Earth:Nacimiento Formation

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Nacimiento Formation
Stratigraphic range: Paleocene
~65.7–61 Ma
The badlands below Angel Peak are made up of Nacimiento Formation rocks
TypeFormation
UnderliesSan Jose Formation
OverliesOjo Alamo Formation
Thickness254 m (833 ft)
Lithology
PrimaryShale
OtherSiltstone, sandstone
Location
Coordinates [ ⚑ ] 36°00′46″N 106°59′08″W / 36.0126977°N 106.9855481°W / 36.0126977; -106.9855481
RegionNew Mexico
CountryUnited States
Type section
Named forNacimiento (now Cuba, New Mexico)
Named byJ.H. Gardner
Year defined1910
Nacimiento Formation is located in the United States
Nacimiento Formation
Nacimiento Formation (the United States)
Nacimiento Formation is located in New Mexico
Nacimiento Formation
Nacimiento Formation (New Mexico)

The Nacimiento Formation is a sedimentary rock formation in the San Juan Basin of western New Mexico (United States).[1] It is dated to approximately 61.0–65.7 million years ago, corresponding to the early and middle Paleocene. The formation has yielded numerous fossil assemblages from the interval following the Cretaceous–Paleogene extinction event, which have been used to study biotic recovery and the early diversification of mammals after the extinction.[1]

Description

The Nacimiento Formation is a heterogeneous nonmarine formation composed of shale, siltstone, and sandstone,[2] deposited in floodplain, fluvial and lacustrine settings,[3] and made up of sediment shed from the San Juan uplift to the north and the Brazos-Sangre de Cristo uplift to the east.[4] It was deposited mostly between ~65.7 and ~61 million years ago, during the early and middle Paleocene.[5] The climate was humid and warm to hot[6] and stable, but with a distinct dry season.[7] This unit interbeds with the underlying Ojo Alamo Formation but is separated by an unconformity from the overlying San Jose Formation.[3]

The Nacimiento Formation is divided into several subunits known as members. In outcrops in southern areas of the formation, the Puercan fauna is found in the Arroyo Chijuillita Member, the Torrejonian fauna is found in the Ojo Encino Member, and the uppermost Escavada Member lacks age-diagnostic fossils.[1] In northern outcrops, the two lower members are indistinguishable, and are called the "main body".[5] Above them are two more informal members. These preserve a younger, Tiffanian fauna.[8] The Puercan and Torrejonian faunas are further subdivided into several biostratigraphic zones.[5]

Fossils

Many fossils are known from the Nacimiento Formation, although bone is often altered into phosphatic concretions.[9] Fossils belonging to a number of different organisms have been found here, including: plants (mostly dicotyledonous angiosperms),[6][10] gastropods, freshwater bivalves,[11] cartilaginous fish and bony fish, salamanders, turtles, champsosaurs, amphisbaenians, lizards, snakes, crocodilians,[12] birds,[13] and a variety of archaic mammals. Mammalian groups represented include multituberculates,[14] plesiadapiforms,[15] didelphid marsupials, insectivorans, carnivorans, taeniodonts, mesonychians, condylarths, and cimolestans.[1] Fossil remains found in the formation support the validity of the genus Thylacodon and the species T. montanensis.[16]

These fossils provide important clues to the impact of the Cretaceous–Paleogene extinction event on mammals[16] and to the recovery, evolution, and turnover of mammals shortly after the event. The formation and its fossils provide a particularly clear record of the To2-To3 turnover event, allowing the timing of the event to be constrained to between 62.59 and 62.47 million years ago. The event may have been associated with climate change or with the rapid development of a river system across the San Juan basin, which caused a temporary pause in sediment deposition that separates the Nacimiento Formation from the San Jose Formation.[17]

Fossil content

Based on the Paleobiology Database:[18]

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Actinopterygii

Boney Fish of the Nacimiento Formation
Genus Species Location Stratigraphic position Material Notes Images
Lepisosteidae indet. Torrejonian A gar of uncertain affinities.

Amphibians

Amphibians of the Nacimiento Formation
Genus Species Location Stratigraphic position Material Notes Images
Habrosaurus H. dilatus Torrejonian A sirenid salamander.

Reptiles

Crocodyliformes

Crocodyliformes of the Nacimiento Formation
Genus Species Location Stratigraphic position Material Notes Images
Akanthosuchus A. langstoni Puercan & Torrejonian An alligatorid.
Listrognathosuchus L. multidentatus Torrejonian An alligatoroid.
Navajosuchus L. mooki Puercan & Torrejonian An alligatorid. frameless

Turtles

Turtles of the Nacimiento Formation
Genus Species Location Stratigraphic position Material Notes Images
Adocus A. hesperius Puercan An adocid turtle. frameless
A. onerosus Torrejonian
Axestemys A. montinsana Puercan[19] A softshell turtle. frameless
Compsemys C. victa Puercan/Torrejonian A compsemydid turtle.
"Emys" "E." sp. Puercan/Torrejonian A pond turtle.
Hutchemys H. acupictus Puercan[19] A plastomenid stem-softshell turtle.
Hoplochelys H. crassa Puercan/Torrejonian A kinosternoid turtle. Type locality of species.
Neurankylus N. cf. eximius Puercan A baenid turtle. Type locality of N. torrejonensis. frameless
N. torrejonensis Torrejonian, potentially Puercan
Plastomenus P. vegetus Puercan A plastomenid stem-softshell turtle.[20]
"Platypeltis" "P." antiqua Torrejonian A stem-softshell turtle. Nomen dubium.[19]
"Trionyx" (=Paleotrionyx) "T." eloisae Puercan A softshell turtle. Type locality of species.[19]
"T." reesidei
"T." singularis

Squamates

Squamates of the Nacimiento Formation
Genus Species Location Stratigraphic position Material Notes Images
Glyptosaurinae indet. Puercan & Torrejonian A glyptosaurine anguid of uncertain affinities.
Helagras H. prisciformis Puercan A snake, the largest known Early Paleocene snake.[21]
Machaerosaurus M. torrejonensis Torrejonian A glyptosaurine anguid.
Odaxosaurus O. piger Puercan A glyptosaurine anguid.
Parodaxosaurus P. sanjuanensis Torrejonian A glyptosaurine anguid.
Plesiorhineura P. tsentasi Torrejonian A rhineurid worm lizard.
Proxestops P. silberlingii Puercan & Torrejonian A glyptosaurine anguid.

Birds

Birds of the Nacimiento Formation
Genus Species Location Stratigraphic position Material Notes Images
Tsidiiyazhi T. abini Torrejonian A relative of the mousebirds.[22]

Mammals

Primary based on Kondrashov and Lucas (2015), with changes where necessary:[23]

Archaic Ungulates

'Condylarths' of the Nacimiento Formation
Genus Species Stratigraphic position Notes Images
Anisonchus A. sectorius Torrejonian 1-3 A periptychid
Carsioptychus [24] C. coarctatus Puercan 2-3 A periptychid originally described as a species of the genus Periptychus.
Conacodon C. cophater Puercan 2-3 A periptychid
C. entoconus Puercan 2
C. kohlbergeri Puercan 2
Ectoconus E. ditrigonus Puercan 2-3 A periptychid
Haploconus H. augustus Torrejonian 1-2 A periptychid
Hemithlaeus H. kowalewskianus Puercan 2 A periptychid
Mithrandir[25] M. gillianus Puercan 2-3 A periptychid originally described as a species of the genus Anisochus.
Oxyacodon O. agapetillus Puercan 2 A periptychid
O. apiculatus Puercan 2-3
O. priscilla Puercan 2
Periptychus P. carinidens Torrejonian 1-3 A periptychid

Arctocyonia

Arctocyonians of the Nacimiento Formation
Genus Species Stratigraphic position Notes Images
Arctocyon A. corrugatus Torrejonian 2 An arctocyonid
A. ferox Torrejonian 2-3
Chriacus C. baldwini Torrejonian 2-3 An arctocyonid
C. pelvidens Torrejonian 2-3
Colpoclaenus C. procyonoides Torrejonian 3 An arctocyonid
Desmatoclaenus D. dianae Puercan 2-3 An arctocyonid
D. protogonoides Puercan 2-3
Deuterogonodon D. montanus Torrejonian 1-2 An arctocyonid
Loxolophus L. hyattianus Puercan 2-3 An arctocyonid
L. pentacus Puercan 2-3
L. priscus Puercan 3
Prothryptacodon P. ambiguus Torrejonian 3 An arctocyonid
Tricentes T. subtrigonus Torrejonian 1-3 An arctocyonid

Carnivorans

Carnivorans of the Nacimiento Formation
Genus Species Stratigraphic position Notes Images
Bryanictis B. paulus Torrejonian 2 A viverravid carnivoran
Intyrictis I. vanvaleni Torrejonian 2 A viverravid carnivoran
Protictis P. haydenianus Torrejonian 2-3 A viverravid carnivoran
P. minor Torrejonian 2
P. simpsoni Torrejonian 2

Cimolesta

Cimolestans of the Nacimiento Formation
Genus Species Stratigraphic position Notes Images
Acemeodon A. secans Torrejonian 2-3 A cimolestid cimolestan
Betonnia B. tsosia Puercan 2-3 A cimolestid cimolestan
Chacopterygus C. minutus Puercan 2-3 A cimolestid cimolestan
Palaeoryctes P. puercensis Torrejonian 2-3 A palaeoryctid cimolestan
Puercolestes P. simpsoni Puercan 2-3 A cimolestid cimolestan

Dermoptera

Dermopterans of the Nacimiento Formation
Genus Species Stratigraphic position Notes Images
Mixodectes M. malaris Torrejonian 2-3 A mixodectid dermopteran
M. pungens Torrejonian 3

Eutheria Incertae sedis

Other Eutherians of the Nacimiento Formation
Genus Species Stratigraphic position Notes Images
Deltatherium[26] D. dandreae Torrejonian 2-3 A eutherian mammal of unknown placement
D. fundaminis Torrejonian 4-5

Leptictida

Leptictidans of the Nacimiento Formation
Genus Species Stratigraphic position Notes Images
Prodiacodon P. puercensis Torrejonian 1-3 A leptictiid leptictidan

Mesonychians

Mesonychians of the Nacimiento Formation
Genus Species Stratigraphic position Notes Images
Ankalagon A. saurognathus Torrejonian 2-3 A mesonychid mesonychian
Carcinodon C. antiquus Puercan 3 A triisodontid mesonychian
C. olearyi Puercan 2-3
C. simplex Puercan 2
Dissacus D. navajovius Torrejonian 2-3 A mesonychid mesonychian
Eoconodon E. coryphaeus Puercan 2-3 A triisodontid mesonychian
E. gaudrianus Puercan 2
E. ginibitohia Puercan 2
Goniacodon G. levisanus Torrejonian 2-3 A triisodontid mesonychian
Oxyclaenus O. cuspidatus Torrejonian 2-3 A triisodontid mesonychian
Triisodon T. crassicuspis Torrejonian 2 A triisodontid mesonychian
T. quivirensis Torrejonian 2

Metatheria

Metatherians of the Nacimiento Formation
Genus Species Stratigraphic position Notes Images
Peradectes P. coprexeches Torrejonian 2-3 A peradectid metatherian
P. minor Puercan 2-3
Swaindelphys S. encinensis Torrejonian 2 A herpetotheriid metatherian
S. johnsoni Torrejonian 3
Thylacodon T. pusillus Puercan 2-3 A herpetotheriid metatherian

Multituberculata

Multituberculates of the Nacimiento Formation
Genus Species Stratigraphic position Notes Images
Anconodon A. gidleyi Torrejonian 2 A neoplagiaulacid multituberculate
Catopsalis C. fissidens Torrejonian 2 A taeinolabid multituberculate
C. foliatus Puercan 2
Ectypodus E. szalayi Torrejonian 2 A neoplagiaulacid multituberculate
Eucosmodon E. americanus Puercan 2 An eucosmodontid multituberculate
E. molestus Torrejonian 2
E. primus Puercan 2
Kimbetohia K. campi Puercan 2 A ptilodontid multituberculate
Kimbetopsalis[27] K. simmonsae Puercan 2 A taeinolabid multituberculate
Krauseia K. clemensi Torrejonian 2 A neoplagiaulacid multituberculate
Liotomus L. vanvaleni Puercan 2 A cimolodontid multituberculate
Mesodma M. formosa Puercan 2 A neoplagiaulacid multituberculate
M. thompsoni Puercan 2
Mimetodon M. krausei Torrejonian 2 A neoplagiaulacid multituberculate
Neoplagiaulax N. macintyrei Puercan 2 A neoplagiaulacid multituberculate
N. macrotomeus Torrejonian 2
Parectypodus P. trovessartianus Torrejonian 2 A neoplagiaulacid multituberculate
Ptilodus P. mediaevus Torrejonian 2-3 A ptilodontid multituberculate
P. tsosiensis Puercan 2
Stygimys S. teilhardi Torrejonian 2 An eucosmodontid multituberculate
Taeniolabis T. simmonsae Puercan 3 A taeinolabid multituberculate
T. taoensis Puercan 3

Palaeanodonta

Palaeanodonta of the Nacimiento Formation
Genus Species Stratigraphic position Notes Images
Escavadodon E. zygus Torrejonian 3 An escavadodontid palaeanodont

Panperissodactyla

Panperissodactyls of the Nacimiento Formation
Genus Species Stratigraphic position Notes Images
Bomburodon B. priscus Puercan 2-3 A hyopsodontid
Choeroclaenus C. turgidunculus Puercan 2-3 A hyopsodontid
Chacomylus C. sladei Puercan 3 A hyopsodontid
Litomylus L. osceolae Torrejonian 2-3 A hyopsodontid
Mioclaenus M. turgidus Torrejonian 1-3 A hyopsodontid
Promioclaenus P. wilsoni Puercan 2-3 A hyopsodontid
Protoselene P. bombadili Puercan 2-3 A hyopsodontid
P. opisthacus Torrejonian 2-3
Tetraclaenodon T. puercensis Torrejonian 1-3 A phenacodontid
Tiznatzinia T. vanderhoofi Puercan 2-3 A hyopsodontid

Pantodonts

Pantodonts of the Nacimiento Formation
Genus Species Stratigraphic position Notes Images
Pantolambda P. bathmodon Torrejonian 3 A pantolambdid pantodont
P. cavirictum Torrejonian 2-3
P. intermedium Torrejonian 2

Pantolesta

Pantolestans of the Nacimiento Formation
Genus Species Stratigraphic position Notes Images
Coriphagus C. encinensis Torrejonian 2-3 A pentacodontid pantolestan
Pentacodon P. inversus Torrejonian 2 A pentacodontid pantolestan
P. occultus Torrejonian 3 A pentacodontid pantolestan

Plesiadapiforms

Plesiadapiforms of the Nacimiento Formation
Genus Species Stratigraphic position Notes Images
Anasazia A. williamsoni Torrejonian 2 A palaechthonid plesiadapiform
Palaechthon P. woodi Tiffanian 1 A palaechthonid plesiadapiform
Paromomys P. libedianus Torrejonian 2 A paromomyid plesiadapiform
Picrodus P. calgariensis Torrejonian 2 A picrodontid plesiadapiform
Plesiolestes P. nacimienti Torrejonian 2-3 A palaechthonid plesiadapiform
P. torrejonius Torrejonian 2
Torrejonia T. wilsoni Torrejonian 3 A palaechthonid plesiadapiform

Taeniodonta

Taeniodonts of the Nacimiento Formation
Genus Species Stratigraphic position Notes Images
Conorystes C. comma Torrejonian 2-3 A conoryctid taeniodont
Conoryctella C. pattersoni Torrejonian 2 A conoryctid taeniodont
Huerfanodon H. torrejonius Torrejonian 2 A conoryctid taeniodont
Onychodectes O. tisonensis Puercan 2-3 A conoryctid taeniodont
Psittacotherium P. multifragum Torrejonian 2-3 A stylinodontid taeniodont
Wortmania W. otariidens Puercan 2-3 A stylinodontid taeniodont

Molluscs

Bivalves

Bivalves of the Nacimiento Formation
Genus Species Location Stratigraphic position Material Notes Images

Gastropods

Gastopods of the Nacimiento Formation
Genus Species Location Stratigraphic position Material Notes Images

History of investigation

Edward Drinker Cope described the fossils during the Wheeler Survey.[28]

Workers in the early 1900s divided the rocks of the Nacimiento Formation into two formations, the lower Puerco Formation and the upper Torrejon Formation.[13] This was rejected on the grounds that there were no lithological differences between the two, only differences in fossil faunas, making determination of which formation was present in a given area impossible if fossils could not be found.[9] The Puerco and Torrejon were retained as zones within the Nacimiento Formation, and their faunas became the basis of the Puercan and Torrejonian North American Land Mammal Ages.[29]

Footnotes

  1. 1.0 1.1 1.2 1.3 Williamson & Lucas 1992.
  2. Schoch 1981.
  3. 3.0 3.1 Lucas & Ingersoll 1981.
  4. Lucas 1984.
  5. 5.0 5.1 5.2 Williamson 1996.
  6. 6.0 6.1 Tidwell, Ash & Parker 1981.
  7. Davis et al. 2016.
  8. Libed 2005.
  9. 9.0 9.1 Simpson 1959.
  10. Anderson 1960.
  11. Hartman 1981.
  12. Sullivan & Lucas 1986.
  13. 13.0 13.1 Gardner 1910.
  14. Williamson et al. 2016.
  15. Silcox & Williamson 2012.
  16. 16.0 16.1 Williamson et al. 2012.
  17. Leslie et al. 2018.
  18. "PBDB Strata Results". https://paleobiodb.org/classic/displayStrata?geological_group=&formation=Nacimiento&group_formation_member=Nacimiento. 
  19. 19.0 19.1 19.2 19.3 Vitek, Natasha S.; Joyce, Walter G. (2015). "A Review of the Fossil Record of New World Turtles of the Clade Pan-Trionychidae" (in en). Bulletin of the Peabody Museum of Natural History 56 (2): 185–244. doi:10.3374/014.056.0204. ISSN 0079-032X. Bibcode2015BPMNH..56..185V. http://www.bioone.org/doi/10.3374/014.056.0204. 
  20. Lyson, Tyler R.; Petermann, Holger; Miller, Ian M. (2021-01-02). "A new plastomenid trionychid turtle, Plastomenus joycei , sp. nov., from the earliest Paleocene (Danian) Denver Formation of south-central Colorado, U.S.A.". Journal of Vertebrate Paleontology 41 (1). doi:10.1080/02724634.2021.1913600. ISSN 0272-4634. Bibcode2021JVPal..41E3600L. 
  21. Longrich, Nicholas R.; Bhullar, Bhart-Anjan S.; Gauthier, Jacques A. (2012-12-26). "Mass extinction of lizards and snakes at the Cretaceous–Paleogene boundary". Proceedings of the National Academy of Sciences 109 (52): 21396–21401. doi:10.1073/pnas.1211526110. PMID 23236177. Bibcode2012PNAS..10921396L. 
  22. Ksepka, Daniel T.; Stidham, Thomas A.; Williamson, Thomas E. (2017-07-25). "Early Paleocene landbird supports rapid phylogenetic and morphological diversification of crown birds after the K–Pg mass extinction". Proceedings of the National Academy of Sciences 114 (30): 8047–8052. doi:10.1073/pnas.1700188114. PMID 28696285. Bibcode2017PNAS..114.8047K. 
  23. Kondrashov, Peter E.; Lucas, Spencer G. (2015). "Paleocene vertebrate faunas of the San Juan Basin, New Mexico". Vertebrate Paleontology in New Mexico 68. 
  24. Shelley, Sarah L.; Williamson, Thomas E.; Brusatte, Stephen L. (2018). "The osteology of Periptychus carinidens: A robust, ungulate-like placental mammal (Mammalia: Periptychidae) from the Paleocene of North America". PLOS ONE 13 (7). doi:10.1371/journal.pone.0200132. ISSN 1932-6203. Bibcode2018PLoSO..1300132S. 
  25. Atteberry, Madelaine R.; Eberle, Jaelyn J. (2021-04-18). "New earliest Paleocene (Puercan) periptychid 'condylarths' from the Great Divide Basin, Wyoming, USA". Journal of Systematic Palaeontology 19 (8): 565–593. doi:10.1080/14772019.2021.1924301. ISSN 1477-2019. Bibcode2021JSPal..19..565A. 
  26. Lucas, Spencer G.; Kondrashov, Peter E. (2004). "A new species of Deltatherium (Mammalia, Tillodontia) from the Paleocene of New Mexico". New Mexico Museum of Natural History and Science Bulletin 26. 
  27. Williamson, Thomas E.; Brusatte, Stephen L.; Secord, Ross; Shelley, Sarah (May 2016b). "A new taeniolabidoid multituberculate (Mammalia) from the middle Puercan of the Nacimiento Formation, New Mexico, and a revision of taeniolabidoid systematics and phylogeny: Revision of Taeniolabidoidea" (in en). Zoological Journal of the Linnean Society 177 (1): 183–208. doi:10.1111/zoj.12336. https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/zoj.12336. 
  28. Wheeler, George; White, Charles; Cope, Edward (1877). "Report upon United States geological surveys west of the one hundredth meridian, Volume IV: Paleontology". US Government Printing Office. https://pubs.er.usgs.gov/publication/70039253. 
  29. Fassett 1992.

References

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