Biology:Almond

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Short description: Species of nut

Almond
Botanical illustration
1897 illustration[1]
Branch of tree with green fruits.
Almond tree with ripening nuts.
Scientific classification edit
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Clade: Rosids
Order: Rosales
Family: Rosaceae
Genus: Prunus
Subgenus: Prunus subg. Amygdalus
Species:
P. amygdalus
Binomial name
Prunus amygdalus
Batsch, 1801
Synonyms[2][3]

The almond (Prunus amygdalus, syn. Prunus dulcis) is a species of small tree from the genus Prunus, cultivated worldwide for its seed, a culinary nut. Along with the peach, it is classified in the subgenus Amygdalus, distinguished from the other subgenera by corrugations on the shell (endocarp) surrounding the seed.[4]

The fruit of the almond is a drupe, consisting of an outer hull and a hard shell with the seed, which is not a true nut. Shelling almonds refers to removing the shell to reveal the seed. Almonds are sold shelled or unshelled. Blanched almonds are shelled almonds that have been treated with hot water to soften the seedcoat, which is then removed to reveal the white embryo. Once almonds are cleaned and processed, they can be stored over time. Almonds are used in many cuisines, often featuring prominently in desserts, such as marzipan.

The almond tree prospers in a moderate Mediterranean climate with cool winter weather. Native to Iran and surrounding countries[5] including the Levant, today it is rarely found wild in its original setting.[6] Almonds were one of the earliest domesticated fruit trees, due to the ability to produce quality offspring entirely from seed, without using suckers and cuttings. Evidence of domesticated almonds in the Early Bronze Age has been found in the archeological sites of the Middle East, and subsequently across the Mediterranean region and similar arid climates with cool winters.

California produces over half of the world's almond supply. Due to high acreage and water demand for almond cultivation, and need for pesticides, California almond production may be unsustainable, especially during the persistent drought and heat from climate change in the 21st century.[7] Droughts in California have caused some producers to leave the industry, leading to lower supply and increased prices.[7]

Description

The almond is a deciduous tree growing to 4–12.2 metres (13–40 feet) in height,[8] with a trunk of up to 30 centimetres (12 inches) in diameter. The young twigs are green at first, becoming purplish where exposed to sunlight, then gray in their second year. The leaves are 8–13 cm (3–5 in) long,[9] with a serrated margin and a 2.5 cm (1 in) petiole.

The flowers are white to pale pink, 3–5 cm (1–2 in) diameter with five petals, produced singly or in pairs and appearing before the leaves in early spring.[10][11] Almond grows best in Mediterranean climates with warm, dry summers and mild, wet winters. The optimal temperature for their growth is between 15 and 30 °C (59 and 86 °F) and the tree buds have a chilling requirement of 200 to 700 hours below 7.2 °C (45.0 °F) to break dormancy.[12]

Almonds begin bearing an economic crop in the third year after planting. Trees reach full bearing five to six years after planting. The fruit matures in the autumn, 7–8 months after flowering.[11][13]

The almond fruit is 3.5–6 cm (1 3⁄8–2 3⁄8 in) long. It is not a nut but a drupe. The outer covering, consisting of an outer exocarp, or skin, and mesocarp, or flesh, fleshy in other members of Prunus such as the plum and cherry, is instead a thick, leathery, gray-green coat (with a downy exterior), called the hull. Inside the hull is a woody endocarp which forms a reticulated, hard shell (like the outside of a peach pit) called the pyrena. Inside the shell is the edible seed, commonly called a nut. Generally, one seed is present, but occasionally two occur. After the fruit matures, the hull splits and separates from the shell, and an abscission layer forms between the stem and the fruit so that the fruit can fall from the tree.[14]

  • Almond blossoms

  • Young almond fruit

  • Green almonds

  • Mature almond nut

  • Almond shell

  • A rare double-seeded shell

  • Harvested almonds

  • Blanched almonds

Taxonomy

Sweet and bitter almonds

Flowering (sweet) almond tree
Blossoming of bitter almond tree

The seeds of Prunus dulcis var. dulcis are predominantly sweet[15][16] but some individual trees produce seeds that are somewhat more bitter. The genetic basis for bitterness involves a single gene, the bitter flavor furthermore being recessive,[17][18] both aspects making this trait easier to domesticate. The fruits from Prunus dulcis var. amara are always bitter, as are the kernels from other species of genus Prunus, such as apricot, peach and cherry (although to a lesser extent).

The bitter almond is slightly broader and shorter than the sweet almond and contains about 50% of the fixed oil that occurs in sweet almonds. It also contains the enzyme emulsin which, in the presence of water, acts on the two soluble glucosides amygdalin and prunasin[19] yielding glucose, cyanide and the essential oil of bitter almonds, which is nearly pure benzaldehyde, the chemical causing the bitter flavor. Bitter almonds may yield 4–9 milligrams of hydrogen cyanide per almond[20] and contain 42 times higher amounts of cyanide than the trace levels found in sweet almonds.[21] The origin of cyanide content in bitter almonds is via the enzymatic hydrolysis of amygdalin.[21] P450 monooxygenases are involved in the amygdalin biosynthetic pathway. A point mutation in a bHLH transcription factor prevents transcription of the two cytochrome P450 genes, resulting in the sweet kernel trait.[22]

Etymology

The word almond comes from Old French almande or alemande,[23] Late Latin amandula, amindula, derived from amygdala from the Ancient Greek amygdálē (ἀμυγδάλη)[23][24] (cf. amygdala, an almond-shaped portion of the brain).[25] Late Old English had amygdales, "almonds".[24]

The adjective amygdaloid (literally 'like an almond') is used to describe objects which are roughly almond-shaped, particularly a shape which is part way between a triangle and an ellipse. For example, the amygdala of the brain uses a direct borrowing of the Greek term amygdalē.[26]

Distribution and habitat

Almond is native to Iran and its surrounding regions,[5] including the Levant area.[6] It was spread by humans in ancient times along the shores of the Mediterranean into northern Africa and southern Europe, and more recently transported to other parts of the world, notably California, United States.[27] The wild form of domesticated almond grows in parts of the Levant.[28]

Selection of the sweet type from the many bitter types in the wild marked the beginning of almond domestication. It is unclear as to which wild ancestor of the almond created the domesticated species. The species Prunus fenzliana may be the most likely wild ancestor of the almond, in part because it is native to Armenia and western Azerbaijan, where it was apparently domesticated.[6] Wild almond species were grown by early farmers, "at first unintentionally in the garbage heaps, and later intentionally in their orchards".[29]

Cultivation

Persian miniature depiction of the almond harvest at Qand-i Badam, Fergana Valley (16th century)[30]
A grove of almond trees
An almond shaker before and during a tree's harvest

Almonds were one of the earliest domesticated fruit trees, due to "the ability of the grower to raise attractive almonds from seed.[6] Thus, in spite of the fact that this plant does not lend itself to propagation from suckers or from cuttings, it could have been domesticated even before the introduction of grafting".[28] Domesticated almonds appear in the Early Bronze Age (3000–2000 BC), such as the archaeological sites of Numeira (Jordan),[6] or possibly earlier. Another well-known archaeological example of the almond is the fruit found in Tutankhamun's tomb in Egypt (c. 1325 BC), probably imported from the Levant.[28] An article on almond tree cultivation in Spain is brought down in Ibn al-'Awwam's 12th-century agricultural work, Book on Agriculture.[31]

Of the European countries that the Royal Botanic Garden Edinburgh reported as cultivating almonds, Germany[32] is the northernmost, though the domesticated form can be found as far north as Iceland.[33]

Varieties

Almond trees are small to medium sized but commercial cultivars can be grafted onto a different root-stock to produce smaller trees. Varieties include:

  • Nonpareil – originates in the 1800s. A large tree that produces large, smooth, thin-shelled almonds with 60–65% edible kernel per nut. Requires pollination from other almond varieties for good nut production.[34]
  • Tuono – originates in Italy. Has thicker, hairier shells with only 32% of edible kernel per nut. The thicker shell gives some protection from pests such as the navel orangeworm. Does not require pollination by other almond varieties.[34]
  • Mariana – used as a rootstock to result in smaller trees

Breeding

Breeding programmes have found the high shell-seal trait.[35]

Pollination

The most widely planted varieties of almond are self-incompatible; hence these trees require pollen from a tree with different genetic characters to produce seeds. Almond orchards therefore must grow mixtures of almond varieties. In addition, the pollen is transferred from flower to flower by insects; therefore commercial growers must ensure there are enough insects to perform this task.[36] The large scale of almond production in the U.S. creates a significant problem of providing enough pollinating insects. Additional pollinating insects are therefore brought to the trees. The pollination of California's almonds is the largest annual managed pollination event in the world, with 1.4 million hives (nearly half of all beehives in the US) being brought to the almond orchards each February.[37]

Much of the supply of bees is managed by pollination brokers, who contract with migratory beekeepers from at least 49 states for the event. This business was heavily affected by colony collapse disorder at the turn of the 21st century, causing a nationwide shortage of honey bees and increasing the price of insect pollination. To partially protect almond growers from these costs, researchers at the Agricultural Research Service, part of the United States Department of Agriculture (USDA), developed self-pollinating almond trees that combine this character with quality characters such as a flavor and yield.[34] Self-pollinating almond varieties exist, but they lack some commercial characters. However, through natural hybridisation between different almond varieties, a new variety that was self-pollinating with a high yield of commercial quality nuts was produced.

Diseases

Main page: Biology:List of almond diseases

Almond trees can be attacked by an array of damaging microbes, fungal pathogens, plant viruses, and bacteria.[38]

Pests

Pavement ants (Tetramorium caespitum), southern fire ants (Solenopsis xyloni), and thief ants (Solenopsis molesta) are seed predators.[38] Bryobia rubrioculus mites are most known for their damage to this crop.[39]

Sustainability

Almond production in California is concentrated mainly in the Central Valley,[40] where the mild climate, rich soil, abundant sunshine and water supply make for ideal growing conditions. Due to the persistent droughts in California in the early 21st century, it became more difficult to raise almonds in a sustainable manner.[41][37] The issue is complex because of the high amount of water needed to produce almonds: a single almond requires roughly 1.1 US gallons (0.92 imperial gallons; 4.2 litres) of water to grow properly.[40][41][42] Regulations related to water supplies are changing so some growers have destroyed their current almond orchards to replace with either younger trees or a different crop such as pistachio that needs less water.[43]

Almond tree with blossoming flowers, Valley of Elah, Israel

Sustainability strategies implemented by the Almond Board of California and almond farmers include:[37][44][45]

  • tree and soil health, and other farming practices
  • minimizing dust production during the harvest
  • bee health
  • irrigation guidelines for farmers
  • food safety
  • use of waste biomass as coproducts with a goal to achieve zero waste
  • use of solar energy during processing
  • job development
  • support of scientific research to investigate potential health benefits of consuming almonds
  • international education about sustainability practices

Production

Almonds (with shell)
Production in 2020
Country Tonnes
 United States 2,370,021
 Spain 416,950
 Australia 221,886
 Iran 164,348
 Turkey 159,187
 Morocco 134,436
World
 4,140,043
Source: FAOSTAT of the United Nations[46]

In 2020, world production of almonds was 4.1 million tonnes, led by the United States providing 57% of the world total (table). Other leading producers were Spain, Australia, and Iran.

United States

In the United States, production is concentrated in California where 1,000,000 acres (400,000 ha) and six different almond varieties were under cultivation in 2017, with a yield of 2.25 billion pounds (1.02 billion kilograms) of shelled almonds.[47] California production is marked by a period of intense pollination during late winter by rented commercial bees transported by truck across the U.S. to almond groves, requiring more than half of the total U.S. commercial honeybee population.[48] The value of total U.S. exports of shelled almonds in 2016 was $3.2 billion.[49]

All commercially grown almonds sold as food in the U.S. are sweet cultivars. The U.S. Food and Drug Administration reported in 2010 that some fractions of imported sweet almonds were contaminated with bitter almonds, which contain cyanide.[50]

Spain

Spain has diverse commercial cultivars of almonds grown in Catalonia, Valencia, Murcia, Andalusia, and Aragón regions, and the Balearic Islands.[51] Production in 2016 declined 2% nationally compared to 2015 production data.[51]

The 'Marcona' almond cultivar is recognizably different from other almonds and is marketed by name.[52] The kernel is short, round, relatively sweet, and delicate in texture. Its origin is unknown and has been grown in Spain for a long time; the tree is very productive, and the shell of the nut is very hard.[52]

Australia

Australia is the largest almond production region in the Southern Hemisphere. Most of the almond orchards are located along the Murray River corridor in New South Wales, Victoria, and South Australia.[53][54]

Toxicity

Bitter almonds contain 42 times higher amounts of cyanide than the trace levels found in sweet almonds.[21] Extract of bitter almond was once used medicinally but even in small doses, effects are severe or lethal, especially in children; the cyanide must be removed before consumption.[21] The acute oral lethal dose of cyanide for adult humans is reported to be 0.5–3.5 mg/kg (0.2–1.6 mg/lb) of body weight (approximately 50 bitter almonds), so that for children consuming 5–10 bitter almonds may be fatal.[21] Symptoms of eating such almonds include vertigo and other typical cyanide poisoning effects.[50]

Almonds may cause allergy or intolerance. Cross-reactivity is common with peach allergens (lipid transfer proteins) and tree nut allergens. Symptoms range from local signs and symptoms (e.g., oral allergy syndrome, contact urticaria) to systemic signs and symptoms including anaphylaxis (e.g., urticaria, angioedema, gastrointestinal and respiratory symptoms).[55]

Almonds are susceptible to aflatoxin-producing molds.[56] Aflatoxins are potent carcinogenic chemicals produced by molds such as Aspergillus flavus and Aspergillus parasiticus.[57] The mold contamination may occur from soil, previously infested almonds, and almond pests such as navel-orange worm. High levels of mold growth typically appear as gray to black filament-like growth. It is unsafe to eat mold-infected tree nuts.

Some countries have strict limits on allowable levels of aflatoxin contamination of almonds and require adequate testing before the nuts can be marketed to their citizens. The European Union, for example, introduced a requirement since 2007 that all almond shipments to the EU be tested for aflatoxin. If aflatoxin does not meet the strict safety regulations, the entire consignment may be reprocessed to eliminate the aflatoxin or it must be destroyed.[58][59]

Breeding programs have found the high shell-seal trait.[35] High shell-seal provides resistance against these Aspergillus species and so against the development of their toxins.[35]

Mandatory pasteurization in California

After tracing cases of salmonellosis to almonds, the USDA approved a proposal by the Almond Board of California to pasteurize almonds sold to the public. After publishing the rule in March 2007, the almond pasteurization program became mandatory for California companies effective 1 September 2007.[60] Raw, untreated California almonds have not been available in the U.S. since then.

California almonds labeled "raw" must be steam-pasteurized or chemically treated with propylene oxide (PPO). This does not apply to imported almonds[61] or almonds sold from the grower directly to the consumer in small quantities.[62] The treatment also is not required for raw almonds sold for export outside of North America.

The Almond Board of California states: "PPO residue dissipates after treatment". The U.S. Environmental Protection Agency has reported: "Propylene oxide has been detected in fumigated food products; consumption of contaminated food is another possible route of exposure". PPO is classified as Group 2B ("possibly carcinogenic to humans").[63]

The USDA-approved marketing order was challenged in court by organic farmers organized by the Cornucopia Institute, a Wisconsin-based farm policy research group which filed a lawsuit in September 2008. According to the institute, this almond marketing order has imposed significant financial burdens on small-scale and organic growers and damaged domestic almond markets. A federal judge dismissed the lawsuit in early 2009 on procedural grounds. In August 2010, a federal appeals court ruled that the farmers have a right to appeal the USDA regulation. In March 2013, the court vacated the suit on the basis that the objections should have been raised in 2007 when the regulation was first proposed.[64]

Uses

Nutrition

Almonds
Mandorle sgusciate.jpg
Nutritional value per 100 g (3.5 oz)
Energy2,423 kJ (579 kcal)
21.6 g
Starch0.7 g
Sugars 4.4 g
0.00 g
Dietary fiber12.5 g
49.9 g
Saturated3.8 g
Monounsaturated31.6 g
Polyunsaturated12.3 g
21.2 g
Tryptophan0.214 g
Threonine0.598 g
Isoleucine0.702 g
Leucine1.488 g
Lysine0.580 g
Methionine0.151 g
Cystine0.189 g
Phenylalanine1.120 g
Tyrosine0.452 g
Valine0.817 g
Arginine2.446 g
Histidine0.557 g
Alanine1.027 g
Aspartic acid2.911 g
Glutamic acid6.810 g
Glycine1.469 g
Proline1.032 g
Serine0.948 g
VitaminsQuantity %DV†
Vitamin A equiv.
0%
1 μg
1 μg
Vitamin A1 IU
Thiamine (B1)
18%
0.211 mg
Riboflavin (B2)
85%
1.014 mg
Niacin (B3)
23%
3.385 mg
Pantothenic acid (B5)
9%
0.469 mg
Vitamin B6
11%
0.143 mg
Folate (B9)
13%
50 μg
Choline
11%
52.1 mg
Vitamin C
0%
0 mg
Vitamin D
0%
0 μg
Vitamin E
171%
25.6 mg
Vitamin K
0%
0.0 μg
MineralsQuantity %DV†
Calcium
26%
264 mg
Copper
50%
0.99 mg
Iron
29%
3.72 mg
Magnesium
75%
268 mg
Manganese
109%
2.285 mg
Phosphorus
69%
484 mg
Potassium
15%
705 mg
Selenium
4%
2.5 μg
Sodium
0%
1 mg
Zinc
32%
3.08 mg
Other constituentsQuantity
Water4.4 g
Link to USDA Database entry
†Percentages are roughly approximated using US recommendations for adults.
Source: USDA Nutrient Database
Amandines de Provence, poster by Leonetto Cappiello, 1900, which shows a woman eating almond biscuits (almond cookies)

Almonds are 4% water, 22% carbohydrates, 21% protein, and 50% fat. In a 100-gram (3 1⁄2-ounce) reference amount, almonds supply 2,420 kilojoules (579 kilocalories) of food energy. The almond is a nutritionally dense food, providing a rich source (20% or more of the Daily Value, DV) of the B vitamins riboflavin and niacin, vitamin E, and the essential minerals calcium, copper, iron, magnesium, manganese, phosphorus, and zinc. Almonds are a moderate source (10–19% DV) of the B vitamins thiamine, vitamin B6, and folate, choline, and the essential mineral potassium. They also contain substantial dietary fiber, the monounsaturated fat, oleic acid, and the polyunsaturated fat, linoleic acid. Typical of nuts and seeds, almonds are a source of phytosterols such as beta-sitosterol, stigmasterol, campesterol, sitostanol, and campestanol.[65]

Health

Almonds are included as a good source of protein among recommended healthy foods by the U.S. Department of Agriculture (USDA).[66] A 2016 review of clinical research indicated that regular consumption of almonds may reduce the risk of heart disease by lowering blood levels of LDL cholesterol.[67][68]

Culinary

While the almond is often eaten on its own, raw or toasted, it is also a component of various dishes. Almonds are available in many forms, such as whole, slivered, and ground into flour. Almond pieces around 2–3 millimetres (1⁄16–1⁄8 in) in size, called "nibs", are used for special purposes such as decoration.[69]

Almonds are a common addition to breakfast muesli or oatmeal.

  • Almond cream cake covered in slivered almonds

  • Colomba di Pasqua, traditional Italian Easter bread

  • Danish cream cake covered with marzipan

Desserts

A wide range of classic sweets feature almonds as a central ingredient. Marzipan was developed in the Middle Ages. Since the 19th century almonds have been used to make bread, almond butter, cakes and puddings, candied confections, almond cream-filled pastries, nougat, cookies (macaroons, biscotti and qurabiya), and cakes (financiers, Esterházy torte), and other sweets and desserts.[70]

The young, developing fruit of the almond tree can be eaten whole (green almonds) when they are still green and fleshy on the outside and the inner shell has not yet hardened. The fruit is somewhat sour, but is a popular snack in parts of the Middle East, eaten dipped in salt to balance the sour taste. Also in the Middle East they are often eaten with dates. They are available only from mid-April to mid-June in the Northern Hemisphere; pickling or brining extends the fruit's shelf life.

Marzipan

Marzipan, a smooth, sweetened almond paste, is used in a number of elegant cakes and desserts. Princess cake is covered by marzipan (similar to fondant), as is Battenberg cake. In Sicily, sponge cake is covered with marzipan to make cassatella di sant'Agata and cassata siciliana, and marzipan is dyed and crafted into realistic fruit shapes to make frutta martorana. The Andalusian Christmas pastry pan de Cádiz is filled with marzipan and candied fruit.

World cuisines

  • In French cuisine, alternating layers of almond and hazelnut meringue are used to make the dessert dacquoise. Pithivier is one of many almond cream-filled pastries.
  • In Germany, Easter bread called Deutsches Osterbrot is baked with raisins and almonds.
  • In Greece almond flour is used to make amygdalopita, a glyka tapsiou dessert cake baking in a tray. Almonds are used for kourabiedes, a Greek version of the traditional quarabiya almond biscuits. A soft drink known as soumada is made from almonds in various regions.
  • In Saudi Arabia, almonds are a typical embellishment for the rice dish kabsa.[71][72]
  • In Iran, green almonds are dipped in sea salt and eaten as snacks on street markets; they are called chaqale bâdam. Candied almonds called noghl are served alongside tea and coffee. Also, sweet almonds are used to prepare special food for babies, named harire badam. Almonds are added to some foods, cookies, and desserts, or are used to decorate foods. People in Iran consume roasted nuts for special events, for example, during New Year (Nowruz) parties.
  • In Italy, colomba di Pasqua is a traditional Easter cake made with almonds. Bitter almonds are the base for amaretti cookies, a common dessert. Almonds are also a common choice as the nuts to include in torrone.
  • In Morocco, almonds in the form of sweet almond paste are the main ingredient in pastry fillings, and several other desserts. Fried blanched whole almonds are also used to decorate sweet tajines such as lamb with prunes. Southwestern Berber regions of Essaouira and Souss are also known for amlou, a spread made of almond paste, argan oil, and honey. Almond paste is also mixed with toasted flour and among others, honey, olive oil or butter, anise, fennel, sesame seeds, and cinnamon to make sellou (also called zamita in Meknes or slilou in Marrakech), a sweet snack known for its long shelf life and high nutritive value.
  • In Indian cuisine, almonds are the base ingredients of pasanda-style and Mughlai curries. Badam halva is a sweet made from almonds with added coloring. Almond flakes are added to many sweets (such as sohan barfi), and are usually visible sticking to the outer surface. Almonds form the base of various drinks which are supposed to have cooling properties. Almond sherbet or sherbet-e-badaam, is a popular summer drink. Almonds are also sold as a snack with added salt.
  • In Israel almonds are used as a topping for tahini cookies or eaten as a snack.
  • In Spain Marcona almonds are usually toasted in oil and lightly salted. They are used by Spanish confectioners to prepare a sweet called turrón.
  • In Arabian cuisine, almonds are commonly used as garnishing for Mansaf.

Certain natural food stores sell "bitter almonds" or "apricot kernels" labeled as such, requiring significant caution by consumers for how to prepare and eat these products.[73]

Milk

Main page: Biology:Almond milk

Almonds can be processed into a milk substitute called almond milk; the nut's soft texture, mild flavor, and light coloring (when skinned) make for an efficient analog to dairy, and a soy-free choice for lactose intolerant people and vegans. Raw, blanched, and lightly toasted almonds work well for different production techniques, some of which are similar to that of soy milk and some of which use no heat, resulting in raw milk.

Almond milk, along with almond butter and almond oil, are versatile products used in both sweet and savoury dishes.

In Moroccan cuisine, sharbat billooz, a common beverage, is made by blending blanched almonds with milk, sugar and other flavorings.[74]

Flour and skins

Almond flour or ground almond meal combined with sugar or honey as marzipan is often used as a gluten-free alternative to wheat flour in cooking and baking.[75]

Almonds contain polyphenols in their skins consisting of flavonols, flavan-3-ols, hydroxybenzoic acids and flavanones[76] analogous to those of certain fruits and vegetables. These phenolic compounds and almond skin prebiotic dietary fiber have commercial interest as food additives or dietary supplements.[76][77]

Syrup

Historically, almond syrup was an emulsion of sweet and bitter almonds, usually made with barley syrup (orgeat syrup) or in a syrup of orange flower water and sugar, often flavored with a synthetic aroma of almonds.[21] Orgeat syrup is an important ingredient in the Mai Tai and many other Tiki drinks.[78][79][80]

Due to the cyanide found in bitter almonds, modern syrups generally are produced only from sweet almonds. Such syrup products do not contain significant levels of hydrocyanic acid, so are generally considered safe for human consumption.[21]

Oils

Almond oil
Oil, almond
Nutritional value per 100 g
Energy3,699 kJ (884 kcal)
100 g
Saturated8.2 g
Monounsaturated69.9 g
Polyunsaturated17.4 g
0
17.4 g
VitaminsQuantity %DV†
Vitamin E
261%
39.2 mg
Vitamin K
7%
7.0 μg
MineralsQuantity %DV†
Iron
0%
0 mg
Link to USDA Database entry
†Percentages are roughly approximated using US recommendations for adults.

Almonds are a rich source of oil, with 50% of kernel dry mass as fat (whole almond nutrition table). In relation to total dry mass of the kernel, almond oil contains 32% monounsaturated oleic acid (an omega-9 fatty acid), 13% linoleic acid (a polyunsaturated omega-6 essential fatty acid), and 10% saturated fatty acid (mainly as palmitic acid). Linolenic acid, a polyunsaturated omega-3 fat, is not present (table). Almond oil is a rich source of vitamin E, providing 261% of the Daily Value per 100 millilitres.

When almond oil is analyzed separately and expressed per 100 grams as a reference mass, the oil provides 3,700 kJ (884 kcal) of food energy, 8 grams of saturated fat (81% of which is palmitic acid), 70 grams of oleic acid, and 17 grams of linoleic acid (oil table).

Oleum amygdalae, the fixed oil, is prepared from either sweet or bitter almonds, and is a glyceryl oleate with a slight odour and a nutty taste. It is almost insoluble in alcohol but readily soluble in chloroform or ether. Almond oil is obtained from the dried kernel of almonds.[81] Sweet almond oil is used as a carrier oil in aromatherapy and cosmetics while bitter almond oil, containing benzaldehyde, is used as a food flavouring and in perfume.[36]

In culture

The almond is highly revered in some cultures. The tree originated in the Middle East. In the Bible, the almond is mentioned ten times, beginning with Genesis 43:11, where it is described as "among the best of fruits". In Numbers 17, Levi is chosen from the other tribes of Israel by Aaron's rod, which brought forth almond flowers. The almond blossom supplied a model for the menorah which stood in the Holy Temple, "Three cups, shaped like almond blossoms, were on one branch, with a knob and a flower; and three cups, shaped like almond blossoms, were on the other … on the candlestick itself were four cups, shaped like almond blossoms, with its knobs and flowers" (Exodus 25:33–34; 37:19–20). Many Sephardic Jews give five almonds to each guest before special occasions like weddings.[82]

Similarly, Christian symbolism often uses almond branches as a symbol of the virgin birth of Jesus; paintings and icons often include almond-shaped haloes encircling the Christ Child and as a symbol of Mary. The word "luz", which appears in Genesis 30:37, sometimes translated as "hazel", may actually be derived from the Aramaic name for almond (Luz), and is translated as such in the New International Version and other versions of the Bible.[83] The Arabic name for almond is لوز "lauz" or "lūz". In some parts of the Levant and North Africa, it is pronounced "loz", which is very close to its Aramaic origin.

The Entrance of the flower (La entrada de la flor) is an event celebrated on 1 February in Torrent, Spain, in which the clavarios and members of the Confrerie of the Mother of God deliver a branch of the first-blooming almond-tree to the Virgin.[84]

See also

References

  1. ↑ illustration from Franz Eugen Köhler, Köhler's Medizinal-Pflanzen, 1897
  2. ↑ "Prunus amygdalus Batsch" (in en). Kew Science. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:60439867-2. 
  3. ↑ "The Plant List, Prunus dulcis (Mill.) D.A.Webb". http://www.theplantlist.org/tpl1.1/record/rjp-375. 
  4. ↑ "Almond Tree – Learn About Nature" (in en-US). 27 July 2016. https://www.learnaboutnature.com/plants/trees/almond-tree/. 
  5. ↑ 5.0 5.1 "BĀDĀM – Encyclopaedia Iranica". http://www.iranicaonline.org/articles/badam-almond. "The Amygdalus communis (or Prunus amygdalus), though undoubtedly native to the Iranian land-mass, is seldom found in natural stands there today." 
  6. ↑ 6.0 6.1 6.2 6.3 6.4 G. Ladizinsky (1999). "On the origin of almond". Genetic Resources and Crop Evolution 46 (2): 143–147. doi:10.1023/A:1008690409554. 
  7. ↑ 7.0 7.1 "Climate Change In California Is Threatening The World's Top Almond Producer" (in en). NPR. Associated Press. 17 August 2021. https://www.npr.org/2021/08/17/1028452988/climate-change-california-drought-heat-almond-production. 
  8. ↑ U.S. Department of the Army (2019). The Official U.S. Army Illustrated Guide to Edible Wild Plants. Guilford, CT: Lyons Press. pp. 10. ISBN 978-1-4930-4039-1. OCLC 1043567121. https://books.google.com/books?id=T2p7DwAAQBAJ&pg=PA10. 
  9. ↑ Bailey, L.H.; Bailey, E.Z.; the staff of the Liberty Hyde Bailey Hortorium. 1976. Hortus third: A concise dictionary of plants cultivated in the United States and Canada. Macmillan, New York.
  10. ↑ Rushforth, Keith (1999). Collins wildlife trust guide trees: a photographic guide to the trees of Britain and Europe. London: Harper Collins. ISBN 0-00-220013-9. 
  11. ↑ 11.0 11.1 Griffiths, Mark D.; Anthony Julian Huxley (1992). The New Royal Horticultural Society dictionary of gardening. London: Macmillan Press. ISBN 0-333-47494-5. 
  12. ↑ "Fruit Cultural Data — P – California Rare Fruit Growers, Inc." (in en-US). https://crfg.org/home/library/crfg-fruit-list/fruit-cultural-data-2/fruit-cultural-data-p/. 
  13. ↑ "University of California Sample Cost Study to Produce Almonds". http://coststudies.ucdavis.edu/files/almondvs08sprink.pdf. 
  14. ↑ Doll, David (22 June 2009). "The Seasonal Patterns of Almond Production". University of California Cooperative Extension. https://thealmonddoctor.com/2009/06/22/the-seasonal-patterns-of-almond-production/. 
  15. ↑ Karl-Franzens-Universität (Graz). "Almond (Prunus dulcis [Mill. D. A. Webb.)"]. http://www.uni-graz.at/~katzer/engl/Prun_dul.html. 
  16. ↑ "Almond and bitter almond". from Quirk Books: www.quirkbooks.com. http://www.chow.com/ingredients/265. 
  17. ↑ Heppner, Myer J (7 April 1923). "The factor for bitterness in the sweet almond". Genetics 8 (4): 390–392. doi:10.1093/genetics/8.4.390. PMID 17246020. 
  18. ↑ Dicenta, Federico; Ortega, Encarnacion; Martinez-Gomez, Pedro (January 2007). "Use of recessive homozygous genotypes to assess genetic control of kernel bitterness in almond". Euphytica (Springer) 153 (1–2): 221–225. doi:10.1007/s10681-006-9257-6. 
  19. ↑ "Prunasin hydrolases during fruit development in sweet and bitter almonds". Plant Physiology 158 (4): 1916–32. April 2012. doi:10.1104/pp.111.192021. PMID 22353576. 
  20. ↑ "Cyanide poisoning after bitter almond ingestion". West. J. Med. 136 (1): 65–9. January 1982. PMID 7072244. 
  21. ↑ 21.0 21.1 21.2 21.3 21.4 21.5 21.6 "Potential Toxic Levels of Cyanide in Almonds (Prunus amygdalus), Apricot Kernels (Prunus armeniaca), and Almond Syrup". ISRN Toxicol 2013 (19 September): 610648. 2013. doi:10.1155/2013/610648. PMID 24171123. 
  22. ↑ Sánchez-Pérez, R.; Pavan, S.; Mazzeo, R.; Moldovan, C.; Aiese Cigliano, R.; Del Cueto, J.; Ricciardi, F.; Lotti, C. et al. (14 June 2019). "Mutation of a bHLH transcription factor allowed almond domestication" (in en). Science 364 (6445): 1095–1098. doi:10.1126/science.aav8197. ISSN 0036-8075. PMID 31197015. Bibcode: 2019Sci...364.1095S. 
  23. ↑ 23.0 23.1 Chisholm, Hugh, ed (1911). "Almond". Encyclopædia Britannica. 1 (11th ed.). Cambridge University Press. p. 716. 
  24. ↑ 24.0 24.1 "Almond". Online Etymology Dictionary, Douglas Harper. 2023. https://www.etymonline.com/search?q=almond. 
  25. ↑ "Almond". Almond. http://dictionary.reference.com/browse/almond. Retrieved 16 May 2012. 
  26. ↑ "The Amygdaloid Complex: Anatomy and Physiology". Physiological Reviews 83 (3): 803–834. 1 July 2003. doi:10.1152/physrev.00002.2003. PMID 12843409. 
  27. ↑ Rieger, Mark (2006). "Almond – Prunus dulcis". http://www.uga.edu/fruit/almond-prunus-dulcis/. 
  28. ↑ 28.0 28.1 28.2 Zohary, Daniel; Maria Hopf (2000). Domestication of plants in the old world: the origin and spread of cultivated plants in West Asia, Europe, and the Nile Valley. Oxford University Press. p. 186. ISBN 0-19-850356-3. 
  29. ↑ Diamond, Jared M. (1997). Guns, germs, and steel: the fates of human societies. New York: W.W. Norton. p. 118. ISBN 0-393-03891-2. 
  30. ↑ Bhawani (1590s). "Harvesting of the almond crop at Qand-i Badam". Baburnama. http://warfare.atspace.eu/Moghul/Baburnama/Harvesting_of_the_almond_crop_at_Qand-i_Badam.htm. 
  31. ↑ Ibn al-'Awwam, Yaḥyá (1864) (in fr). Le livre de l'agriculture d'Ibn-al-Awam (kitab-al-felahah). Paris: A. Franck. pp. 260–263 (ch. 7 – Article 20). OCLC 780050566. https://archive.org/details/lelivredelagric00algoog/page/n14/mode/2up.  (pp. 260–263 (Article XX)
  32. ↑ "Flora Europaea Search Results". Royal Botanic Garden Edinburgh. http://193.62.154.38/cgi-bin/nph-readbtree.pl/feout?FAMILY_XREF=&GENUS_XREF=Prunus&SPECIES_XREF=dulcis&TAXON_NAME_XREF=&RANK=. 
  33. ↑ "Prunus dulcis". Plants for a Future. http://www.ibiblio.org/pfaf/cgi-bin/arr_html?Prunus+dulcis. 
  34. ↑ 34.0 34.1 34.2 Alfredo Flores (6 April 2010). "ARS Scientists Develop Self-pollinating Almond Trees". USDA Agricultural Research Service. http://www.ars.usda.gov/is/pr/2010/100406.htm. 
  35. ↑ 35.0 35.1 35.2 Robens, Jane; Cary, Jeffrey W.; Campbell, Bruce C. (25–27 October 2000). "Introduction". Aflatoxin/Fumonisin Workshop 2000. Yosemite, California, USA. p. 17. http://www.ars.usda.gov/pacific-west-area/albany-ca/wrrc/ftdp/docs/aflatoxin-elimination-workshop-proceedings/. 
  36. ↑ 36.0 36.1 van Wyk, Ben-Erik (2019). Food plants of the world (2nd ed.). CABI. p. 342. ISBN 9781789241303. 
  37. ↑ 37.0 37.1 37.2 Alan Bjerga; Donna Cohen; Cindy Hoffman. "California Almonds Are Back After Four Years of Brutal Drought". Bloomberg.com (Bloomberg). https://www.bloomberg.com/graphics/2018-thirst-for-almonds/. 
  38. ↑ 38.0 38.1 "Almond – Diseases and Pests, Description, Uses, Propagation". http://plantvillage.psu.edu/topics/almond/infos. 
  39. ↑ "Brown Mite / Almond / Agriculture: Pest Management Guidelines / UC Statewide IPM Program (UC IPM)". UC Agriculture and Natural Resources (UC ANR). http://www.ipm.ucanr.edu/agriculture/almond/Brown-Mite/. 
  40. ↑ 40.0 40.1 Brodwin E; Lee S (8 April 2015). "Chart shows how some of your favorite foods could be making California's drought worse" (in en). Business Insider. http://www.businessinsider.com/amount-of-water-needed-to-grow-one-almond-orange-tomato-2015-4. 
  41. ↑ 41.0 41.1 "How Almonds Became A Scapegoat For California's Drought". US National Public Radio. 16 April 2015. https://www.npr.org/sections/thesalt/2015/04/16/399958203/how-almonds-became-a-scapegoat-for-californias-drought. 
  42. ↑ Mekonnen, M. M.; Hoekstra, A. Y.. "The green, blue and grey water footprint of crops and derived crop products". Twente Water Centre, University of Twente. https://hess.copernicus.org/articles/15/1577/2011/hess-15-1577-2011.pdf. 
  43. ↑ "Why Are Almond Growers Uprooting Their Orchards?". 2 July 2021. https://www.atlasobscura.com/articles/uprooting-almond-orchard. 
  44. ↑ "Annual report: Growing Good – Almond Sustainability 2018". Almond Board of California. 2018. https://newsroom.almonds.com/sites/default/files/pdf_file/Growing%20Good%202018.pdf. 
  45. ↑ "Almond industry forerunner of future farm practices, sustainability program internationally recognized". Western FarmPress. 6 March 2018. https://www.farmprogress.com/tree-nuts/almond-industry-forerunner-future-farm-practices-sustainability-program-internationally. 
  46. ↑ "Almonds (in shells) production in 2020, Crops/Regions/World list/Production Quantity (pick lists)". UN Food and Agriculture Organization, Corporate Statistical Database (FAOSTAT). 2020. http://www.fao.org/faostat/en/#data/QC. 
  47. ↑ Averill, Travis (6 July 2017). "2017 Almond Forecast". National Agricultural Statistics Service, US Department of Agriculture. http://www.almonds.com/sites/default/files/content/attachments/almond_objective_report_2017_presentation.pdf. 
  48. ↑ Ginger Zee; David Miller; Kelly Harold; Andrea Miller (16 January 2018). "Growing California almonds takes more than half of US honeybees". ABC News. https://abcnews.go.com/US/growing-california-almonds-takes-half-us-honeybees/story?id=52265334. 
  49. ↑ Workman, Daniel (25 July 2017). "Top Almonds Exporters by Country in 2016". World's Top Exports. http://www.worldstopexports.com/top-almonds-exporters-by-country/. 
  50. ↑ 50.0 50.1 "Cyanide and amygdalin as indicators of the presence of bitter almonds in imported raw almonds". Journal of Forensic Sciences 57 (5): 1313–7. September 2012. doi:10.1111/j.1556-4029.2012.02138.x. PMID 22564183. https://zenodo.org/record/1230689. 
  51. ↑ 51.0 51.1 "Tree nuts annual; Almonds, shelled basis; Report number SP1619". GAIN Report, US Department of Agriculture. 15 September 2016. https://gain.fas.usda.gov/Recent%20GAIN%20Publications/Tree%20Nuts%20Annual_Madrid_EU-28_9-8-2016.pdf. 
  52. ↑ 52.0 52.1 Gradziel, T.M. (2011). "Origin and Dissemination of Almonds". in J. Janick. Horticultural Reviews. 38. Wiley-Blackwell. p. 55. doi:10.1002/9780470872376.ch2. ISBN 9780470872376. https://eurekamag.com/research/038/900/038900919.php. Retrieved 10 April 2018. 
  53. ↑ "Where are Australian Almonds grown?". http://australianalmonds.com.au/enjoy/australian_almonds/where-are-australian-almonds-grown. 
  54. ↑ Gibson, Chris (5 February 2014). "Agri-comeback kids of 2014". Sydney Morning Herald. http://www.theland.com.au/story/3580611/agri-comeback-kids-of-2014/. 
  55. ↑ "Almond allergy". Food-info.net. 26 July 2001. http://www.food-info.net/uk/intol/almond.htm. 
  56. ↑ "The high cost of aflatoxins". Almond Board of California. 2009. http://www.almondboard.com/Handlers/Documents/The-High-Cost-of-Aflatoxins.pdf. 
  57. ↑ Rushing, Blake R.; Selim, Mustafa I. (2019). "Aflatoxin B1: A review on metabolism, toxicity, occurrence in food, occupational exposure, and detoxification methods". Food and Chemical Toxicology 124: 81–100. doi:10.1016/j.fct.2018.11.047. ISSN 0278-6915. PMID 30468841. 
  58. ↑ "Aflatoxins in food". European Food Safety Authority. 2010. http://www.efsa.europa.eu/en/topics/topic/aflatoxins-food. 
  59. ↑ "New EU Aflatoxin Levels and Sampling Plan". USDA Foreign Agricultural Service. 2010. http://gain.fas.usda.gov/Recent%20GAIN%20Publications/New%20EU%20Aflatoxin%20Levels%20and%20Sampling%20Plan_Brussels%20USEU_EU-27_3-9-2010.pdf. 
  60. ↑ "The Food Safety Program & Almond Pasteurization" (Press release). Almond Board of California. 17 September 2010. Archived from the original on 25 January 2010. Retrieved 17 September 2010.
  61. ↑ Agricultural Marketing Service (8 November 2006) "Almonds Grown in California: Changes to Incoming Quality Control Requirements" (71 FR -FR-65373 65373, 71 FR -FR-65374 65374, 71 FR -FR-65375 65375 and 71 FR -FR-65376 65376)
  62. ↑ Burke, Garance (29 June 2007). "Almond pasteurization rubs some feelings raw". Associated Press. http://www.insidebayarea.com/business/ci_6260038. 
  63. ↑ Harris LJ, ed (2013). Improving the Safety and Quality of Nuts. Elsevier, Woodhead Publishing Series in Food Science, Technology and Nutrition. pp. 36–37. ISBN 978-0-85709-748-4. https://books.google.com/books?id=wlpEAgAAQBAJ&pg=PA36. 
  64. ↑ "The Authentic Almond Project". The Cornucopia Institute.. http://www.cornucopia.org/almonds/. 
  65. ↑ "Effects of almond consumption on the reduction of LDL-cholesterol: a discussion of potential mechanisms and future research directions". Nutrition Reviews 69 (4): 171–85. April 2011. doi:10.1111/j.1753-4887.2011.00383.x. PMID 21457263. 
  66. ↑ "Protein foods: nutrients and health benefits". ChooseMyPlate.gov, USDA. 4 October 2018. https://www.choosemyplate.gov/protein-foods-nutrients-health. 
  67. ↑ Musa-Veloso, Kathy; Paulionis, Lina; Poon, Theresa; Lee, Han Youl (16 August 2016). "The effects of almond consumption on fasting blood lipid levels: a systematic review and meta-analysis of randomised controlled trials". Journal of Nutritional Science 5: e34. doi:10.1017/jns.2016.19. ISSN 2048-6790. PMID 27752301. 
  68. ↑ "Almonds". TH Chan School of Public Health, Harvard University. 2019. https://www.hsph.harvard.edu/nutritionsource/food-features/almonds/. 
  69. ↑ Sinclair, Charles (1 January 2009) (in en). Dictionary of Food: International Food and Cooking Terms from A to Z. A&C Black. pp. 45. ISBN 9781408102183. https://books.google.com/books?id=Mdwm7jI9J10C&pg=PT45. 
  70. ↑ Dolby, Richard (1830). The Cook's Dictionary: A New Family Manual of Cookery and Confectionery. 
  71. ↑ El Masri, Arwa (27 September 2011). Tea with Arwa: A Memoir of Family, Faith and Finding a Home in Australia. Hachette Australia. ISBN 9780733628528. https://books.google.com/books?id=6Gg0AgAAQBAJ&pg=PT22. 
  72. ↑ Salloum, Habeeb (28 February 2012). The Arabian Nights Cookbook: From Lamb Kebabs to Baba Ghanouj, Delicious Homestyle Arabian Cooking. Tuttle Publishing. ISBN 9781462905249. https://books.google.com/books?id=v5TTAgAAQBAJ&dq=Kabsa+rice+almonds&pg=PA74. 
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  75. ↑ Amsterdam, Elana (2009). The Gluten-Free Almond Flour Cookbook: Breakfasts, Entrees, and More. Random House of Canada. ISBN 978-1-58761-345-6. 
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  77. ↑ "Prebiotic effects of almonds and almond skins on intestinal microbiota in healthy adult humans". Anaerobe 26 (4): 1–6. 2014. doi:10.1016/j.anaerobe.2013.11.007. PMID 24315808. 
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  82. ↑ "Jewish Sephardi Wedding Recipes and Traditions" (in en-US). 13 July 2017. https://www.myjewishlearning.com/2017/07/13/jewish-sephardi-wedding-recipes-and-traditions/. 
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External links

Wikidata ☰ Q39918 entry



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