Biology:Dinichthys

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Short description: Extinct genus of placoderm fish

Dinichthys
Temporal range: Late Devonian: Famennian, 371.1–359.3 Ma
Scientific classification e
Kingdom: Animalia
Phylum: Chordata
Class: †Placodermi
Order: †Arthrodira
Clade: †Phlyctaenioidei
Suborder: †Brachythoraci
Clade: †Eubrachythoraci
Clade: †Pachyosteomorphi
Clade: †Aspinothoracidi
Genus: †Dinichthys
Newberry, 1868
Species
  • Dinichthys herzeri Newberry, 1868 (type)
Synonyms
  • Ponerichthys Miller, 1892

Dinichthys (from Greek: δεινός deinós, 'terrible' and Greek: ἰχθύς ichthys 'fish') is an extinct monospecific genus of large marine arthrodire placoderm from the Late Devonian (Famennian stage) measuring around 3 metres (9.8 ft) long.[1] Fossils were recovered from the Ohio Shale Formation along the Olentangy River in Delaware County, Ohio.

Classification History

Dinichthys was originally described in 1868 by John Newberry on the basis of an incomplete skull roof and mandibles (holotype AMNH 81). Subsequently, many unrelated large arthrodires were originally classified together within this genus, including species now assigned to Dunkleosteus, Eastmanosteus, and Titanichthys. Notably, the type species of Dunkleosteus was originally described as Dinichthys terrelli by Newberry in 1873, and was later separated into Dunkleosteus by Jean-Pierre Lehman in 1956. Dunkleosteus was still thought to be closely related to Dinichthys, and they were grouped together in the family Dinichthyidae. However, in the 2010 Carr & Hlavin phylogenetic study, Dunkleosteus and Dinichthys were found to belong to two separate clades. Carr & Hlavin resurrected the family Dunkleosteidae and placed Dunkleosteus, Eastmanosteus, and a few other genera from Dinichthyidae within it.[2] Dinichthyidae, in turn, is left a monospecific family[3] and dismissed as a family grouping,[4] and the genus Dinichthys is now considered a monotypic genus, containing only the type species, D. herzeri.[2]

Phylogeny

Dinichthys is a member of the clade Aspinothoracidi, which belongs to the clade Pachyosteomorphi, one of the two major clades within Eubrachythoraci. The cladogram below shows the phylogeny of Dinichthys:[5]

Eubrachythoraci 
 

Coccosteomorphi

 
 Pachyosteomorphi 
 

Rhachiosteus pterygiatus

 
 
Dunkleosteoidea 
 

Eastmanosteus calliaspis

 
 
 
 

Eastmanosteus pustulosus

 
 

Kiangyousteus yohii

 
 
 
 

Golshanichthys asiatica

 
 
 

Westralichthys uwagedensis

 
 
 
 

Dunkleosteus raveri

 
 

Dunkleosteus terrelli

 
 
 
 

Dunkleosteus amblyodoratus

 
 Heterostiidae 
 

Heterosteus ingens

 
 

Yinosteus major

 
 
 
 
 
 
 
 
Aspinothoracidi
 

Tapinosteus heintzi

 
 
 
 

Bullerichthys fascidens

 
 
 

Kendrickichthys cavernosus

 
 

Bruntonichthys multidens

 
 
 
 
 

Dinichthys herzeri

 
 
 

Hadrosteus rapax

 
 
 

Gorgonichthys clarki

 
 Selenosteidae 
 

Heintzichthys gouldii

 
 

Pachyosteus bulla

 
 

Gymnotrachelus hydei

 
 

Stenosteus angustopectus

 
 

Brachyosteus dietrichi

 
 
 

Melanosteus occitanus

 
 

Rhinosteus parvulus

 
 
 
 
 
 
 
 
 
 
 

References

  1. ↑ Bashford Dean (1895). Fishes, Living and Fossil: An Outline of Their Forms and Probable Relationships. Macmillan and Company. p. 130. https://books.google.com/books?id=KlA9AAAAYAAJ&dq=Titanichthys+length&pg=PA130. Retrieved 2022-08-28. 
  2. ↑ 2.0 2.1 Carr, R.K.; Hlavin, W.J. (2010). "Two new species of Dunkleosteus Lehman, 1956, from the Ohio Shale Formation (USA, Famennian) and the Kettle Point Formation (Canada, Upper Devonian), and a cladistic analysis of the Eubrachythoraci (Placodermi, Arthrodira)". Zoological Journal of the Linnean Society 159 (1): 195–222. doi:10.1111/j.1096-3642.2009.00578.x. 
  3. ↑ Carr, Robert K.; William J. Hlavin (September 2, 1995). "Dinichthyidae (Placodermi):A paleontological fiction?". Geobios 28: 85–87. doi:10.1016/S0016-6995(95)80092-1. 
  4. ↑ You-An Zhu; Min Zhu (2013). "A redescription of Kiangyousteus yohii (Arthrodira: Eubrachythoraci) from the Middle Devonian of China, with remarks on the systematics of the Eubrachythoraci". Zoological Journal of the Linnean Society 169 (4): 798–819. doi:10.1111/zoj12089. 
  5. ↑ Zhu, You-An; Zhu, Min; Wang, Jun-Qing (1 April 2016). "Redescription of Yinostius major (Arthrodira: Heterostiidae) from the Lower Devonian of China, and the interrelationships of Brachythoraci". Zoological Journal of the Linnean Society 176 (4): 806–834. doi:10.1111/zoj.12356. ISSN 0024-4082. 

Wikidata ☰ Q112325116 entry