Biology:Selenosteidae

From HandWiki
Short description: Extinct family of fishes

Selenosteidae
Temporal range: Late Devonian, 382.7–358.9 Ma
Driscollichthys, Amazichthys, Draconichthys
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Class: †Placodermi
Order: †Arthrodira
Suborder: †Brachythoraci
Clade: †Eubrachythoraci
Clade: †Pachyosteomorphi
Clade: †Aspinothoracidi
Family: †Selenosteidae
Dean, 1901
Type species
Selenosteus brevis
(Claypole), 1869
Genera
Synonyms
  • Pachyosteidae Gross, 1932
  • Braunosteidae Stensiö, 1959
  • Rhinosteidae Stensiö, 1963
Life restoration of Amazichthys

Selenosteidae is an extinct family of small to large-sized arthrodire placoderms from the Late Devonian. With the exception of the Chinese Phymosteus, selenosteids lived in shallow seas in what is now Eastern North America (the Cleveland Shale), Eastern Europe (Holy Cross Mountains, Poland , and the Kellwasserkalk fauna of Bad Wildungen), and Northeastern Africa (the Anti-Atlas Mountains, Morocco).

Selenosteids have, in cross section, a rounded body, a blunt snout, and tremendous orbits. The lower jaws were slender, the inferognathals usually either being finely serrated, or adapted for crushing,[1] though, in Draconichthys, the inferognathals had long prongs for seizing prey.[2] The rostrum is very small.

Taxonomy

Selenosteidae is a member of the clade Aspinothoracidi, which belongs to the clade Pachyosteomorphi, one of the two major clades within Eubrachythoraci. Gorgonichthys is closely related to the family Selenosteidae, and could possibly be included in the family. The cladogram below shows the placement of Selenosteidae within Eubrachythoraci:[3]

Eubrachythoraci 
 

Coccosteomorphi

 
 Pachyosteomorphi 
 

Rhachiosteus pterygiatus

 
 
Dunkleosteoidea 
 

Eastmanosteus calliaspis

 
 
 
 

Eastmanosteus pustulosus

 
 

Kiangyousteus yohii

 
 
 
 

Golshanichthys asiatica

 
 
 

Westralichthys uwagedensis

 
 
 
 

Dunkleosteus raveri

 
 

Dunkleosteus terrelli

 
 
 
 

Dunkleosteus amblyodoratus

 
 Heterostiidae 
 

Heterosteus ingens

 
 

Yinosteus major

 
 
 
 
 
 
 
 
Aspinothoracidi
 

Tapinosteus heintzi

 
 
 
 

Bullerichthys fascidens

 
 
 

Kendrickichthys cavernosus

 
 

Bruntonichthys multidens

 
 
 
 
 

Dinichthys herzeri

 
 
 

Hadrosteus rapax

 
 
 

Gorgonichthys clarki

 
 Selenosteidae 
 

Heintzichthys gouldii

 
 

Pachyosteus bulla

 
 

Gymnotrachelus hydei

 
 

Stenosteus angustopectus

 
 

Brachyosteus dietrichi

 
 
 

Melanosteus occitanus

 
 

Rhinosteus parvulus

 
 
 
 
 
 
 
 
 
 
 

Below is a more detailed cladogram of Selenosteidae, from Rücklin (2011):[2]

Aspinothoracidi
 

Parabelosteus

 
 

Brachydeirus

 
Selenosteidae
 

Pachyosteus

 
 
 
 

Selenosteus

 
 
 

Stenosteus

 
 

Gymnotrachelus

 
 
 
 
 

Microsteus

 
 
 

Walterosteus

 
 
 

Draconichthys

 
 

Enseosteus

 
 
 

Melanosteus

 
 

Rhinosteus

 
 
 
 
 
 
 
 

Denison (1978)[1] separated Selenosteidae into two informal groups, the "American" genera (i.e., those taxa from the Upper Famennian Cleveland Shales, such as Selenosteus and Gymnotrachelus), and the "European" genera (i.e., those taxa from the Upper Frasnian Kellwasserkalk facies of Germany, and later, Morocco, like Rhinosteus and Microsteus). Denison notes that the differences between the two groups, such as how the American genera tend to have slightly smaller orbits, and slightly longer cheeks, may suggest the similarities between them are due to parallel evolution.[1]

Rücklin (2011), on the other hand, places Pachyosteus as the most basal selenosteid, then groups the American genera (with Selenosteus grouped as the sister taxon of Gymnotrachelus and Stenosteus) as being the sister group of the remaining European/Kellwasserkalk genera, plus Draconichthys.[2]

See also

  • List of placoderms

References

  1. ↑ 1.0 1.1 1.2 Denison, Robert (1978). Handbook of Paleoichthyology, Volume 2, Placodermi. New York: Gustav Fischer Verlage. pp. 94. ISBN 9780895740274. 
  2. ↑ 2.0 2.1 2.2 RÜCKLIN, MARTIN (January 14, 2011). "First selenosteid placoderms from the eastern Anti-Atlas of Morocco; osteology, phylogeny and palaeogeographical implications". Palaeontology 56 (1): 25–62. doi:10.1111/j.1475-4983.2010.01026.x. 
  3. ↑ Zhu, You-An; Zhu, Min; Wang, Jun-Qing (1 April 2016). "Redescription of Yinostius major (Arthrodira: Heterostiidae) from the Lower Devonian of China, and the interrelationships of Brachythoraci". Zoological Journal of the Linnean Society 176 (4): 806–834. doi:10.1111/zoj.12356. ISSN 0024-4082. 

Wikidata ☰ Q7447910 entry