Biology:Rheiformes

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Rheiformes is an order that contains the family Rheidae (rheas).[1] It is in the infraclass Paleognathae, which contains all ratites. Extant members are found in South America. While the IOC World Bird List and the Clements Checklist categorise Rheiformes as its own order,[1][2] the BirdLife Data Zone includes rheas, along with ostriches, tinamous, cassowaries, emu, and kiwis, in the order Struthioniformes.[3] Of the two extant species of rheas recognized by the IUCN Red List, as of 2022, Rhea americana is listed as near threatened,[4] while Rhea pennata is listed as least concern.[5] From 2014 to 2022, the IUCN recognised Rhea tarapacensis as a separate species, and listed it as near threatened in its last assessment in 2020;[6] in 2022, it was again recognised as a subspecies of R. pennata.[5]: Taxonomy

Today, the order is represented by the sole living genus Rhea, though it contains 4-5 genera in total depending on the affiliation of the extinct genus Diogenornis. The taxonomy of the order is as follows:[7]

Order Rheiformes (Forbes, 1884) Furbringer, 1888 [Rheimorphae Bonaparte, 1849; Rheae Forbes 1884][8][9][10][11][12]

  • Family Rheidae (Bonaparte 1849) Bonaparte, 1853


Extant species summary

The IOC World Bird List (version 15.1) recognizes 2 species of Rheiformes.[14] As of January 2026, IUCN/BirdLife International have assessed both species within the order, but neither have a global population estimate.

Common name Binomial name Population Status Trend Notes Image
Greater rhea Rhea americana unknown[4] Template:IUCN-NT[4] Decrease[4] 175px
Lesser rhea

(Darwin's rhea)

Rhea pennata unknown[5] Template:IUCN-LC[5] Decrease[5] The populations of subspecies R. p. tarapacensis and R. p. garleppi are estimated to total 1,000-2,499 mature individuals. The population of the nominate subspecies, R. p. pennata, is expected to be much larger but has not been quantified.[5] 175px

References

  1. 1.0 1.1 Gill, Frank; Donsker, David; Rasmussen, Pamela, eds (11 August 2022). "Ratites: Ostriches to tinamous". IOC World Bird List. https://www.worldbirdnames.org/new/bow/ratites/. 
  2. Clements, James F.; Schulenberg, Thomas S.; Iliff, Marshall J.; Fredericks, Thomas A.; Gerbracht, Jeff A.; LePage, Denis; Billerman, Shawn M.; Sullivan, Brian L. et al. (25 October 2022). "The eBird/Clements checklist of Birds of the World". Cornell Lab of Ornithology. https://www.birds.cornell.edu/clementschecklist/updateindex/october-2022/. 
  3. "HBW and BirdLife Taxonomic Checklist". BirdLife International. July 2022. http://datazone.birdlife.org/species/taxonomy. 
  4. 4.0 4.1 4.2 4.3 BirdLife International. (2022). "Rhea americana". IUCN Red List of Threatened Species 2022. doi:10.2305/IUCN.UK.2022-2.RLTS.T22678073A219615764.en. https://www.iucnredlist.org/species/22678073/219615764. Retrieved 7 July 2024. 
  5. 5.0 5.1 5.2 5.3 5.4 5.5 BirdLife International. (2022). "Rhea pennata". IUCN Red List of Threatened Species 2022. doi:10.2305/IUCN.UK.2022-2.RLTS.T22678081A217016710.en. https://www.iucnredlist.org/species/22678081/217016710. Retrieved 7 July 2024. 
  6. BirdLife International. (2020). "Rhea tarapacensis". IUCN Red List of Threatened Species 2020. doi:10.2305/IUCN.UK.2020-3.RLTS.T22728206A177987446.en. https://www.iucnredlist.org/species/22728206/177987446. Retrieved 7 July 2024. 
  7. 7.0 7.1 Picasso, Mariana B.J.; Hospitaleche, Carolina Acosta; Mosto, María C. (2022). "An overview and update of South American and Antarctic fossil rheidae and putative ratitae (Aves, Palaeognathae)". Journal of South American Earth Sciences 115. doi:10.1016/j.jsames.2022.103731. ISSN 0895-9811. 
  8. Haaramo, Mikko (2007). "Paleognathia - paleognathous modern birds". http://www.helsinki.fi/~mhaaramo/metazoa/deuterostoma/chordata/archosauria/aves/palaeognathia/paleognathia_1.html. 
  9. "Taxonomic lists - Aves". http://www.paleofile.com/. 
  10. "Part 7 - Vertebrates". http://mave.tweakdsl.nl/tn/genera7.html. 
  11. Çınar, Ümüt (November 2015). "01 → Pᴀʟᴇᴏɢɴᴀᴛʜᴀᴇ: Sᴛʀᴜᴛʜɪᴏɴɪfᴏʀᴍᴇs, Rʜᴇɪfᴏʀᴍᴇs, Cᴀsᴜᴀʀɪɪfᴏʀᴍᴇs, Aᴘᴛᴇʀʏɢɪfᴏʀᴍᴇs, Aᴇᴘʏᴏʀɴɪᴛʜɪfᴏʀᴍᴇs, Dɪɴᴏʀɴɪᴛʜɪfᴏʀᴍᴇs, Lɪᴛʜᴏʀɴɪᴛʜɪfᴏʀᴍᴇs, Tɪɴᴀᴍɪfᴏʀᴍᴇs & Rᴇfᴇʀᴇɴᴄᴇs". http://kmoksy.com/zobot/birds_English.html. 
  12. Brodkob, Pierce (1963). "1- Archaeopterygiformes through Ardeiformes". Biological sciences. Catalogue of fossil birds (Bulletin of the Florida State Museum) 7 (4): 180–293. http://ufdc.ufl.edu/UF00001514/00001. Retrieved 30 December 2015. 
  13. Cite error: Invalid <ref> tag; no text was provided for refs named Tambussi2023
  14. "Ratites: Ostriches to tinamous – IOC World Bird List". https://www.worldbirdnames.org/bow/ratites/. 

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