N/A Preoccupied;[9] referred to Tylosaurus
Rikisaurus Wiffen 1990 Jr. synonym N/A N/A Junior synonym of Moanasaurus
Rikkisaurus Bell et al. 1999 Lapsus calami N/A N/A Lapsus calami of Rikisaurus, a junior synonym of Moanasaurus
Romeosaurus Palci et al. 2013 Valid Late Cretaceous  Italy None of the described specimens has enough well-preserved postcranial material, making it somewhat difficult to make any good judgements of the genus' full anatomy 200px
Russellosaurus Polcyn & Bell 2005 Valid Late Cretaceous  United States One of the oldest mosasaurs described from North America 200px
Sarabosaurus Polcyn et al. 2023 Valid Late Cretaceous  United States Based on remains of a mature animal as indicated by its preserved growth rings
Saurochampsa Wagler 1830 Jr. synonym N/A N/A Junior synonym of Mosasaurus
Selmasaurus Wright & Shannon 1988 Valid Late Cretaceous  United States Would have been unable to widen its jaws to attack large prey due to its uniquely akinetic skull 200px
Sironectes Cope 1840 Jr. synonym N/A N/A Junior synonym of Platecarpus
Stelladens Longrich et al. 2023 Valid Late Cretaceous  Morocco Named for its star-shaped teeth 200px
Taniwhasaurus Hector 1874 Valid Late Cretaceous Antarctica
 Japan?
 New Zealand
 South Africa?
May have possessed an electro-sensitive organ in its snout foramina that was likely capable of detecting movements of prey underwater 200px
Tetrapodophis Martill et al. 2015 Valid Early Cretaceous  Brazil Originally described as a basal snake but later often reinterpreted as a dolichosaurid 200px
Tethysaurus Bardet, Pereda-Suberbiola & Jalil 2003 Valid Late Cretaceous  Morocco Exhibits a mixture of primitive and advanced features 200px
Thalassotitan Longrich et al. 2022 Valid Late Cretaceous  Morocco Its discovery shows that mosasaurs evolved to be the apex predators in the Late Cretaceous oceans. Closely related to Prognathodon 200px
Tylosaurus Marsh 1872 Valid Late Cretaceous  Angola
 Belgium
 Canada
 Democratic Republic of the Congo?
 France
 Morocco
 Sweden
 United States
Some species are among the largest mosasaurs yet known 200px
Vallecillosaurus Smith & Buchy 2008 Valid Late Cretaceous  Mexico One of the oldest mosasauroids ever described 200px
Xenodens Longrich et al. 2021 Valid Late Cretaceous  Morocco One study considered this taxon a nomen dubium due to the lack of authenticity,[10] but this was refuted by the discovery of additional material and CT scans[11] 200px
Yaguarasaurus Páramo 1994 Valid Late Cretaceous  Colombia
 Mexico
The most completely known South American mosasaur during the time of its description 200px

See also

Notes

  1. Stenocercus at Herpbreeder.com . Retrieved 12-03-2007.
  2. Alessandro Palci, Michael W. Caldwell and Cesare A. Papazzoni (2013). "A new genus and subfamily of mosasaurs from the Upper Cretaceous of northern Italy". Journal of Vertebrate Paleontology 33 (3): 599–612. doi:10.1080/02724634.2013.731024. 
  3. Longrich, Nicholas R.; Polcyn, Michael J.; Jalil, Nour-Eddine; Pereda-Suberbiola, Xabier; Bardet, Nathalie (2024-03-01). "A bizarre new plioplatecarpine mosasaurid from the Maastrichtian of Morocco". Cretaceous Research 160. doi:10.1016/j.cretres.2024.105870. ISSN 0195-6671. Bibcode2024CrRes.16005870L. https://www.sciencedirect.com/science/article/pii/S0195667124000430. 
  4. Sauropterygia at Paleofile.com. Retrieved 12-03-2007.
  5. Konishi, Takuya; Ohara, Masaaki; Misaki, Akihiro; Matsuoka, Hiroshige; Street, Hallie P.; Caldwell, Michael W. (2023). "A new derived mosasaurine (Squamata: Mosasaurinae) from south-western Japan reveals unexpected postcranial diversity among hydropedal mosasaurs" (in en). Journal of Systematic Palaeontology 21 (1). doi:10.1080/14772019.2023.2277921. ISSN 1477-2019. 
  6. Páramo-Fonseca, María Eurídice; Narváez-Rincón, José Alejandro; Benavides-Cabra, Cristian David (2025-10-29). "Oneirosaurus caballeroi gen. et sp. nov., a new mosasaur from Colombia" (in en). Earth Sciences Research Journal 29 (3): 231–247. doi:10.15446/esrj.v29n3.117157. ISSN 2339-3459. https://revistas.unal.edu.co/index.php/esrj/article/view/117157. 
  7. Dutchak, Alex R.; and Caldwell, Michael W. (2009)."A redescription of Aigialosaurus (= Opetiosaurus) bucchichi (Kornhuber, 1901) (Squamata: Aigialosauridae) with comments on mosasauroid systematics". Journal of Vertebrate Paleontology 29 (2): 437-452.
  8. Madzia, D., Cau, A. (2017). Inferring "weak spots" in phylogenetic trees: application to mosasauroid nomenclature . PeerJ 5:e3782
  9. Edward Drinker Cope "On the structure of the Pythonomorpha". (1872) Proceedings of the Academy of Natural Sciences of Philadelphia, 24:140-141
  10. Sharpe, Henry S.; Powers, Mark J.; Caldwell, Michael W. (2024-12-16). "Reassessment of Xenodens calminechari with a discussion of tooth morphology in mosasaurs" (in en). The Anatomical Record. doi:10.1002/ar.25612. ISSN 1932-8486. PMID 39682068. 
  11. Longrich, N. R.; Bardet, N.; Jalil, N.-E.; Pereda-Suberbiola, X.; Schulp, A.; Ghamizi, M. (2025). "New Information on the Morphology and Tooth Replacement of Xenodens calminechari (Squamata: Mosasauridae), a Unique Mosasaurid from the Maastrichtian Phosphates of Morocco". Diversity 17 (12). doi:10.3390/d17120819. https://www.mdpi.com/1424-2818/17/12/819. 

References

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