Biology:Rhinosteus

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Short description: Genus of fishes (fossil)

Rhinosteus
Temporal range: Late Devonian: Frasnian, 382.7–372.2 Ma
R. tuberculatus, R. traquairi and R. parvulus
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Class: †Placodermi
Order: †Arthrodira
Suborder: †Brachythoraci
Family: †Selenosteidae
Genus: †Rhinosteus
Jaekel, 1911
Species
  • Rhinosteus traquairi Jaekel, 1911 (type)
  • Rhinosteus parvulus (Gross, 1932)
  • Rhinosteus tuberculatus Gross, 1932
Synonyms
  • Platyrhinosteus sp. Jaekel, 1929

Rhinosteus is an extinct genus of small to medium selenosteid arthrodire placoderms of the Late Devonian known from the Upper Frasnian Kellwasserkalk facies of Bad Wildungen, Germany [1] and Morocco.[2]

Description

Rhinosteus is a typical genus of Kellwasserkalk selenosteids, with short cheeks, and slender inferognathal (lower jaw) plates. However, two of the species, R. traquairi and R. tuberculatus, have long, pointed rostrums, and tubercles on the plates. In R. traquairi, the rostrum is sharply pointed, extending beyond the snout, and the tubercles are small and irregularly placed. In R. tuberculatus, the rostrum is bluntly pointed, and the tubercles are large and plentiful. The species R. parvulus has a blunt rostrum that does not extend beyond the snout, and lacks tuberculation altogether.[1] The average skull length of R. traquairi is about 11 centimetres. The average skull length of R. tuberculatus is 15 centimetres, while that of R. parvulus is 4 to 6 centimetres.

Distribution

Fossils of R. traquairi and R. tuberculatus are found only in Bad Wildungen, Germany . R. parvulus is also found in Bad Wildungen. Rücklin (2011) discusses how specimens of small Rhinosteus have been found in the Kellwasserkalk facies of the Anti-Atlas Mountains of Morocco, and are tentatively referred to R. parvulus due primarily to their small size.[2]

Phylogeny

Rhinosteus is a member of the family Selenosteidae of the clade Aspinothoracidi, which belongs to the clade Pachyosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny of Rhinosteus:[3]

Eubrachythoraci 
 

Coccosteomorphi

 
 Pachyosteomorphi 
 

Rhachiosteus pterygiatus

 
 
Dunkleosteoidea 
 

Eastmanosteus calliaspis

 
 
 
 

Eastmanosteus pustulosus

 
 

Kiangyousteus yohii

 
 
 
 

Golshanichthys asiatica

 
 
 

Westralichthys uwagedensis

 
 
 
 

Dunkleosteus raveri

 
 

Dunkleosteus terrelli

 
 
 
 

Dunkleosteus amblyodoratus

 
 Heterostiidae 
 

Heterosteus ingens

 
 

Yinosteus major

 
 
 
 
 
 
 
 
Aspinothoracidi
 

Tapinosteus heintzi

 
 
 
 

Bullerichthys fascidens

 
 
 

Kendrickichthys cavernosus

 
 

Bruntonichthys multidens

 
 
 
 
 

Dinichthys herzeri

 
 
 

Hadrosteus rapax

 
 
 

Gorgonichthys clarki

 
 Selenosteidae 
 

Heintzichthys gouldii

 
 

Pachyosteus bulla

 
 

Gymnotrachelus hydei

 
 

Stenosteus angustopectus

 
 

Brachyosteus dietrichi

 
 
 

Melanosteus occitanus

 
 

Rhinosteus parvulus

 
 
 
 
 
 
 
 
 
 
 

References

  1. ↑ 1.0 1.1 Denison, Robert (1978). Handbook of Paleoichthyology, Volume 2, Placodermi. New York: Gustav Fischer Verlage. pp. 97. ISBN 9780895740274. 
  2. ↑ 2.0 2.1 RÜCKLIN, MARTIN (January 14, 2011). "First selenosteid placoderms from the eastern Anti-Atlas of Morocco; osteology, phylogeny and palaeogeographical implications". Palaeontology 56 (1): 25–62. doi:10.1111/j.1475-4983.2010.01026.x. 
  3. ↑ Zhu, You-An; Zhu, Min; Wang, Jun-Qing (1 April 2016). "Redescription of Yinostius major (Arthrodira: Heterostiidae) from the Lower Devonian of China, and the interrelationships of Brachythoraci". Zoological Journal of the Linnean Society 176 (4): 806–834. doi:10.1111/zoj.12356. ISSN 0024-4082. 

Wikidata ☰ Q7320683 entry